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Published: 2016-06-14
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FAMILY CERATOPOGONIDAE

División Entomología, Museo de La Plata, Paseo del Bosque s/n, 1900 La Plata, Argentina.
Grupo de Entomologia, Instituto de Biologia, Universidad de Antioquia, Medellín-Colombia.
Diptera catalogue Ceratopogonidae Colombia distribution localities species list

Abstract

Two hundred and thirty five species of Ceratopogonids arranged in 28 genera are listed for Colombia, mostly Culicoides (112 species) and Forcipomyia (50 species). For each listed species, information on type locality, type category and sex, location of type material, species distribution, localities for Colombia and main references, is provided. Localities are not specified for 17 species in which the literature searching was negative. Forcipomyia squamitibia Lutz and Bezzia clavipennis are excluded from the list.

 

References

  1. Aitken, T.H.G., Wirth, W.W., Williams, R.W, Davies, J.B. & Tikasingh, E.S. (1975) A review of the bloodsucking midges of Trinidad and Tobago, West Indies (Diptera: Ceratopogonidae). Journal of Entomology (B), 44, 101–144.

    Barbosa, F.A.S. (1947) Culicoides (Diptera: Heleidae) da região neotropica. Annais da Sociedade de Biologia de Pernambuco, 7, 3–30. [Nov.]

    Barbosa, F.A.S. (1951) A change of specific name in the genus Culicoides (Diptera, Heleidae). Proceedings of the Entomological Society of Washington, 53, 163.

    Barbosa, F.A.S. (1952) Novos subsídios para o conhecimento dos Culicoides neotrópicos (Diptera: Heleidae). Ph.D. Thesis, Univ. Recife, Recife (Brasil): Imprensa Industrial, 21 pp.

    Barreto, P. (1986) Catálogo de los Culicoides (Diptera, Ceratopogonidae) de Colombia. Colombia Médica, 17, 140–150.

    Barretto, M.P. (1944) Sobre o género Culicoides Latreille, 1809, com a descrição de trés novas espécies (Diptera, Ceratopogonidae). Annais da Faculdade de Medicina (da Universidade) de São Paulo, 20, 89–105, pls. 1–4.

    Blanton, F.S. & Wirth, W.W. (1979) The sand flies (Culicoides) of Florida (Diptera: Ceratopogonidae). Arthropods of Florida and neighboring land areas, 10, i–xv + 1–204.

    Borkent, A. (2015) World species of biting midges (Diptera: Ceratopogonidae), 241 pp. Available from: http://wwx.inhs.illinois.edu/files/1114/2384/5200/CeratopogonidCatalog.pdf (accessed 1 February 2015)

    Borkent, A. & Spinelli, G.R. (2007) Neotropical Ceratopogonidae (Diptera: Insecta). In: Adis J., Arias J.R., Rueda-Delgado G., Wantzen K.M. (Eds.), Aquatic Biodiversity in Latin America (ABLA). Vol. 4. Pensoft, Sofia-Moscow, pp. 1–198.

    Borkent, A., Spinelli, G.R. & Grogan, W.L. (2009) Ceratopogonidae (Biting midges, purrujas). In: Browne, V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 1. NRC Research Press, Ottawa, Ontario, 714 pp.

    Borkent, A. & Wirth, W.W. (1997) World species of biting midges (Diptera: Ceratopogonidae). Bulletin of the American Museum of Natural History, 233, 1–257.

    Brèthes, J. (1912) Descripción de un nuevo género y especie nueva de Chironomidae (Dipt). Anales del Museo Nacional de Buenos Aires, 15, 451–453.

    Brèthes, J. (1914) Descripción de seis Cécidomyidae (Dipt.) de Buenos Aires. Anales del Museo Nacional de Buenos Aires, 26, 151–156.

    Browne, J.E. (1980) New species of biting midges of the genus Culicoides from Colombia and the first description of the male of C. florenciae (Diptera: Ceratopogonidae). Journal of Medical Entomology, 17, 533–544.
    http://dx.doi.org/10.1093/jmedent/17.6.533

    Bystrak, P.G. & Wirth, W.W. (1978) The North American species of Forcipomyia, subgenus Euprojoannisia (Diptera: Ceratopogonidae). Unites States Department of Agriculture, Technical Bulletin, 1591, 1–51.

    Carter, H.F. (1921) A revision of the genus Leptoconops Skuse. Bulletin of Entomological Research, 12, 1-28. [June]

    Cavalieri, F. (1966) Notas sobre Ceratopogonidae (Diptera, Nematocera) V. Descripción de una nueva especie de Venezuela: Culicoides (Oecacta) birabeni sp. n. Physis, 26, 59–63.

    Chaudhuri, P.K., Das Gupta, S.K. & Chaudhuri, D.K. (1972) Biting midges of the genus Alluaudomyia Kieffer from India (Diptera: Ceratopogonidae). Oriental Insects, 6, 83–110.
    http://dx.doi.org/10.1080/00305316.1972.10434057

    Clastrier, J. (1971) Deux nouveaux Culicoides (Diptera, Ceratopogonidae) de la Guyane Française. Annales de Parasitologie, 46, 285–294.

    Clastrier, J. & Wirth, W.W. (1995) Révision des Forcipomyia du sous-genre Microhelea de la région Néotropicale, parasites de phasmes (Diptera: Ceratopogonidae). Annales de la Société Entomologique de France, 31, 97–150.

    Coquillett, D.W. (1899) A new dipterous family related to the Chironomidae. Entomological News, 10, 60–61.

    Coquillett, D.W. (1910) The Type species of the North American genera of Diptera. Proceedings of the United States National Museum, 37, 499–647.
    http://dx.doi.org/10.5479/si.00963801.37-1719.499

    Costa Lima, A. da (1937) Chave das espécies de Culicoides da região neotropical (Diptera: Ceratopogonidae). Memórias do Instituto Oswaldo Cruz, 32, 411–422.
    http://dx.doi.org/10.1590/S0074-02761937000300003

    Debenham, M.L. (1987) The biting midge genus Forcipomyia (Diptera: Ceratopogonidae) in the Australasian Region (exclusive of New Zealand). I. Introduction, key to subgenera, and the Thyridomyia and Trichohelea groups of subgenera. Invertebrate Taxonomy, 1, 35–119.
    http://dx.doi.org/10.1071/IT9870035

    De Meillon, B. & Wirth, W.W. (1980) A new subgenus of Forcipomyia, with descriptions of eight new species (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington, 82, 9–24.

