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Type: Monograph
Published: 2016-09-01
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A systematic catalogue of butterflies of the former Soviet Union (Armenia, Azerbaijan, Belarus, Estonia, Georgia, Kyrgyzstan, Kazakhstan, Latvia, Lituania, Moldova, Russia, Tajikistan, Turkmenistan, Ukraine, Uzbekistan) with special account to their type specimens (Lepidoptera: Hesperioidea, Papilionoidea)

Nizhny Novgorod Branch, Russian Entomological Society, P.O.Box 97, Nizhny Novgorod 603009 Russia.
Tula Branch, Russian Entomological Society, Pervomayskaya Str. 12-17 Tula, 300034 Russia.
Lepidoptera butterflies Russia East Europe Caucasus Central Asia catalogue nomenclatural acts name-bearing types distribution

Abstract

A catalogue of butterflies of Russia and adjacent countries is given, with special account to the name-bearing types depository. This catalogue contains data about 86 species (3 of them are questionable) of Hesperiidae (22 genera); 47 species of Papilionidae (14 genera); 89 species of Pieridae (5 of them are questionable)  (15 genera); 1 species (1 genus) of Libytheinae(dae); 2 species of Danainae(dae) (2 genera); 160 species of Nymphalinae(dae) (1 of them is questionable) (23 genera); 259 species of Satyrinae(dae) (14 of them are questionable, mainly from genera Oeneis and Pseudochazara) (34 genera); 3 species of Riodinidae (2 genera); 318 species of Lycaenidae (11 of them are questionable, mainly from genera Neolycaena and Plebeius) (57 genera). In total: 965 species of butterflies, 174 genera, by countries: Armenia—244, Azerbaijan—225, Belarus—107, Estonia—113, Georgia—211, Kyrgyzstan—316, Kazakhstan—344, Latvia—115, Lituania—126, Moldova—87, Russia—522, Tajikistan—295, Turkmenistan—159, Ukraine—192, Uzbekistan—241. Detailed distribution and subspecific structure (if present) for every species is provided. Lectotypes of the following species-group taxa are designated: Hesperia poggei Lederer, 1858, Parnassius felderi Bremer, 1861, P. eversmanni Eversmann, 1851, P. boedromius Püngeler, 1901, Limenitis moltrechti Kardakov, 1928, L. sydyi Kindermann, 1853, L. amphyssa Ménétriès, 1859, L. doerriesi Staudinger, 1892, L. helmanni duplicata Staudinger, 1892, L. homeyeri Tancré, 1881, Argynnis penelope Staudinger, 1891, A. thore borealis Staudinger, 1861, Vanessa io geisha Stichel, [1908], Melitaea maturna staudingeri Wnukowsky, 1929 (=uralensis Staudinger, 1871), M. didymina Staudinger, 1895, Papilio fascelis Esper, 1783, Thecla quercivora Staudinger, 1887, Lycaena orion var. ornata Staudinger, 1892. The following nomenclatural acts are established: Neolycaena submontana baitenovi (Zhdanko, 2011), comb. et stat.n. The following new synonymy is provided: Hesperia comma repugnans (Staudinger, 1892) = lena Korshunov et Gorbunov, 1995, syn.n.; Argynnis niobe orientalis Alphéraky, 1881 =ornata Staudinger, 1901, syn.n. =tanjusha Zhdanko, 2011, syn.n.; Boloria frigga gibsoni (Barnes & Benjamin, 1926) = kosarevi Korb, 2011, syn.n., B. erubescens houri Wyatt, 1961 =ancilla Churkin, 2004, syn.n.; Melitaea fergana maracandica Staudinger, 1882 = irinae Churkin, Kolesnichenko et Tremasov, 2012, syn.n.; M. asteroida clara Staudinger, 1887 =ludmilla Churkin, Kolesnichenko et Tuzov, 2000, syn.n.; Paralasa jordana jordana (Staudinger, 1882) =khramovi Churkin et Pletnev, 2012, syn.n.; P. jordana subocellata (Staudinger, 1901) =kipnisi Churkin et Pletnev, 2012, syn.n.; P. kusnezovi kusnezovi (Avinov, 1910) =bosbutaensis Churkin et Pletnev, 2012, syn.n.; Erebia meta Staudinger, 1886 =gertha Staudinger, 1886, syn.n.; Oeneis ammon ammon Elwes, 1899 =smirnovi Yakovlev, 2011, syn.n.; O. norna tundra A.Bang-Haas, 1912 =ivonini Yakovlev, 2011, syn.n.; Chazara briseis ianthe (Pallas, 1771) =lyrnessus Fruhstorfer, 1908, syn.n., Plebejides stekolnikovi (Stradomsky et Tikhonov, 2015), comb.n.

 

References

  1. Achtelik, G. (1999) Vier neue Arten aus der «Melitaea sultanensis-Gruppe» (Staudinger, 1886) (Lepidoptera, Nymphalidae). Atalanta, 30 (1/4), 63–86.

    Ackery, P.R. (1973) A list of the type-specimens of Parnassius (Lepidoptera: Papilionidae) in the British Museum (Natural History). Bulletin of the British Museum (Natural History), Entomology, 29, 1–35.

