Abstract
Five species groups of Dichotomius (Selenocopris) are defined and separated in an illustrated key in the first effective attempt of systematization of the subgenus. Four new brachypterous species of Selenocopris are described and the also brachypterous Dichotomius ingens (Luederwaldt) is redescribed. These species are diagnosed, illustrated and assigned to the quadraticeps species group. Taxonomic issues on the subgenus are discussed.
References
Arias-Buriticá, J.A. & Vaz-de-Mello, F.Z. (2012) Redescripción de Dichotomius camposeabrai y D. nemericola (Coleoptera: Scarabaeidae: Scarabaeinae), con apuntes sobre su posición sistemática. Revista Mexicana de Biodiversidad, 83, 387–395.
Arias-Buriticá, J.A. & Vaz-de-Mello, F.Z. (2013) Dichotomius ribeiroi (Pereira, 1954) (Coleoptera: Scarabaeidae: Scarabaeinae): Redescripción y Anotaciones taxonómicas de la especie Dichotomius ribeiroi (Pereira 1954) (Coleoptera: Scarabaeidae: Scarabaeinae): redescription and taxonomic annotations of the species. Caldasia, 35 (1), 209–217.
Blackwelder, R.E. (1944) Checklist of the Coleopterous insects of Mexico, Central America, the West Indies, and South America. United States National Museum Bulletin, 185, 189–341.
Burmeister, H. (1846) Genera Quaedam Insectorum. Iconibus Illustravit et Descripsit. Vol. I. Continet Tabulas XL. Sumtibus A. Burmeister, Berlim, no pagination.
http://dx.doi.org/10.5962/bhl.title.8144Culot, L., Bovy, E., Vaz-de-Mello, F.Z., Guevara, R. & Galetti, M. (2013) Selective defaunation affects dung beetle communities in continuous Atlantic rainforest. Biological Conservation, 163, 79–89.
http://dx.doi.org/10.1016/j.biocon.2013.04.004Daniel, G.M., Nunes, L.G.O.A. & Vaz-de-Mello, F.Z. (2014) Species composition and functional guilds of dung beetles (Insecta: Coleoptera: Scarabaeidae: Scarabaeinae) in different vegetational types in the Brazilian-Shield-Chacoan Depression Border. Annales de la Société entomologique de France, 50 (2), 183–190.
http://dx.doi.org/10.1080/00379271.2014.938936Darwin, C. (1859) On the Origin of Species by Means of Natural Selection, or the Preservation of Favored Races in the Struggle for Life. John Murray, London, 502 pp.
Dinghi, P., Sánchez, M.V., Cantil, L.F., Sarzetti, L.C. & Genise, J.F. (2013) Leaf-Litter Brood Chambers in Dichotomius (Luederwaldtinia) Carbonarius (Mannerheim, 1829) (Coleoptera: Scarabaeidae): A Novel Behaviour for Dung Beetles. The Coleopterists Bulletin, 67 (3), 388–396.
http://dx.doi.org/10.1649/0010-065X-67.3.388Felsche, C. (1901) Beschereibungen coprophage Scarabaeiden. Deutsche Entomologische Zeitschrift, 2, 135–155.
Génier, F. (2000) Dichotomius comarapensis sp. nov., une nouvelle espèce bolivienne de scarabée brachyptère (Coleoptera: Scarabaeidae: Scarabaeinae). Fabreries, 25, 25–31.
Germar, E.F. (1824) Coleopterum species novae aut minus cognitae, descriptionibus illustratae. Imprensis J. C. Hendelli et Filli, Halae, 624 pp.
Harold, E. (1867) Diagnosen neuer Coprophagen. Coleopterologische Hefte, 5, 94–100.
Harold, E. (1869) Révision des espèces qui rentrent dans le genre Pinotus Erichson. L'Abeille 6, 123–144.
Hope, F.W. (1838) The Coleopterists Manual containing the insects of Linneus and Fabricius. Henry G. Bohn, London, 156 pp.
Howden, H. & Young, O.P. (1981) Panamanian Scarabaeinae: Taxonomy, Distributio, and Habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18 (1), 1–204.
Kohlmann, B. & Solís, A. (1997) El género Dichotomius en Costa Rica. Giornale italiano di Entomologia, 8, 343–382.