    Díaz, F., Ronderos, M.M. & Spinelli, G.R. (2005) The immatures of the Neotropical species Culicoides venezuelensis Ortíz and Mirsa (Diptera: Ceratopogonidae). Transactions of the American Entomological Society, 131, 375–385.

    Díaz, F., Spinelli, G.R, Donato, M. & Ronderos, M.M. (2014) A taxonomic revision of the Patagonian species of the Dasyhelea mutabilis species group with a phylogenetic analysis of New World species (Diptera: Ceratopogonidae). Journal of Natural History, 48, 2117–2175.
    http://dx.doi.org/10.1080/00222933.2014.909062

    Díaz, F., Spinelli, G.R. & Ronderos, M.M. (2010) A revision of the species allied to Dasyhelea patagonica Ingram & Macfie (Diptera: Ceratopogonidae). Journal of Natural History, 44, 2825–2849.
    http://dx.doi.org/10.1080/00222933.2010.512424

    Dow, M.I. & Turner, E.C. (1976) A revision of the Nearctic species of the genus Bezzia (Diptera: Ceratopogonidae). Occassional Papers from the Department of Entomology, Virginia Polytechnic Institute and State University, Research Division Bulletin, 103, 1–162.

    Duret, J. P. & Lane, J. (1955) Novas Heteromyia da Argentina (Diptera. Ceratopogonidae). Dusenia, 6, 35–40.

    Edwards, F.W. (1922) On some Malayan and other species of Culicoides, with a note on the genus Lasiohelea. Bulletin of Entomological Resarch, 13, 161–167.
    http://dx.doi.org/10.1017/S0007485300028030

    Enderlein, G. (1912) Paryphoconus, eine neue Chironomidengattung aus Brasilien. Stettiner Entomologische Zeitung, 73, 57–60.

    Enderlein, G. (1936) Ordnung: Zweiflügler, Diptera. Abt. 16. In: Brohmer, P., Ehrmann, P., Ulmer, G. (Eds.), Die Tierwelt Mitteleuropas 6: Insekten III Teil. Quello & Meier, Leipzig.

    Ewen, A.B. & Saunders, L.G. (1958) Contributions toward a revision of the genus Atrichopogon based on characters of all stages (Diptera, Heleidae). Canadian Journal of Zoology, 36, 671–724.
    http://dx.doi.org/10.1139/z58-061

    Felippe-Bauer, M.L., Pinto Damasceno, C., Py-Daniel, V. & Spinelli, G.R. (2009) Culicoides baniwa sp. nov. From the brazilian Amazon region with a synopsis of the hylas species group (Diptera: Ceratopogonidae). Memórias do Instituto Oswaldo Cruz, 104, 851–857.

    http://dx.doi.org/10.1590/S0074-02762009000600005

    Felippe-Bauer, M.L. & Wirth, W.W. (1987) A new Culicoides of the stigmalis group (Diptera: Ceratopogonidae). Memórias do Instituto Oswaldo Cruz, 82, 415–419.
    http://dx.doi.org/10.1590/S0074-02761987000300014

    Forattini, O.P. (1954) Nova espécie de Culicoides (Diptera, Ceratopogonidae) do Brasil Central. Arquivos da Faculdade de Higiene e Saúde Pública da Universidade de São Paulo, 8, 189–192.
    http://dx.doi.org/10.11606/issn.2358-792X.v8i2p189-192

    Forattini, O.P. (1956) A new Culicoides from Panama (Diptera, Ceratopogonidae). Proceedings of the Entomological Society of Washington, 58, 35–36.

    Forattini, O.P. (1957) Culicoides da região neotropical (Diptera, Ceratopogonidae). Arquivos da Faculdade de Higiene e Saúde Pública da Universidade de São Paulo, 11, 159–526.
    http://dx.doi.org/10.11606/issn.2358-792X.v11i2p161-526

    Forattini, O.P. (1958) Nova espécie de Leptoconops s. str. da região neotropical (Diptera, Ceratopogonidae). Revista Brasileira de Entomologia, 8, 37–41.

    Fox, I. (1946) A review of the species of biting midges or Culicoides from the Caribbean Region (Diptera: Ceratopogonidae). Annals of the Entomological Society of America, 39, 248–258.
    http://dx.doi.org/10.1093/aesa/39.2.248

    Fox, I. (1947) Two new Central American biting midges or Culicoides (Diptera: Ceratopogonidae). Kuba, 3, 90–91. [June]

    Fox, I. (1948) Hoffmania, a new subgenus in Culicoides (Diptera: Ceratopogonidae). Proceedings of the Biological Society of Washington, 61, 21–28.

    Fox, I. (1952) Six new Neotropical species of Culicoides (Diptera: Ceratopogonidae). Annals of the Entomological Society of America, 45, 364–368.
    http://dx.doi.org/10.1093/aesa/45.3.364

    Fox, I. (1955) A catalogue of the bloodsucking midges of the Americas (Culicoides, Leptoconops, and Lasiohelea) with keys to the subgenera and Nearctic species, a geographic index, and bibliography. Journal of Agriculture of the University of Puerto Rico, 39, 214–285.

    Fox, I. & Hoffman, W.A. (1944) New Neotropical biting sand flies of the genus Culicoides (Diptera: Ceratopogonidae). Puerto Rico Journal of Public Health and Tropical Medicine, 20, 108–111.

    Garrett, C.B.D. (1925) Seventy new Diptera. Cranbrook, British Columbia, 16 pp.

    Goeldi, E. (1905) Os mosquitos no Pará. Reunião de quatro trabalhos sobre os mosquitos indígenas, principalmente as espécies que molestam o homem. Memórias do Museu Goeldi (Museu Paraense) de Historia Natural e Ethnographie, 4, 1–154, pls. 1–5.

    Grogan, W.L. & Wirth, W.W. (1980) Nannohelea, a new genus of biting midges of the tribe Ceratopogonini, related to Baeohelea Wirth and Blanton (Diptera: Ceratopogonidae). Journal of Kansas Entomological Society, 53, 373–385.

    Grogan, W.L. & Wirth, W.W. (1981) A new American genus of predaceous midges related to Palpomyia and Bezzia (Diptera: Ceratopogonidae). Proceedings of the Biological Society of Washington, 94, 1279–1305.

    Haeselbarth, E. (1965) Phaenobezzia, a new genus of biting midges (Diptera: Ceratopogonidae), with a review of the African species. Zeitschrift für Angewandte Zoologie, 52, 297–324.

    Hoffman, W.A. (1925) A review of the species of Culicoides of North and Central America and the West Indies. American Journal of Hygiene, 5, 274–301.

    Hoffman, W.A. (1939) Culicoides filariferus, new species. Puerto Rico Journal of Public Health and Tropical Medicine, 15, 172–176.