    Antonova, E.M. (1981) The type specimens of Lepidoptera, preserved in collections of Zoological Museum of the Moscow University. Sbornik Trudov zoologicheskogo museya MGU, 19, 208–221 [in Russian]

    Asahi, J., Kanda, S., Kawata, M., Kohara, Y. & Fujioka, T. (1999) The butterflies of Sakhalin in nature. Tokyo, 312 pp. [in Japanese]

    Azarova, N.A. (1986) Type material of butterflies of A.I.Kurentzov (Lepidoptera, Rhopalocera) in the collections of Institute of Soil and Biology in Vladivostok. Sistematika I ekologiya cheshuekrylykh Dalnego Vostoka. Nauka Press, Vladivostok, 121–128 [in Russian]

    Back, W. (1990) Taxonomische Untersuchungen innerhalb der Artengruppe um Euchloe ausonia (Hübner, 1804) (Lepidoptera, Pieridae). Atalanta, 21 (3/4), 187–206.

    Back, W. (2012) Phaenotypische und genotypische Abgrenzung der Arten und Unterarten der Gattung Zegris Boisduval, 1836 (Lepidoptera, Pieridae). Atalanta, 43 (1/2), 77–86.

    Back, W. (2013) Verbreitung von Euchloe ausonia (Hübner, 1804), Euchloe daphalis (Moore, 1865) und Euchloe persica Verity, 1908, stat.nov. im Iran (Lepidoptera, Pieridae). Atalanta, 44, 109–117.

    Back, W., Knebelsberger, T. & Miller, M.A. (2006) Molecular investigation of the species and subspecies of the genus Anthocharis Boisduval, Rambur, Duméril & Graslin, 1833 with special focus on the cardamines-group (Lepidoptera: Pieridae). Linneana Belgica, 20 (6), 245–256.

    Back, W., Knebelsberger T. & Miller, M.A. (2008) Molekularbiologische Untersuchungen und Systematik der palaearktischen Arten von Euchloe Hübner, [1819] (Lepidoptera: Pieridae). Entomologische Zeitschrift, 118 (4), 151–169.

    Bálint, Z. (1994) Adalékok a Balkan boglarkalepke-faunajahoz (Lepidoptera, Lycaenidae). A Janus Pannonius Mûzeum Évkônyve, 39, 69–77 [in Hungarian]

    Bálint, Z. (1999) Annotated list of type specimens of Polyommatus sensu Eliot of the Natural History Museum, London (Lepidoptera, Lycaenidae). Neue entomologische Nachrichten, 46, 3–89

    Balint, Z. (2001) Questions and answers concerning Polyommatus (Agrodiaetus) iphigenia Herrich-Schäffer, [1847] (Lepidoptera: Lycaenidae). Linneana Belgica, 18 (2), 73–74.

    Bálint, Z., Kertezs, A. & Lukhtanov, V.A. (1992) A review of blue butterflies of the subgenus Plebejides Sauter, 1968 (Lepidoptera, Lycaenidae). Entomologicheskoe obozreniye, 71 (4), 863–886 [in Russian]

    Bálint, Zs., Guppy, C.S., Kondla, N.G., Johnson, K. & Durden, Ch. J. (2001) Plebeius Kluk, 1780 or Plebejus Kluk, 1780. Folia entomologica Hungarica, 62, 177–184.

    Berg, L.S. (1931) Landscape-geographical zones of USSR. Gosgeografgiz, Moscow-Leningrad, 401 pp. [in Russian]

    Berg, L.S. (1947) Geographical zones of Soviet Union. Geografgiz, Moscow, 397 pp. [in Russian]

    Bernardi, G. (1964) Lépidoptères Lycaenidae (sauf Agrodiaetus) récoltés en Iran par H. de Lesse en 1955 et 1958. Alexanor, 3 (5), 209–216, 273–278.

    Blanchard, E. (1840) Histoire naturelle des insects Orthoptères, Névroptères, Hémiptères, Hyménoptères, Lépidoptères et Diptères. Dumeril, Paris, 3, 672 pp.

    Blume, C., Etzbauer, C., Gaedike, R., Heidenreich, U., Kleikamp, U., Schnug, L., Spiedel, W. & Stüning, D. (2010) Type material of Lepidoptera and Trichoptera in the ZFMK collection, Bonn. Bonner zoologische Bulletin, 58, 169–215.

    Bogdanov, P.V. (2001) A review of the genus Karanasa Moore (Lepidoptera, Satyridae) of Pamir-Alai and Tian-Shan. Trudy Gosudarstvennogo Darvinovskogo Museya, 4, 73–103 [in Russian]

    Bogdanov, P.V. (2003) The type material of Clossiana frigga annae (Suschkin, 1906) (Lepidoptera, Nymphalidae). Trudy Gosudarstvennogo Darvinovskogo Museya, 6, 125–126 [in Russian]

    Bogdanov, P.V. (2004) Classification of the genus Pseudochazara de Lesse, 1951 (Lepidoptera, Satyridae) of Central and Middle Asia. Trudy Gosudarstvennogo Darvinovskogo Museya, 8, 88–181 [in Russian]

    Bogdanov, P.V. (2011) New data on systematics of Karanasa butterflies (Lepidoptera, Satyridae). Trudy Gosudarstvennogo Darvinovskogo Museya, 15, 134–175 [in Russian]

    Bolotov, I.N. (2011) Fauna and ecology of butterflies (Lepidoptera, Rhopalocera) from Kanin peninsula and Kolguev island. Zoologicheskiy zhurnal, 90 (11), 1365–1373 [in Russian]

    Bolshakov, L.V. (2013) Discovery of some Euphydryas Scudder, 1871 species (Lepidoptera: Nymphalidae) in the Alai and Terskei Ala-Too Mountains (Kirgizia). Eversmannia, 35, 50 [in Russian]

    Bolshakov, L.V. & Korb, S.K. (2012) To the systematics and nomenclature of checkerspot butterflies of the group Euphydryas aurinia (Rottemburg, 1775) from Siberia and Central Asia (Lepidoptera: Nymphalidae). Eversmannia, 31-32, 38–48 [in Russian].