Köller, W.W., Gomes, A., Rodrigues, S.A. & Goiozo, P.F.I. (2007) Scarabaeidae e Aphodiidae coprófagos em pastagens cultivadas em área do cerrado sul-matogrossense. Revista Brasileira de Zôociências, 9 (1), 81–93.
Louzada, J.N.C., Lopes, F.S. & Vaz-de-Mello, F.Z. (2007) Structure and composition of a dung beetle community (Coleoptera, Scarabaeidae) in a small forest patch from Brazilian Pantanal. Revista Brasileira de Zoociências, 9 (2), 199–203.
Luederwaldt, H. (1929) As espécies Brasileiras do gênero Pinotus. Revista do Museu Paulista, Tomo XVI, 1–173.
Luederwaldt, H. (1931) Novos subsídios para o conhecimento do gênero Pinotus. Revista de Entomologia, 1, 297–304.
Lüderwaldt, H. (1935) Zur kenntnis der gattung Pinotus. Revista de Entomologia, 5 (3), 334–342.
Martínez, A. (1951) La invalidez del nombre genérico Pinotus Erichson y dos nuevas sinonímias (Col. Scarab.). Anales de La Sociedad Científica Argentina, 152, 138–142.
Nunes, R.V., Frizzas, M.R. & Vaz-de-Mello, F.Z. (2012) Scarabaeinae (Coleoptera: Scarabaeidae) of a rupestrian field at Cafuringa, Distrito Federal, Brazil: commented list of species. Biota Neotropica, 12, 125–129.
http://dx.doi.org/10.1590/S1676-06032012000400013Nunes, R.V. & Vaz-de-Mello, F.Z. (2013) New brachypterous species of Dichotomius Hope, with taxonomic notes in the subgenus Luederwaldtinia Martínez (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa, 3609 (4), 411–420.
http://dx.doi.org/10.11646/zootaxa.3609.4.3Pereira, F.S. (1947) Pinotus da secção Batesi. Arquivos do Museu Paranaense, 6, 317–328.
Pereira, P.F.S. & D’Andretta, M.A.V. (1955) Novos Escarabeídeos e Novas Sinonímias (Col. Scarabaeidae). Papeis avulsos do Departamento de Zoologia, 12 (11), 247–264.
Scheffler, P.Y. (2005) Dung beetle (Coleoptera: Scarabaeidae) diversity and community structure across three disturbance regimes in eastern Amazonia. Journal of Tropical Ecology, 21, 9–19.
http://dx.doi.org/10.1017/S0266467404001683Tarasov, S. & Génier, F. (2015) Innovative bayesian and parsimony phylogeny of dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) enhanced by ontology-based partitioning of morphological characters. PLOS ONE, 10 (3), e0116671.
http://dx.doi.org/10.1371/journal.pone.0116671Vaz-de-Mello, F.Z. (1999) Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) de um Fragmento de Floresta Amazônica no Estado do Acre, Brasil. 1. Taxocenose. Anais da Sociedade Entomologica Brasileira, 28 (3), 447–453.
http://dx.doi.org/10.1590/S0301-80591999000300009Vaz-de-Mello, F.Z. (2000) Estado atual de conhecimento dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil. In: Martín-Piera, F., Morrone, J.J. & Melic, A. (Eds.), Hacia un Proyecto CYTED para el Inventario y Estimación de la Diversidad Entomológica en Iberoamérica: PrIBES-2000. Monografias Tercer Milênio, Zaragoza, pp. 183–195.
Vaz-de-Mello, F.Z., Louzada, J.N.C. & Gavino, M. (2001) Nova espécie de Dichotomius Hope, 1838 (Coleoptera, Scarabaeidae) do Espírito Santo, Brasil. Revista Brasileira de Entomologia, 45 (2), 99–102.
Vaz-de-Mello, F.Z., Edmonds, W.D., Ocampo, F.C. & Schoolmesters, P. (2011) A multilingual key to the genera and subgenera of the subfamiliy Scarabaeinae of the new world (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa, 2854, 1–73.
Vernalha, M.M. (1952) Descrição de uma espécie nova do gênero Dichotomius, Hope 1838 (Col. Scarabaeidae). Arquivos de Biologia e Tecnologia, 7 (8), 39–44.