    Huerta, H. (1996) Los Ceratopogónidos colectados por Alfonso Dampf en Chiapas, Mexico (Diptera: Ceratopogonidae). Thesis. Universidad Nacional Autónoma de Mexico, Mexico City, xi + 199 pp.

    Huerta, H., Felippe-Bauer, M.L. & Spinelli, G.R. (2012) A new species and new records of Downeshelea Wirth & Grogan in Neotropical México (Diptera: Ceratopogonidae). Zootaxa, 3394, 64–68.

    Iches, L. (1906) Sobre cinco dípteros nuevos del Chaco Austral. Boletín de Ministerio de Agricultura y Ganadería, Argentina, 6, 262–273.

    Ingram, A. & Macfie, J.W.S. (1931) Ceratopogonidae. In: Diptera of Patagonia and South Chile, based mainly on material in the British Museum (Natural History). Part II. Fasc. 4. British Museum (Natural History), London, pp. 155–232.

    Johannsen, O.A. (1938) New species of Nemocera from Puerto Rico. Journal of Agriculture of the University of Puerto Rico, 22, 219–225.

    Karsch, F. (1886) Ein neues märkisches Dipteron (Ceratopogon crudelis n. sp.). Berliner Entomologische Zeitschrift, 30, xvii–xviii.

    Kieffer, J.J. (1899) Description d'un nouveau genre et tableau des genres européens de la famille des Chironomides (Dipt.). Bulletin de la Société Entomologique de France, 1899, 66–70.

    Kieffer, J.J. (1906) Diptera. Fam. Chironomidae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. Bruxelles 42, pp. 1–78 pp., pls. 1–4.

    Kieffer, J.J. (1911a) Description de nouveaux chironomides de l'Indian Museum de Calcutta. Records of the Indian Museum, 6, 113–177, pls. 6-7.

    Kieffer, J.J. (1911b) Les chironomides de Tendipedidae de l'Himalaya et d'Assam. Records of the Indian Museum, 6, 319–349.

    Kieffer, J.J. (1911c) Nouvelles descriptions de chironomides obtenus d'éclosion. Bulletin de la Société d'Histoire Naturelle de Metz, 27, 1–60. [Jan]

    Kieffer, J.J. (1913) Chironomidae et Cecidomyidae. In: Alluaud, C.A. & Jeannel, R. (Eds.), Voyage de Ch. Alluaud et R. Jeannel en Afrique orientale (1911-1912). Resultats scientifiques. Diptera, 5, pp. 1–43. [Paris]

    Kieffer, J.J. (1917a) Chironomides d'Australie conservés au Musée National Hongrois de Budapest. Annales Historico-Naturales Musei Nationalis Hungarici, 15, 175–228.

    Kieffer, J.J. (1917b) Chironomides d'Amérique conservés au Musée National Hongrois de Budapest. Annales Historico-Naturales Musei Nationalis Hungarici, 15, 292–364.

    Kieffer, J.J. (1919) Observations sur les chironomides (Dipt.) décrits par J.R. Malloch. Bulletin de la Société Entomologique de France, 1919, 191–194.

    Kieffer, J.J. (1921a) Chironomides de l'Afrique Équatoriale. Annales de la Société Entomologique de France, 90, 1–56.

    Kieffer, J.J. (1921b) Sur quelques Diptères piqueurs de la tribu des Ceratopogoninae. Archives de l'Institut Pasteur de l'Afrique du Nord, 1, 107–115. [March]

    Knab, F. (1914) Ceratopogoninae sucking the blood of caterpillars. Proceedings of the Entomological Society of Washington, 16, 63–66.

    Lane, J. (1944) As espécies neotropicais do gênero Clinohelea Kieffer, 1917 (Diptera, Ceratopogonidae). Revista de Entomologia, 15, 249–261.

    Lane, J. (1946) New Neotropical Ceratopogonidae (Heleidae). (Diptera, Nemocera). Revista de Entomologia, 17, 202–215.

    Lane, J. (1948) Novas Ceratopogonídeos do Brasil (Diptera, Ceratopogonidae) (Heleidae). Arquivos da Faculdade de Higiene e Saúde Pública da Universidade de São Paulo, 1, 225–239, figs. 1–8.

    Lane, J. (1956a) On "Paryphoconus" and Stenoxenus (Diptera, Ceratopogonidae). Revista Brasileira de Biologia, 16, 299–308.

    Lane, J. (1956b) The genus "Macfiehelea" Lane, 1946 (Diptera, Ceratopogonidae). Revista Brasileira de Biologia, 16, 435–438.

    Lane, J. (1958) On Neotropical Bezzia. Revista Brasileira de Entomologia, 8, 25–36.

    Lane, J. (1960) Additional data on "Palpomyia" (Diptera, Ceratopogonidae). Revista Brasileira de Biologia, 20, 381–389.

    Lane, J. & Duret, J.P. (1954) Clinohelea da Argentina e do Brasil (Diptera, Ceratopogonidae). Dusenia, 5, 247–253.

    Lane, J. & Forattini, O.P. (1958) Neotropical Stilobezzia II. Fourteen new species, chiefly from Panama (Diptera, Ceratopogonidae). Revista Brasileira de Entomologia, 8, 203–224.

    Lane, J. & Forattini, O.P. (1961) Neotropical Stilobezzia, 1911 III. Key for the the adults of this genus and description of one new species (Diptera, Ceratopogonidae). Revista Brasileira de Entomologia, 10, 83–94.

    Lane, J. & Wirth, W.W. (1964) The biting midge genus Monohelea Kieffer in the Neotropical region (Diptera, Ceratopogonidae). Studia Entomologica, 7, 209–236.

    Latreille, P.A. (1809) Genera crustaceorum et insectorum secundum ordinem naturalem in familias disposita, iconibus exemplisque plurimis explicata. Vol. 4. Paris and Strasbourg, 399 pp., 4 pls.

    Lee, V.H. (1968) Parthenogenesis and autogeny in Culicoides bambusicola Lutz (Ceratopogonidae, Diptera). Journal of Medical Entomology, 5, 91–93.

    Lenz, F. (1934) 13a. Heleidae (Ceratopogonidae). In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region. 3 (Lfg. 78). Schweizerbart, Stuttgart, pp. 95–133.

    Loew, H. (1861) Diptera Americae septentrionalis indigena. Berliner Entomologische Zeitschrift, 5, 307–359.