    Bolshakov, L.V. & Okulov, V.S. (2014) Оn the fauna of higher Lepidoptera of the Udmurtia. Addition 2. Eversmannia, 37, 27–31 [in Russian]

    Bozano, G.C. (2002) Guide to the butterflies of the palaearctic region. Satyrinae part III. Tribe Satyrini. Subtribes Melanargiina and Coenonymphina. Omnes Artes, Milano, 62 pp.

    Bozano, G.C. (2008) Guide to the butterflies of the Palaearctic region. Nymphalidae part III. Subfamily Limenitinae Tribe Neptini. Omnes Artes, Milano, 69 pp.

    Bozano, G.C. & Weidenhoffer, Z. (2001) Guide to the butterflies of the palaearctic region. Lycaenidae part I. Subfamily Lycaeninae. Omnes Artes, Milano, 62 pp.

    Bremer, O. (1867) Catalog der Lepidopteren-Sammlung des Professors Dr. Eversmann. Horae Societatis Entomologicae Rossicae, 5, 1–23.

    Bryk, F. (1935) Parnassiidae pars II (Subfam. Parnassiinae). Tierreich, 65, 1–790.

    Burnasheva, A.P. (2012) Butterflies (Lepidoptera, Rhopalocera) of the steppe associations in the Middle Lena River valley. Amurian zoological Journal, 4 (3), 277–283.

    Butler, A.G. (1868) An essay towards an arrangement of the genera of the family Satyridae. Entomologist’s Monthly Magasine, 4, 193–197.

    Churkin, S.V. (2000) Taxonomical notes on Melitaea ambrisia Higgins, 1935 and Melitaea alrashid Higgins, 1941, stat.n. (Lepidoptera, Nymphalidae) from the Central Asia. Helios, 1, 88–98.

    Churkin, S. (2002a) Review of the Erebia ocnus - sibo complex with the description of new taxa (Lepidoptera, Satyridae). Helios, 3, 3–49.

    Churkin, S. (2002b) Taxonomic notes on the Thersamonia solskyi species group (Lepidoptera, Lycaenidae) with the description of a new subspecies. Helios, 3, 172–196.

    Churkin, S.V. (2002c) Review of the Erebia meta species-group from the Tian-Shan and Alai regions with some notes on zoogeography and evolution (Lepidoptera, Satyridae). Helios, 3, 50–93.

    Churkin, S.V. 2002d. Some notes on the genus Athamanthia Zhdanko (Lepidoptera, Lycaenidae). Helios, 3, 115–120.

    Churkin, S. (2004) A review of Neolycaena de Niceville, 1890 from South Siberia, Mongolia and North China with the descriptions of new taxa (Lepidoptera, Lycaenidae). Helios, 5, 153–218.

    Churkin, S. (2006) Taxonomic notes on Neolycaena de Niceville, 1890 from Tian-Shan with the descriptions of new taxa (Lepidoptera, Lycaenidae). Helios, 7, 105–141.

    Churkin, S. (2009) Notes on Parnassius Latreille, 1804 from Tian-Shan and Alai. Part 3: Parnassius charltonius Gray, 1852 (Lepidoptera, Papilionidae). Atalanta, 40 (3/4), 411–434.

    Churkin, S. & Kolesnichenko, K. (2002) A study on Melitaea infernalis Grum-Grshimailo, 1891 (Lepidoptera, Nymphalidae). Helios, 3, 143–157.

    Churkin, S., Grieshuber, J., Bogdanov, P. & Zamolodchikov, D. (2001) Taxonomic notes on Colias tyche Böber, 1812 and Colias nastes Boisduval, 1832 (Lepidoptera, Pieridae) from the Russian Far east with the descriptions of new subspecies. Helios, 2, 103–116.

    Churkin, S.V., Kolesnichenko, K.A. & Tuzov, V.K. (2000) Revision of the Melitaea asteroida species-group (Lepidoptera, Nymphalidae) with new taxa descriptions. Helios, 1, 61–87.

    Churkin, S.V. & Pletnev, V.A. (2012) A review of the Paralasa jordana-complex from Central Asia with descriptions of new taxa (Lepidoptera, Satyridae). Atalanta, 43 (1/2), 120–144.

    Churkin, S.V. & Tuzov, V.K. (1998) Revision of the Paralasa hades species-group from Ghissaro-Darvaz and Pamiro-Alai regions (Lepidoptera, Satyridae). Neue entomologische Nachrichten, 41, 183–195.

    Churkin, S.V. & Tuzov, V.K. (2000) Revision of the Erebia radians species-group (Lepidoptera, Satyridae) from the Tian-Shan and Pamirs-Alai Regions. Helios, 1, 3–28.

    Churkin, S. & Zamolodchikov, D. (2011) The first record of Aglais milberti (Godart, 1819) from the territory of Russia (Lepidoptera, Nymphalidae). Atalanta, 42, 101.

    Churkin, S.V. & Zhdanko, A.B. (2003) A review of the Plebeius idas - subsolanus complex of the Asian part of Russia and Mongolia with the descriptions of new taxa (Lepidoptera, Lycaenidae). Helios, 4, 3–74.