    Loew, H. (1866) Diptera Americae septentrionalis indigena. Berliner Entomologische Zeitschrift, 9, 127–186.
    http://dx.doi.org/10.1002/mmnd.18650090303

    Lutz, A. (1913) Contribuição para o estudo das Ceratopogoninas hematófagas do Brazil. Parte sistemática. Segunda Memória. Memórias do Instituto Oswaldo Cruz, 5, 45–73, pls. 6–8.
    http://dx.doi.org/10.1590/s0074-02761913000100005

    Lutz, A. (1914) Contribuição para o conhecimento das "Ceratopogoninas" do Brasil. Tercera memória. Aditamento terceiro e descrição de espécies que não sugam sangue. Memórias do Instituto Oswaldo Cruz, 6, 81–99, pls. 8–9.

    Macfie, J.W.S. (1932a) Ceratopogonidae from the wings of dragonflies. Tijdschrift voor Entomologie, 75, 265–283.

    Macfie, J.W.S. (1932b) Some new or little known Ceratopogonidae. Annual Magazine of Natural History, 9, 485–499.
    http://dx.doi.org/10.1080/00222933208673523

    Macfie, J.W.S. (1934) A new British Forcipomyia (Diptera, Ceratopogonidae). Entomologist's Monthly Magazine, 70, 144. [June]

    Macfie, J.W.S. (1935) Ceratopogonidae (Dipt.) from the river Amazon. Stylops, 4, 49–56.
    http://dx.doi.org/10.1111/j.1365-3113.1935.tb00555.x

    Macfie, J.W.S. (1937) Notes on Ceratopogonidae (Diptera). Proceedings of the Royal Entomological Society of London (B), 6, 111–118.
    http://dx.doi.org/10.1111/j.1365-3113.1937.tb00451.x

    Macfie, J.W.S. (1938) Notes on Ceratopogonidae (Diptera). Proceedings of the Royal Entomological Society of London (B), 7, 157–166.
    http://dx.doi.org/10.1111/j.1365-3113.1938.tb01271.x

    Macfie, J.W.S. (1939a) 5. Ceratopogonidae. In: Ruwenzori Expedition 1934-5. 1 (5). British Mus. (Nat. Hist.), London, pp. 81–102.

    Macfie, J.W.S. (1939b) A report on a collection of Brazilian Ceratopogonidae. Revista de Entomologia, 10, 137–219.

    Macfie, J.W.S. (1940a) Ceratopogonidae (Diptera) from north-eastern Brazil. Proceedings of the Royal Entomological Society of London (B), 9, 73–79.

    Macfie, J.W.S. (1940b) Ceratopogonidae (Diptera) from British Guiana and Tinidad. Part 2. Proceedings of the Royal Entomological Society of London (B), 9, 187–195.

    Macfie, J.W.S. (1940c) A report on a collection of Ceratopogonidae (Diptera) from British Guiana. Entomologist's Monthly Magazine, 76, 23–32.

    Macfie, J.W.S. (1940d) The genera of Ceratopogonidae. Annals of Tropical Medicine and Parasitology, 34, 13–30.

    Macfie, J.W.S. (1948) Some species of Culicoides (Diptera, Ceratopogonidae) from the state of Chiapas, Mexico. Annals of Tropical Medicine and Parasitology, 42, 67–87.

    Macfie, J.W.S. (1949) Notes on Ceratopogonidae. Proceedings of the Royal Entomological Society of London (B), 18, 109–115.
    http://dx.doi.org/10.1111/j.1365-3113.1949.tb01431.x

    Malloch, J.R. (1915a) The Chironomidae, or midges, of Illinois, with particular reference to the species occurring in the Illinois River. Bulletin of the Illinois State Laboratory of Natural History, 10, 275–543, 24 pls.

    Malloch, J.R. (1915b) Some additional records of Chironomidae for Illinois and notes on other Illinois Diptera. Bulletin of the Illinois State Laboratory of Natural History, 11, 305–363, pls. 80–84.

    Marinkelle, C.J. & Germán, E. (1970) Mansonelliasis in the Comisaria del Vaupés. Tropical and Geographical Medicine, 22, 101–111.

    Marino, P.I. & Spinelli, G.R. (1999) The species groups of the subgenus Forcipomyia (Forcipomyia) in the Neotropics, with a description of a new species of the genualis group (Diptera: Ceratopogonidae). Transactions of the American Entomological Society, 125, 445–452. [Dec.]

    Marino, P.I. & Spinelli, G.R. (2001) Las especies del subgénero Forcipomyia (Forcipomyia) en la Patagonia (Diptera, Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 60, 99–124.

    Marino, P.I. & Spinelli, G.R. (2002) A revision of the Forcipomyia squamitibia group in the Neotropics with the description of three new species (Diptera: Ceratopogonidae). Insect Science and its Application, 22, 307–319.
    http://dx.doi.org/10.1017/s1742758400020944

    Mayer, K. (1937) Beitrag zur Kenntnis der deutschen Ceratopogoniden. (Dipt.) IX. Stettiner Entomologische Zeitung , 98, 301–304.

    Meigen, J.G. (1818) Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten. Vol. 1. Bei Friedrich Wilhelm Forstmann, Aache, xxxvi + 333 pp., pls. 1–11.

    Messersmith, D.H. (1972) A new species of Culicoides from Colombia (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington, 74, 165–169.

    Moncada, L., Carrasquiila, M.C., Spinelli, G.R., Lotta, I. & Matta, N. (2010) Description of Culicoides lisicarruni (Diptera: Ceratopogonidae), a new species from Cundinamarca, Colombia. Memórias do Instituto Oswaldo Cruz, 105, 978–980.
    http://dx.doi.org/10.1590/S0074-02762010000800005

    Newman, E. (1834) Attempted division of British insects into natural orders. Entomological Magazine, 2, 379–431.

    Noè, G. (1905) Un nuovo genere appartenente alla famiglia Chironomidae. Rendiconti, Accademia Nazionale dei Lincei, 14, 114–120.

    Noè, G. (1907) Due nuove specie di ditteri appartenenti ad un genere nuovo. Archivio Zoologico Italiano; Pubblicato Sotto gli Auspicii Della Unione Zoologica, 3, 101–164, pls. 1–3.

    Ortíz, I. (1950a) Estudios en Culicoides IV. Revisión de las especies americanas del sub-género Hoffmania Fox 1948, con la descripción de dos nuevas especies. Revista de Sanidad y Asistencia Social, 15, 437–460.

    Ortíz, I. (1950b) Estudios en Culicoides V. Informes sobre una nueva especie y lista de los machos cuyas genitalias son conocidas. Revista de Sanidad y Asistencia Social, 15, 461–465.