    Condamine, F.L., Sperling, F.A.H., Wahlberg, N., Rasplus, J.-Y. & Kergoat, G.J. (2012) What causes latitudinal gradients in species diversity? Evolutionary processes and ecological constraints on swallowtail biodiversity. Ecology Letters, 15, 267–277.
    http://dx.doi.org/10.1111/j.1461-0248.2011.01737.x

    Coutsis, J.G. (2005) Revision of the Turanana endymion species- group (Lycaenidae). Nota lepidopterologica, 27 (4), 251–272.

    Coutsis, J. G. & Borie, J.-P. (2007) Turanana dushak new to Iran, a re-description of its valva and of the valve of Turanana panaegides, and new sampling locality data for Turanana endymion endimion and Turanana endymion ahasveros (Lepidoptera: Lycaenidae). Phegea, 35 (2), 57–63.

    Crotch, G.R. (1872) On the generic nomenclature of Lepidoptera. Cistula Entomologica, 1, 59–71, 91–92.

    Curtis, J. (1823–1840) British entomology; being illustrations and descriptions of The genera of insects found in Great Britain and Ireland: containing coloured figures from nature of the most rare and beautiful species and in many instances of the plants upon which they are found. Curtis Press, London, 8 vols.

    Dalman, J.W. (1816) Fȯrsȯk till Systematisk uppstållning af Sveriges fjårillar. Kongliae Vetenska Akademiae Handliae, 1, 199–220.

    D’Abrera, B. (1993) Butterflies of the Holarctic region, Part III, Nymphalidae (concl.), Libytheidae, Riodinidae et Lycaenidae. Victoria, I–VII+335–524 pp.

    Dantchenko, A. (2003) On the taxonomy of Plebeius tancrei (Graeser, [1888]) (Lepidoptera, Lycaenidae). Helios, 4, 342–349.

    Dantchenko, A.V. & Lukhtanov, V.A. (1993) Zur Systematik und Verbreitung der Arten der Polyommatus (Agrodiaetus) damone-Gruppe Südosteuropas und Südwestsibiriens (Lepidoptera, Lycaenidae). Atalanta, 24 (1/2), 75–83.

    Davenport, D. (1941) The butterflies of the satyrid genus Coenonympha. Bulletin of the Museum of Comparative Zoology at Harvard College, 87 (4), 213–349.

    Della Bruna, C., Gallo, E. & Sbordoni, V. (2004) Guide to the butterflies of the Palaearctic region. Pieridae part I. Subfamily Pierinae Tribe Pierini (partim). Delias, Aporia, Mesapia, Baltia, Pontia, Belenois, Talbotia. Omnes Artes, Milano, 80 pp.

    Denis, M. & Schiffermüller, I. (1775) Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend. Wien, 322 S.

    Devyatkin, A.L. (1990) To the systematics of skippers of Muschampia poggei group (Lepidoptera, Hesperiidae). Zoologicheskiy Zhurnal, 69 (9), 51–56 [in Russian]

    Devyatkin, A.L. (1992) To the systematics and distribution of the skippers of Muschampia staudingeri Speyer group (Lepidoptera, Hesperiidae). Bulleten Moskovskogo Obstshestva Ispytateley Prirody, 97 (3), 43–56 [in Russian]

    Devyatkin, A.L. (1996) Taxonomic notes on the genus Erynnis Schrank, 1801 (Lepidoptera, Hesperiidae). Atalanta, 27 (3/4), 605–614.

    Devyatkin, A.L. & Tuzov, V.K. (1988) Melitaea lutko (Lepidoptera, Nymphalidae) – a new species for the USSR fauna. Zoologicheskiy zhurnal, 63 (11), 1748–1751 [in Russian]

    Dincă, V., Lukhtanov, V.A., Talavera, G. & Vila R. (2011) Unexpected layers of cryptic diversity in wood white Leptidea butterflies. Nature communications, 8 pp.
    http://dx.doi.org/10.1038/ncomms1329

    Dubatolov, V.V. (2002) The type specimens of two little-known species of Erebia (Lepidoptera, Satyridae) from North-Eastern Siberia. Euroasian entomological Journal, 1 (2), 143–146.

    Dubatolov, V.V. & Lvovsky, A.L. (1997) What is true Yphthima motshulskyi (Lepidoptera, Satyridae)? Transactions of the Lepidopterological Society of Japan, 48 (4), 191–198.

    Dubatolov, V.V., Korb, S.K. & Yakovlev, R.V. (2016) A review of the genus Triphysa Zeller, 1858 (Lepidoptera, Satyridae). Biological Bulletin of Bogdan Chmelnitskiy Melitopol State Pedagogical University, 6 (1), 445–497.
    http://dx.doi.org/10.15421/201628

    Dubatolov, V.V., Korshunov, Yu.P., Gorbunov, P.Yu., Kosterin, O.E. & Lvovsky, A.L. 1998. A review of the Erebia ligea- complex (Lepidoptera, Satyridae) from Eastern Asia. Transactions of the Lepidopterological Society of Japan, 49 (3), 177–193.

    Dubatolov, V.V., Mutin, V.A., Novomodny, E.V. & Dolgikh, A.M. (2010) Distributional limits of butterflies (Insecta, Lepidoptera: Hesperioidea, Papilionoidea) of subboreal and south representatives of temperate complex in Lower Priamurye. Amurian zoological Journal, 2 (3), 253–275.