    Ortíz, I. (1950c) Descripción de Stilobezzia glauca venezuelensis n. vr. y Monohelea mayeri n. sp. Memórias de la Sociedad de Ciencias Naturales La Salle, 10, 199–204.

    Ortíz, I. (1951) Estudios en Culicoides (Diptera, Ceratopogonidae) 9. Sobre los caracteres diferenciales entre Culicoides paraensis (Goeldi, 1905), C. stellifer (Coquillett, 1901), y C. lanei (Ortíz, 1950). Descipción de cuatro nuevas especies con la redescripción de algunas otras poco conocidas. Revista de Sanidad y Asistencia Social, 16, 573–591.

    Ortíz, I. (1953) Nueva contribución al conocimiento de los caracteres morfológicos externos de las hembras americanas del género Culicoides Ltr. (Diptera, Ceratopogonidae) con una espermateca. Descripción de dos nuevas especies de Venezuela: Culicoides transferrans (C. oublepharus [sic] Ortíz, 1952) y Culicoides mirsae. Revista de Sanidad y Asistencia Social, 18, 797–806.

    Ortíz, I. (1954) Sobre dos nuevos dípteros hematófagos del género Culicoides (Nematocera, Ceratopogonidae). Archivos Venezolanos de Patología Tropical y Parasitología Médica, 2, 221–226.

    Ortíz, I. (1956) Sobre un pequeño díptero de la región del Auyantepuy. Boletín Venezolano de Laboratorio Clínico, 1, 93–96.

    Ortíz, I. & León, L.A. (1955) Los Culicoides (Diptera: Ceratopogonidae) de la República del Ecuador. Boletín de Informaciones Cientificas Nacionales, 7, 564–594. [January–February]

    Ortíz, I. & Mirsa, M. (1950) Estudios en Culicoides (Diptera, Ceratopogonidae) VIII.- Sobre una nueva especie del grupo haematopotus: Culicoides venezuelensis n. sp. Archivos Venezolanos de Patología Tropical y Parasitología Médica, 2, 137–143.

    Ortíz, I. & Mirsa, M. (1952a) Culicoides de Venezuela - Redescripción de 10 especies con la descripción de algunos sexos no conocidos. Revista de Sanidad y Asistencia Social, 17, 257–279.

    Ortíz, I. & Mirsa, M. (1952b) Sobre las especies Americanas del género "Culicoides" Latr., (Diptera, “Ceratopogonidae”) con una espermateca. Acta Científica Venezolana, 2, 125–128.

    Philippi, R.A. (1865) Aufzahlung der chilenischen Dipteren. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft, 15, 595–782. [in Wien]

    Poey, F. (1853) Memórias sobre la historia natural de la Isla de Cuba, acompañadas de sumarios latinos y extractos en francés. Vol. 1. pt. 4. Havanna, 80 pp. [pp. 201–280]

    Rafinesque, C.S. (1815) Analyse de la nature ou tableau de l'univers et des corps organisés. Le nature est mon guide, et Linnéus mon maitre. Privately published, Palmero, 224 pp.

    Rodríguez, M.C. de & Wirth, W.W. (1986) A new species of man-biting Culicoides from the high Andes of Colombia (Diptera: Ceratopogonidae). Florida Entomologist, 69, 311–314.
    http://dx.doi.org/10.2307/3494934

    Ronderos, M.M. (1988) Estudio sistemático de los géneros hematófagos de Ceratopogonidae (Diptera: Nematocera) de la República Argentina. Ph.D. Thesis, Univ. Nac. La Plata, Argentina, 135 pp. + 49 pls.

    Ronderos, M.M. & Spinelli, G.R. (1992) A key to the Neotropical Leptoconops (Diptera: Ceratopogonidae), with the description of a new species from Argentina. Revista de la Sociedad Entomológica Argentina, 51, 41–45.

    Ronderos, M.M. & Spinelli, G.R. (1995) Redescripción de Culicoides lahillei y descripción de Culicoides ichesi, n. sp. de la Argentina, Paraguay y Uruguay (Diptera: Ceratopogonidae). Neotrópica, 41, 77–81.

    Ronderos, M.M. & Spinelli, G.R. (1998) Las especies de Culicoides en el área de influencia de la represa de Yacyretá (Diptera: Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 57, 79–83.

    Ronderos, M.M. & Spinelli, G.R. (1999) On the subgenus Forcipomyia (Lasiohelea) in the Neotropical Region (Diptera: Ceratopogonidae). Transactions of the American Entomological Society, 125, 151–161. [June]

    Santarem A.M.N., Lopes Da Trinidade R., Do Nascimiento Da Silva, T., Castellon, E.G., Antunes de Melo-Patiú. C. & Felippe-Bauer, M.L. (2014) New neotropical Culicoides and redescription of Culicoides reticulatus Lutz (Diptera: Ceratopogonidae). Zootaxa, 3795 (3), 255–274.
    http://dx.doi.org/10.11646/zootaxa.3795.3.2

    Saunders, L.G. (1925) On the life history, morphology and systematic position of Apelma Kieff. and Thyridomyia n. g. (Diptera, Nemat. Ceratopogoninae). Parasitology, 17, 252–277.
    http://dx.doi.org/10.1017/S0031182000004704

    Saunders, L.G. (1957) Revision of the genus Forcipomyia based on characters of all stages (Diptera, Ceratopogonidae). Canadian Journal of Zoology, 34, 657–705. [1956]
    http://dx.doi.org/10.1139/z56-065

    Saunders, L.G. (1959) Methods for studying Forcipomyia midges, with special reference to cacao-pollinating species (Diptera, Ceratopogonidae). Canadian Journal of Zoology, 37, 3–51.
    http://dx.doi.org/10.1139/z59-005

    Say, T. (1825) American entomology, or descriptions of the insects of North America. Vol. 2. S. A. Mitchell, Philadelphia, 121 pp., 18 pls. [pls. 19–36]

    Skuse, F.A.A. (1889) Diptera of Australia. Part VI. - The Chironomidae. Proceedings of the Linnean Society of New South Wales, 4, 215–311, pls. 11–14.

    Soria, S. & Bystrak, P.G. (1975) A new species of Forcipomyia (Diptera, Ceratopogonidae) described in all stages, with an account of its role as a cacao pollinator. Revista Theobroma, 5, 3–11.

    Soria, S. & Wirth, W.W. (1979) Ceratopogonid midges (Diptera: Nematocera) collected from cacao flowers in Palmira, Colombia: an account of their pollinating abilities. Revista Theobroma, 9, 77–84.

    Spinelli, G.R. & Duret, J.P. (1993) Las especies neotropicales de Clinohelea (Diptera: Ceratopogonidae). Graellsia, 49, 39–50.

    Spinelli, G.R., Greiner, E.C. & Wirth, W.W. (1993) The Neotropical bloodsucking midges of the Culicoides guttatus group of the subgenus Hoffmania (Diptera: Ceratopogonidae). Contributions of the American Entomological Insitute, 27, 1–91.

    Spinelli, G.R. & Grogan, W.L. (1987) A revision of the Neotropical species of Parabezzia (Diptera: Ceratopogonidae). Biología Acuática, 11, 1–45.

    Spinelli, G.R. & Grogan, W.L. (1989) Dos especies nuevas de Palpomyia, grupo distincta de la región Neotropical (Diptera: Ceratopogonidae). Neotrópica, 35, 3–8.

    Spinelli, G.R. & Grogan, W.L. (1994) Two new Neotropical species of the subgenus Isohelea of Brachypogon, with the description of the female of B. (I.) misionensis and a key to the Neotropical species (Diptera: Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 53, 1–8.

    Spinelli, G.R. & Grogan, W.L. (1998) A revision of the Neotropical predaceous midges of Brachypogon (Brachypogon) Kieffer (Diptera: Ceratopogonidae). Insecta Mundi, 12, 59–79.

    Spinelli, G.R., Ronderos, M.M., Díaz, F. & Marino, P.I. (2005) The bloodsucking biting midges of Argentina (Diptera: Ceratopogonidae). Memórias do Instituto Oswaldo Cruz, 100, 137–150.
    http://dx.doi.org/10.1590/S0074-02762005000200006

    Spinelli, G.R., Santamaría, E., Cabrera, O.L., Ronderos, M.M. & Suárez, M.F. (2009) Five new species of Culicoides Latreille described from Colombia, yielding a new species list and country records (Diptera: Ceratopogonidae). Memórias do Instituto Oswaldo Cruz, 104, 81–92.
    http://dx.doi.org/10.1590/S0074-02762009000100013

    Spinelli, G.R. & Wirth, W.W. (1984a) A review of the Neotropical predaceous midge genus Paryphoconus (Diptera: Ceratopogonidae). Proceedings of the Biological Society of Washington, 97, 882–908.

    Spinelli, G.R. & Wirth, W.W. (1984b) The Neotropical predaceous midges of the genus Alluaudomyia (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington, 86, 673–702.

    Spinelli, G.R. & Wirth, W.W. (1984c) Las especies Neotropicales del género Dasyhelea, grupo cincta (Diptera: Ceratopogonidae). Limnobios, 2, 586–608.

    Spinelli, G.R. & Wirth, W.W. (1986a) The Neotropical species of Phaenobezzia (Diptera: Ceratopogonidae). Florida. Entomologist, 69, 231–236.
    http://dx.doi.org/10.2307/3494765

    Spinelli, G.R. & Wirth, W.W. (1986b) Clave para la identificación de las especies del género Culicoides Latreille presentes al sur de la cuenca Amazónica. Nuevas citas y notas sinonímicas (Diptera: Ceratopogonidae). Revista de la Sociedad Entomológica Argentina, 44, 49–73.

    Spinelli, G.R. & Wirth, W.W. (1989a) Las especies Neotropicales del género Bezzia (Diptera: Ceratopogonidae). II. Los grupos nobilis y punctipennis. Revista de la Sociedad Entomológica Argentina, 45, 109–129.

    Spinelli, G.R. & Wirth, W.W. (1989b) The Neotropical predaceous midges of the genus Bezzia (Diptera: Ceratopogonidae) Part I. The glabra and brevicornis groups. Limnobios, 2, 762–778.

    Spinelli, G.R. & Wirth, W.W. (1990) Neotropical predaceous midges of the genus Bezzia (Diptera: Ceratopogonidae) Part III. The gibbera group of species. Insecta Mundi, 4, 11–32.

    Spinelli, G.R. & Wirth, W.W. (1991) The Neotropical predaceous midges of the genus Bezzia (Diptera: Ceratopogonidae) Part IV. The dentifemur and venustula groups. Insecta Mundi, 5, 1–17.

    Spinelli, G.R. & Wirth, W.W. (1993) Los Ceratopogonidae de la Argentina (Insect: Diptera). In: Castellanos, Z. (Ed.), Fauna de agua dulce de la República Argentina. Volumen 38. Diptera Fascículo 3. Ceratopogonidae. Profadu-Conicet, Buenos Aires, 124 pp.

    Staeger, R.C. (1839) Systematisk fortegnelse over de i Danmark hidtil fundne Diptera. Naturhistorisk Tidsskrift, 2, 549–600.

    Tavares, O. & Souza, M.A. de (1979) Sobre algumas espécies do gênero Culicoides Latreille, 1809 do Estado do Rio de Janeiro, Brasil (Diptera, Ceratopogonidae). Revista Brasileira de Biologia, 39, 611–613.

    Tidwell, M.A. & Tidwell, M. (1982) Development of Mansonella ozzardi in Simulium amazonicum, S. argentiscutum and Culicoides insinuatus. American Journal of Tropical Medicine and Hygiene, 31, 1137–1141.

    Tidwell, M.A., Tidwell, M., Hoyos, P.M., Corredor, A. & Barreto, P. (1980) Vectores de Onchocerca volvulus y Mansonella ozzardi en Colombia. Acta Médica del Valle, 11, 119–30.

    Townsend, C.H.T. (1893) An interesting blood-sucking gnat of the family Chironomidae. Psyche, 6, 369–371.
    http://dx.doi.org/10.1155/1893/58983

    Utmar, J.A. & Wirth, W.W. (1976) A revision of the new world species of Forcipomyia, subgenus Caloforcipomyia (Diptera: Ceratopogonidae). Florida Entomologist, 59, 109–133.
    http://dx.doi.org/10.2307/3493959

    Vargas, L. (1954) Dos nuevas especies de Culicoides Mexicanos (Diptera, Heleidae). Revista del Instituto de Salubridad y Enfermedades Tropicales, 14, 25–32.

    Vargas, L. (1960) The subgenera of Culicoides of the Americas (Diptera, Ceratopogonidae). Revista de Biologia Tropical, 8, 35–47.

    Vitale, G. C., Wirth, W.W. & Aitken, T.H.G. (1981) New species and records of Culicoides reared from arboreal habitats in Panama, with a synopsis of the debilipalpis group (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington, 83, 140–159.

    Walker, F. (1856) Diptera. In: Saunders, W.W. (Ed.), Insecta Saundersiana: Or characters of undescribed insects in the collection of William Wilson Saunders. Vol. 1. London, pp. 415–474.