    Dubatolov, V.V. & Sergeev, M.G. (1987) To the systematics of Lycaenids of the genus Neozephyrus Sibatani et Ito (Lepidoptera, Lycaenidae). Chlenistonogie, klestshi i gelminty, Nauka, Novosibirsk, 18–30 [in Russian]

    Dubrovskiy, D.V. & Vodyanov, K.Y. (2014) Systematical and zoogeographical review of some groups of butterflies (Lepidoptera, Papilionoidea) of Pamir. Quarta, Voronezh, 76 pp. [in Russian]

    Ebert, G. (1971) Die Typen von Parnassius apollonius daubi Fruhstorfer, 1903. Zoologische Mededelingen, 45, 159–160.

    Eckweiler, W. & Bozano, G.C. (2011) Guide to the butterflies of the Palaearctic region. Satyrinae Part IV. Omnes Artes, Milano, 102 pp.

    Eckweiler, W. & Häuser, C.L. (1997) An illustrated checklist of Agrodiaetus Hübner, 1822, a subgenus of Polyommatus Latreille, 1804 (Lepidoptera: Lycaenidae). Nachrichten des entomologischen Vereins Apollo, Suppl. 16, 113–168.

    Eisner, C. (1959) Parnassiana nova XXIV. Kritische Revision der Gattung Tadumia. Zoologische Mededelingen, 34 (10), 165–192.

    Eisner, C. (1966) Parnassiidae-Typen in der Sammlung J.C.Eisner. Zoologische Verhandelingen, 81, 3–190.

    Eisner, C. (1974) Parnassiana Nova XLIX. Die Arten und Unterarten der Baroniidae, Teinopalpidae und Parnassiidae (Erster teil) (Lepidoptera). Zoologische Verhandelingen, 135, 1–96.

    Eitschberger, U. ([1984]) Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.). Herbipoliana, 1, I–XXII, 1–504; 2, 1–601.

    Eitschberger, U. (1989) Siebte Ergänzung zu «Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.)» Pieris bryoniae lappona Rangnow, 1935, comb. nov. prioritätsberechtigt vor Pieris bryoniae bicolorata Petersen, 1947 (Lepidoptera, Pieridae). Atalanta, 20 (1/2), 207–209.

    Eitschberger, U. & Lukhtanov, V.A. (1994) Eine neue Unterart von Oeneis tarpeia Pallas, 1771 aus Buratien im Transbaikal (Lepidoptera, Satyridae). Atalanta, 25 (1/2), 163–166.

    Elwes, H.J. & Edwards, J. (1897) A revision of the oriental Hesperiidae. Transactions of the Royal Zoological Society of London, 14, 101–324.
    http://dx.doi.org/10.1111/j.1096-3642.1897.tb00059.x

    Ermakov, A.I., Olschvang, V.N., Nupponen, K. & Gorbunov, P.Y. (2014) Lepidoptera of the Denezhkin Kamen’ mountain massif of the Urals, Russia. Euroasian entomological Journal, 13 (2), 163–172 [in Russian]

    Ershov, N. & Fil’d, A. (1870) A catalogue of Lepidoptera of the Russian Empire. Horae Societatis Entomologicae Rossicae, 4 (1–2), 130–204.

    Evans, W.H. (1949) A catalogue of the Hesperiidae from Europa, Asia and Australia in the British Museum (Nat. Hist.). Trustees of British Museum, London, 502 pp.

    Forster, W. (1939) Lepidopteren aus Iran. Danish scientific investigations in Iran. Part I. Munsgaard, Copenhagen, 1–7.

    Forster, W. (1960) Bausteine zur Kenntnis der Gattung Agrodiaetus Scudd. (Lep. Lycaen.) II. Zeitschrift der Wiener entomologischen Gesellschaft, 45, 102–145.

    Grum-Grshimaïlo, G.E. (1890) Le Pamir et sa faune lépidoptèrologique. Mémoires sur les Lépidoptères, 4, 1–577.

    Gorbunov, P.Y. (2011) Macrolepidoptera of deserts and southern steppes of Western Kazakhstan. Lisitsina Press, Ekaterinburg, 192 pp. [in Russian]

    Gorbunov, P.Y. & Kosterin, O.E. (2007) The butterflies (Hesperioidea and Papilionoidea) of North Asia in nature. 2. Moscow, 852 pp.

    Gorbunov, P.Y., Olshvang, V.N., Lagunov, A.V., Migranov, M.G. & Gabidullin, A.S. (1992) Butterflies of South Ural. Ekaterunburg, 132 pp. [in Russian].

    Grieshuber, J. (1998) Zur Verbreitung und subspezifischen Gliederung von Colias tyche Boeber, 1812 (Lepidoptera: Pieridae) Teil I. Lambillionea, 98 (1), 126–134.

    Grieshuber, J. (2002) Zur Verbreitung und subspezifischen Gliederung von Colias chrysotheme ( Esper, [1781]) (Lepidoptera: Pieridae). Entomologische Zeitschrift (Stuttgart), 112, 81–88, 299–305.

    Grieshuber, J. (2005) What is Colias tunkuna Austaut, 1912 (Lepidoptera, Pieridae)? Helios, 6, 65–68.

    Grieshuber, J. (2006) The status and type locality of Colias tyche herzi Staudinger, 1901 (Lepidoptera, Pieridae). Helios, 7, 77–83.

    Grieshuber, J. (2014) Guide to the butterflies of Palaearctic region. Pieridae II. Coliadinae. Omnes Artes, Milano, 86 pp.