    Westwood, J.O. (1840) Order XIII. Diptera Aristotle (Antliata Fabricius. Halteriptera Clairv.). In: Westwood, J.O. (Ed.), An introduction to the modern classification of insects; founded on the natural habits and corresponding organisation of the different families. Vol. 2. Synopsis of the genera of British insects. Longman, Orme, Brown, Green, London, pp. 125–154.

    Williston, S.W. (1896) On the Diptera of St. Vincent (West Indies). Transactions of the Entomological Society of London, 1896, 253–446, pls. 8–14.

    Williston, S.W. (1900) Supplement (Part). In: Godman, F.D. & Salvin, O. (Eds.), Biologia Centrali-Americana. Zoologia-Insecta-Diptera. Vol. 1. Taylor & Francis, London, pp. 217–248.

    Wirth, W.W. (1951a) A new biting midge of the genus Leptoconops from Florida, with new records of other American species (Diptera, Heleidae). Proceedings of the Entomological Society of Washington, 53, 281–284.

    Wirth, W.W. (1951b) New species and records of Virginia Heleidae (Diptera). Proceedings of the Entomological Society of Washington, 53, 313–326.

    Wirth, W.W. (1952) Two new species of anthropophilic Culicoides from Guatemala (Diptera: Heleidae). Journal of Parasitology, 38, 245–247.
    http://dx.doi.org/10.2307/3274042

    Wirth, W.W. (1953) American biting midges of the heleid genus Monohelea. Proceedings of the United States National Museum, 103, 135–154.
    http://dx.doi.org/10.5479/si.00963801.103-3320.135

    Wirth, W.W. (1959) Pachyhelea, a new genus of American Ceratopogonidae related to Palpomyia (Diptera). Bulletin of the Brooklyn Entomological Society, 54, 50–52.

    Wirth, W.W. (1960) The genus Pellucidomyia Macfie (Diptera, Ceratopogonidae). Bulletin of the Brooklyn Entomological Society, 55, 1–3.

    Wirth, W.W. (1962) A reclassification of the Palpomyia-Bezzia-Macropeza groups, and a revision of the North American Sphaeromiini (Diptera, Ceratopogonidae). Annals of the Entomological Society of America, 55, 272–287.
    http://dx.doi.org/10.1093/aesa/55.3.272

    Wirth, W.W. (1965) A revision of the genus Parabezzia Malloch (Diptera, Ceratopogonidae). Proceedings of the Entomological Society of Washington, 67, 215–230.

    Wirth, W.W. (1970) The neotropical Forcipomyia midges of the subgenus Thyridomyia Saunders (Diptera, Ceratopogonidae). Studia Entomologica, 13, 429–440.

    Wirth, W.W. (1971) A review of the "stick-ticks", neotropical biting midges of the Forcipomyia subgenus Microhelea parasitic on walking stick insects (Diptera: Ceratopogonidae). Entomological News, 82, 229–245.

    Wirth, W.W. (1972) The neotropical Forcipomyia (Microhelea) species related to the caterpillar parasite F. fuliginosa (Diptera: Ceratopogonidae). Annals of the Entomological Society of America, 65, 564–577.
    http://dx.doi.org/10.1093/aesa/65.3.564

    Wirth, W.W. (1974) 14. Family Ceratopogonidae. In: Papavero, N. (Ed.), A catalog of the diptera of the Americas south of the United States. Museu de Zoologia, São Paulo, 89 pp.

    Wirth, W.W. (1976) Forcipomyia pictoni Macfie and descriptions of two new related species from Florida (Diptera: Ceratopogonidae). Florida Entomologist, 59, 77–84.
    http://dx.doi.org/10.2307/3493176

    Wirth, W.W. (1978) New species and records of intertidal biting midges of the genus Dasyhelea Kieffer from the Gulf of California (Diptera: Ceratopogonidae). Pacific Insects, 18, 191–198.

    Wirth, W.W. (1982) The cacao-pollinating midges of the Forcipomyia argenteola group (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington, 84, 568–585.

    Wirth, W.W. (1991a) The predaceous midge genus Allohelea Kieffer in the western hemisphere (Diptera: Ceratopogonidae). Florida Entomologist, 74, 491–505.
    http://dx.doi.org/10.2307/3495403

    Wirth, W.W. (1991b) New and little-known species of Forcipomyia (Diptera: Ceratopogonidae) associated with cocoa pollination in Brazil. Proceedings of the Entomological Society of Washington, 93, 163–175.

    Wirth, W.W. (1991c) Forcipomyia bicolor and related species of the subgenus Lepidohelea in Brazil (Diptera: Ceratopogonidae). Florida Entomologist, 74, 506–517.
    http://dx.doi.org/10.2307/3495404

    Wirth, W.W. (1994) The western hemisphere species of the predaceous midge genus Echinohelea, with descriptions of six new species (Diptera: Ceratopogonidae). Insecta Mundi, 8, 227–242.

    Wirth, W.W. & Atchley, W.R. (1973) A review of the North American Leptoconops (Diptera: Ceratopogonidae). Graduate Studies Texas Tech University, 5, 1–57.

    Wirth, W.W. & Barreto, P. (1978) New species of Culicoides biting midges (Diptera: Ceratopogonidae) from Colombia. Journal of Medical Entomology, 14, 553–564.
    http://dx.doi.org/10.1093/jmedent/14.5.553

    Wirth, W.W. & Blanton, F.S. (1953a) Studies in Panama Culicoides (Diptera: Heleidae): I, Descriptions of six new species. Journal of the Washington Academy of Sciences, 43, 69–77.
    http://dx.doi.org/10.2307/3273942

    Wirth, W.W. & Blanton, F.S. (1953b) Studies in Panama Culicoides (Diptera, Heleidae). II. Descriptions of six additional new species. Journal of Parasitology, 39, 229–236.
    http://dx.doi.org/10.2307/3273942

    Wirth, W.W. & Blanton, F.S. (1955a) Studies in Panama Culicoides (Diptera, Heleidae) IV. Descriptions of three new species. Bulletin of the Brooklyn Entomological Society, 50, 100–106. [October]

    Wirth, W.W. & Blanton, F.S. (1955b) Studies in Panama Culicoides (Diptera, Heleidae) V. Descriptions of three new species of the subgenus Oecacta Poey. Bulletin of the Brooklyn Entomological Society, 50, 121–127.