    Grieshuber, J. & Churkin, S. (2003) The lectotypes of Colias diva Grum-Grshimailo, 1891, Colias wanda Grum-Grshimailo, 1907, Colias grumi Alpheraky, 1897, Colias cocandica maja Grum-Grshimailo, 1891, Colias cocandica tatarica Bang-Haas, 1915, and Colias tamerlana mongola Alpheraky, 1897 (Lepidoptera: Pieridae). Helios, 4, 244–271.

    Grieshuber, J. & Worthy, B. (2004) Notes on Colias palaeno arctica Nordstrom, 1927 (Lepidoptera, Pieridae). Helios, 5, 32–36.

    Grieshuber, J. & Worthy, B. (2006) A revision of Colias wiskotti Staudinger, 1882, the available names and the type material, with a note on Colias alpherakii Staudinger, 1882 (Lepidoptera, Pieridae). Mitteilungen der Münchener Entomologischen Gesellschaft, 96, 43–75.

    Grieshuber, J., Lukhtanov, V.A. & Lvovsky, A.L. (2001) Die Lectotypen von Colias hyperborea Grum-Grshimailo, 1900, Colias aquilonaris Grum-Grshimailo, 1900, und Colias hecla var. orientalis Grum-Grshimailo, 1900 (Lepidoptera: Pieridae). Entomologische Zeitschrift, 111 (6), 168–172.

    Grieshuber, J., Worthy, B. & Lamas, G. (2012) The genus Colias Fabricius, 1807. Jan Haugum's annotated catalogue of the Old World Colias (Lepidoptera, Pieridae). Tshikolovets Publications, Pardubice-Bad Griesbach-Caretham-Lima, 438 pp.

    Grieshuber, J., Churkin, S., Worthy, B. & Lvovsky, A. (2004) The type material of Colias staudingeri Alpheraky, 1881 and Colias staudingeri emivittata Wagner, 1913 (Lepidoptera, Pieridae). Helios, 5, 37–44.

    Gross, F.J. (1978) Beitrag zur Systematik von Pseudochazara-Arten (Lep. Satryridae). Atalanta, 9 (2), 41–103.

    Grote, A.R. (1873) Kleiner Beitrag zur Kenntniss einiger Nordamerikanischer Lepidoptera. Bulletin of the Buffalo Entomological Society, 1, 168–174.

    Grum-Grshimailo, G. (1887) Bericht über meine Reise in das östliche Buchara. Correspondenz. Nebst Anhang: Diagnosen einiger neuen Spezies. Mémoires sur les Lépidoptères, 3, 357–402.

    Grum-Grshimailo, G. (1888) Novae species et varietates Rhopalocerorum e Pamir. Horae Societatis Entomologicae Rosicae, 22, 303–307.

    Groum-Grshimaïlo, G.E. (1890) Le Pamir et sa faune lépidoptèrologique. Mémoires sur les Lépidoptères, 4, 1–577.

    Hanus, J. & Thèye, M.-L. (2013) Les premiers Parnassius: histoire de leur découverte, de leur description et de l’erreur d’identification de Parnassius phoebus (Fabricius, 1793) (Lepidoptera Papilionidae). Alexanor, 25 (6), 323–354.

    Häuser, C.L. (1993) An annotated checklist of the species of the Parnassiinae (Lepidoptera: Papilionidae). Tijdschrift voor Entomologie, 136, 137–146.

    Häuser, C.L., Bartsch, D., Holstein, J. & Steiner, A. (2003) The Lepidoptera type material of G.A.W.Herrich-Schäffer in the Staatkiches Museum für Naturkunde, Stuttgart. Stuttgarter Beiträge Zur Naturkunde, Serie A (Biologie), 657, 1–78.

    Häuser, C.L. & Eckweiler, W. (1997) A catalogue of the species-group taxa in Agrodiaetus Hübner, 1822, a subgenus of Polyommatus Latreille, 1804 (Lepidoptera: Lycaenidae). Nachrichten des Entomologischen Vereins Apollo, Suppl. 16, 53–112.

    Hemming, F. (1933) On the types of certain butterfly genera. Entomologist, 66, 196–200.

    Hemming, F. (1934) The generic names of the Holarctic butterflies. British Museum, London, 184 pp.

    Hemming, F. (1960) Annotationes Lepidopterologicae. Pts 1-2. Hepburn & Sons, London, 72 pp.

    Hemming, F. (1967) The generic names of the butterflies and their type-species (Lepidoptera: Rhopalocera). Bulletin of the British Museum (Natural History), Entomology, Suppl. 9, 509.

    Hesselbarth, G., van Oorschot, H. & Wagener, S. (1995) Die Tagfalter der Türkei unter Berücksichtigung der angrenzenden Länder. Selbstverlag Sigbert Wagener, Bocholt, 1354 S.

    Higgins, L.G. (1941) An illustrated catalogue of the palaearctic Melitaea (Lep. Rhopalocera). Transactions of the entomological Society of London, 91 (7), 175–365.
    http://dx.doi.org/10.1111/j.1365-2311.1941.tb01045.x

    Higgins, L.G. (1950) A descriptive catalogue of the palaearctic Euphydryas (Lepidoptera: Rhopalocera). Transactions of the Royal Entomological Society of London, 101 (12), 435–499.
    http://dx.doi.org/10.1111/j.1365-2311.1950.tb00745.x

    Higgins, L.G. (1955) A descriptive catalogue of the genus Mellicta Billberg (Lepidoptera: Nymphalidae) and its species, with supplementary Remarks on the genera Melitaea and Euphydryas. Transactions of the Royal Entomological Society, 106, 1–131.
    http://dx.doi.org/10.1111/j.1365-2311.1955.tb01262.x

    Honey, M.R. & Scoble, M.J. 2001. Linnaeus’s butterflies (Lepidoptera: Papilionoidea and Hesperioidea). Zoological Journal of the Linnean Society, 132, 277–399.
    http://dx.doi.org/10.1111/j.1096-3642.2001.tb01326.x

    Humphreys, H.N. & Westwood, J.O. (1841) British butterflies and their transformations. William Smit, London, 310 pp.

    Ivonin, V.V., Kosterin, O.E. & Nikolaev, S.L. (2009) Butterflies of Novosibirsk area (Lepidoptera, Diurna). 1. Hesperiidae, Papilionidae, Pieridae. Euroasian Entomological Journal, 8 (1), 85–104 [in Russian]

    Ivonin, V.V., Kosterin, O.E. & Nikolaev, S.L. (2011) Butterflies (Lepidoptera, Diurna) of Novosibirskaya Oblast, Russia. 2. Lycaenidae. Euroasian entomological Journal, 10 (2), 217–242 [in Russian]

    Ivonin, V.V., Kosterin, O.E. & Nikolaev, S.L. (2013) Butterflies (Lepidoptera, Diurna) of Novosibirskaya Oblast, Russia. 3. Nymphalidae (without Satyrinae). Euroasian entomological Journal, 12 (2), 177–199 [in Russian]

    Jong, R. de. (1972) Systematics and geographic history of the genus Pyrgus in the Palaearctic region (Lepidoptera Hesperiidae). Tijdschrift voor Entomologie, 115, 1–121.

    Jong, R. de. (1974) Systematics and evolution of the Palaearctic Spialia species (Lepidoptera, Hesperiidae). Tijdschrift voor Entomologie, 117, 225–271.

    Jong, R. de. (1975) Notes on the genus Pyrgus (Lepidoptera, Hesperiidae). Zoologische Mededelingen, 49, 1–11.

    Jong, R. de (1979) Revision of the Pyrgus alpinus group (Lepidoptera, Hesperiidae), with notes on phylogeny and character displacement. Zoologische Mededelingen, 54 (4), pp. 53–82.

    Jong, R. de (1984) Notes on the genus Thymelicus Hübner (Lepidoptera, Hesperiidae). Nota lepidopterologica, 7, 148–163.

    Jugovec, J. & Koren, T. (2014) Wing pattern morphology of three closely related Melitaea (Lepidoptera, Nymphalidae) species reveals highly inaccurate external morphology-based species identification. Nota lepidopterologica, 37, 75–90.
    http://dx.doi.org/10.3897/nl.37.7966

    Katoh, T., Chichvarkin, A., Yagi, T. & Omoto, K. (2005) Phylogeny and evolution of butterflies of the genus Parnassius: inferences from mitochondrial 16S and ND1 sequences. Zoological Science, 22, 343–351.
    http://dx.doi.org/10.2108/zsj.22.343

    Kesküla, T. (1993) The occurence of Pontia (daplidice) daplidice (Linnaeus, 1758) and P. (daplidice) edusa (Fabricius, 1777) in the Commonwealth of Indepent States (Lepidoptera, Pieridae). Lepidopteroloogiline Informatioon, 8, 1–4.

    Kirby, W.F. (1894) A handbook to the order Lepidoptera. W.H.Allen and Co., London, 261 pp.

    Kleckova, I., Cesanek, M., Fric, F. & Pellissier, R. (2015) Diversification of the cold-adapted butterfly genus Oeneis related to Holarctic biogeography and climatic niche shifts. Molecular phylogenetics and evolution, 92, 255–265.
    http://dx.doi.org/10.1016/j.ympev.2015.06.012

    Klitin, A.K. (1989) Rare insects of Sakhalin Area. Rare and endangered animals. Nauka, Novosibirsk, 134–137 [in Russian]

    Knyazev, S.A., Teploukhov, V.Y. & Rogalyov, V.V. (2011) The new and interesting records of butterflies and moths (Lepidoptera) in Omsk Province. Eversmannia, 25-26, 75–80 [in Russian]

    Knyazev, S.A., Rogalyov, V.V., Ponomaryov, K.B. & Teploukhov V.Y. (2013) New records of butterflies and moths (Lepidoptera) in Omsk Province. Eversmannia, 36, 42–46 [in Russian].

    Kolesnichenko, K.A. & Bush, M.G. (2015) Geographical distribution and subspecies structure of Mellicta ambigua (Ménétriés, 1859) (Lepidoptera, Nymphalidae. Zoologicheskyi zhurnal, 94 (9), 1051–1064 [in Russian]

    Kolesnichenko, K. & Churkin, S. (2003) A review of the Melitaea minerva (Staudinger, 1881) species-group (Lepidoptera, Nymphalidae). Helios, 4, 301–341.

    Kolesnichenko, K.A., Churkin, S.V. & Berdiev, R.K. (2011) A review of the East Palaearctic taxa of the Melitaea didyma (Esper, [1779])-group. Part II. Neue entomologische Nachrichten, 67, 257–278.

    Korb, S.K. (2000) Une sous-espèce nouvelle de Pyrgus alpinus (Erschoff) du T’ien-chan septentrional (Lepidoptera Hesperiidae). Alexanor, 21 (2), 83–87.

    Korb, S.K. (2005) A catalogue of butterflies of the ex-USSR, with remarks on systematics and nomenclature. Korb press, N.Novgorod, 158 pp.