    Wirth, W.W. & Blanton, F.S. (1956a) A new species of salt-marsh sand fly from Florida, the Bahamas, Panama and Ecuador: its distribution and taxonomic differentiation from Culicoides furens (Poey) (Diptera, Heleidae). Florida Entomologist, 39, 157–162.
    http://dx.doi.org/10.2307/3492592

    Wirth, W.W. & Blanton, F.S. (1956b) Studies in Panama Culicoides VII. The species of the pulicaris and cova-garciai group (Diptera, Heleidae). Proceedings of the Entomological Society of Washington, 58, 211–227.

    Wirth, W.W. & Blanton, F.S. (1956c) Studies in Panama Culicoides (Diptera, Heleidae). IX. Two new species related to leoni Barbosa and reevesi Wirth. Bulletin of the Brooklyn Entomological Society, 51, 45–52.

    Wirth, W.W. & Blanton, F.S. (1959) Biting midges of the genus Culicoides from Panama (Diptera: Heleidae). Proceedings of the United States National Museum, 109, 237–482.
    http://dx.doi.org/10.5479/si.00963801.109-3415.237

    Wirth, W.W. & Blanton, F.S. (1968) A revision of the neotropical biting midges of the hylas group of Culicoides (Diptera: Ceratopogonidae). Florida Entomologist, 51, 201–215.
    http://dx.doi.org/10.2307/3493420

    Wirth, W.W. & Blanton, F.S. (1970a) New genera of Neotropical Ceratopogonidae (Diptera). Florida Entomologist, 53, 7–14.
    http://dx.doi.org/10.2307/3493107

    Wirth, W.W. & Blanton, F.S. (1970b) Notes on Brachypogon Kieffer (Diptera, Ceratopogonidae), a new species, and two new Neotropical genera of the tribe Ceratopogonini. Florida Entomologist, 53, 93–104.
    http://dx.doi.org/10.2307/3493452

    Wirth, W.W. & Blanton, F.S. (1970c) A review of the Culicoides nigrigenus group, with two new species (Diptera: Ceratopogonidae). Entomological News, 81, 141–151.

    Wirth, W.W. & Blanton, F.S. (1972) Lanehelea, a new neotropical genus of Sphaeromiini (Diptera: Ceratopogonidae). Studia Entomologica, 15, 433–438.

    Wirth, W.W. & Blanton, F.S. (1973) A review of the maruins or biting midges of the genus Culicoides (Diptera: Ceratopogonidae) in the Amazon Basin. Amazoniana, 4, 405–470.

    Wirth, W.W. & Blanton, F.S. (1974) The West Indian sandflies of the genus Culicoides (Diptera: Ceratopogonidae). Agricultural Research Service, United States Department of Agriculture, Technical Bulletin, 1474, iv + 98 pp.

    Wirth, W.W. & Dow, M.I. (1972) Studies on the genus Forcipomyia 4. Rhynchoforcipomyia, a new neotropical subgenus in the Trichohelea complex (Diptera: Ceratopogonidae). Annals of the Entomological Society of America, 65, 862–872.
    http://dx.doi.org/10.1093/aesa/65.4.862

    Wirth, W.W., Dyce, A.L. & Spinelli, G.R. (1988) An atlas of wing photographs, with a summary of the numerical characters of the Neotropical species of Culicoides (Diptera: Ceratopogonidae). Contributions of the American Entomological Institute, 25, 1–72

    Wirth, W.W. & Felippe-Bauer, M.L. (1989) The neotropical biting midges related to Culicoides paraensis (Diptera: Ceratopogonidae). Memórias do Instituto Oswaldo Cruz, 84, 551–565.
    http://dx.doi.org/10.1590/S0074-02761989000800096

    Wirth, W.W. & Grogan, W.L. (1988) The predaceous midges of the world (Diptera: Ceratopogonidae; Tribe Ceratopogonini). Flora and Fauna Handbook. 4. E.J. Brill, Leiden, xv + 160 pp.

    Wirth, W.W. & Hubert, A.A. (1960) Ceratopogonidae (Diptera) reared from cacti, with a review of the copiosus group of Culicoides. Annals of the Entomological Society of America, 53, 639–658.
    http://dx.doi.org/10.1093/aesa/53.5.639

    Wirth, W.W. & Lee, V.H. (1967) New species of Culicoides from high altitudes in the Colombian Andes (Diptera: Ceratopogonidae). Proceedings of the United States National Museum 124, 1–22.
    http://dx.doi.org/10.5479/si.00963801.124-3626.1

    Wirth, W.W. & Ratanaworabhan, N.C. (1971) Notes on Neotropical Pellucidomyia (Diptera, Ceratopogonidae). Annals of the Entomological Society of America, 64, 446–448.
    http://dx.doi.org/10.1093/aesa/64.2.446

    Wirth, W.W. & Ratanaworabhan, N.C. (1972a) Neobezzia, a new neotropical biting midge genus of the tribe Sphaeromiini (Diptera: Ceratopogonidae). Journal of the Kansas Entomological Society, 45, 476–490.

    Wirth, W.W. & Ratanaworabhan, N.C. (1972b) A revision of the tribe Stenoxenini (Diptera: Ceratopogonidae). Annals of the Entomological Society of America, 65, 1368–1388.
    http://dx.doi.org/10.1093/aesa/65.6.1368

    Wirth, W.W. & Ratanaworabhan, N.C. (1978) Studies on the genus Forcipomyia. V. Key to subgenera and description of a new subgenus related to Euprojoannisia Brèthes (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington, 80, 493–507.

    Wirth, W.W. & Soria, S.J. (1975) A new Neotropical Forcipomyia midge closely related to F. (F.) genualis (Loew) (Diptera: Ceratopogonidae). Revista Theobroma, 5, 19–27.

    Wirth, W.W. & Soria, S.J. (1980) Studies on the genus Forcipomyia VI. The Neotropical species of the subgenus Warmkea (Diptera: Ceratopogonidae). Revista Theobroma, 9, 137–161.

    Wirth, W.W. & Soria, S.J. (1981) Two Culicoides biting midges reared from inflorescences of Calathea in Brazil and Colombia, and a key to the species of the discrepans group (Diptera: Ceratopogonidae). Revista Theobroma, 11, 107–117.

    Wirth, W.W. & Spinelli, G.R. (1992) Immature stages of Forcipomyia seminole Wirth and a related new neotropical species (Diptera: Ceratopogonidae). Florida Entomologist, 75, 349–356.
    http://dx.doi.org/10.2307/3495856

    Wirth, W.W. & Spinelli, G.R. (1993) The American species of the annulatipes group of the subgenus Lepidohelea, genus Forcipomyia (Diptera: Ceratopogonidae). Insecta Mundi, 6, 109–125. [1992]