    Korb, S.K. (2006) A neotype designation of Maculinea alcon alconides Korb, 1997 (Lepidoptera, Lycaenidae). Bulleten Moskovskogo Obstshestva ispytatelei prirody, Otdel biologicheskiy, 111 (4), 70 [in Russian]

    Korb, S.K. (2009a) A fauna of butterflies (Lepidoptera, Rhopalocera) of Kungey Ala-Too Mts. (North Tian-Shan). Zoologicheskiy Zhurnal, 88 (9), 1079–1089 [in Russian]

    Korb, S.K. (2009b) A new subspecies of Hyponephele kirghisa (Alphéraky, 1881) from the northern slope of Kirghiz Mts (Lepidoptera, Satyridae). Eversmannia, 19-20, 32–35 [in Russian]

    Korb, S.K. (2011a) Relocation of primary types of butterfl ies (Papilionoidea) described by S. K. Korb in the Y. B. Kosarev collection. Nota lepidopterologica, 34 (1), 83–85.

    Korb, S.K. (2011b) A review of the subgenus Maculinea of genus Phengaris (Lepidoptera, Lycaenidae) of Palaearctics. Eversmannia, 27-28, 22–46 [in Russian]

    Korb, S.K. (2012a) Butterflies of North Tian-Shan (Lepidoptera: Papilionoformes). Hesperiidae, Papilionidae, Pieridae, Satyridae. Eversmannia, Suppl. 3, 84 pp. [in Russian]

    Korb, S.K. (2012b) Polycaena Staudinger, 1886 (Lepidoptera: Riodinidae) in Middle Asia: one species or two? Caucasian entomological Bulletin, 8 (1), 141–143 [in Russian]

    Korb, S.K. (2012c) To the systematics of blue butterflies of the genus Glaucopsyche Scudder, 1872 in the Tian-Shan mountains (Lepidoptera, Lycaenidae). Atalanta, 43 (1/2), 173–180.

    Korb, S.K. (2012d) Plebeius maracandicus (Erschoff, 1874) – a new butterfly species for Upper Volga region. Entomologicheskie i parazitologicheskie issledovaniya v Povolzhje, 10: 98–99 [in Russian]

    Korb, S.K. (2012e) The Parnassiinae systematics based on four genes and morphological features. Eversmannia, 31-32, 5–37 [in Russian]

    Korb, S.K. (2013a) Clarification of data on the Far Eastern representatives of genus Hesperia Fabricius, 1793 (Lepidoptera: Hesperiidae). Eversmannia, 33, 3–4 [in Russian]

    Korb, S.K. (2013b) Recension on the book Tshikolovets V.V., Nekrutenko Yu.P. The butterflies of Caucasus and Transcaucasia (Armenia, Azerbaijan, Georgia and Russian Federation). – Pardubice: Tshikolovets Publications, 2012. 423 pp. Eversmannia, 34, 64–66 [in Russian]

    Korb, S.K. (2013c) Subspecific structure of Parnassius apollo (Linnaeus, 1758) (Lepidoptera: Papilionidae). Eversmannia, 33, 5–9 [in Russian]

    Korb, S.K. (2013d) Butterflies of North Tian-Shan (Lepidoptera: Papilionoformes). Nymphalidae, Libytheidae, Riodinidae, Lycaenidae. Eversmannia, Suppl. 4, 74 pp. [in Russian]

    Korb, S.K. (2013e) To the systematics, biology and diagnostics of a little known species Lycaena triphysina Staudinger, 1891 (Lepidoptera: Lycaenidae). Eversmannia, 34, 32–37 [in Russian]

    Korb, S.K. (2013f) Contribution to the systematics and distribution of lycaenid butterflies of the genus Glaucopsyche Scudder, 1872 (Lepidoptera: Lycaenidae). Amurian zoological Journal, 5 (2), 177–179 [in Russian]

    Korb, S.K. (2013g) A revision of the type specimens of Agriades Hübner, 1819 (Lepidoptera, Lycaenidae) deposited in the Museum für Naturkunde an der Humboldt Universität zu Berlin, with notes on its systematics. Amurian zoological Journal, 5 (3), 309–313 [in Russian]

    Korb, S.K. (2013h) A review of blues of the Polyommatus dagmara group (Grum-Grshimailo, 1888) (Lepidoptera: Lycaenidae). Caucasian entomological Bulletin, 9 (1), 175–180 [in Russian]

    Korb, S.K. (2013i) Notes on the nomenclature of the Palaearctic butteflies (Lepidoptera: Papilionoformes) established in the book of H.C.Lang «Rhopalocera Europæ descripta et delineata». Eversmannia, 36, 3–5 [in Russian]

    Korb, S.K. (2013k) Short review of the Middle Asiatic representatives of the Turanana cytis (Christoph, 1877) group (Lepidoptera: Lycaenidae). Caucasian entomological bulletin, 9 (2), 295–298 [in Russian]

    Korb, S.K. (2014a) Une sous-espèce nouvelle de Muschampia antonia (Speyer, 1879) en vallée de Soussamyr (Kirghizie) (Lepidoptera Hesperiidae Pyrginae). Alexanor, 26 (1), 47–51.

    Korb, S.K. (2014b) A catalogue of the type specimens of Colias Fabricius, 1807 deposited in the Museum für Naturkunde an der Humboldt-Universität zu Berlin (Lepidoptera, Pieridae). Amurian Zoological Journal, 6 (2), 182–206.

    Korb, S.K. (2014c) The lectotype designation of Pararge epaminondas Lang, 1884 and Lasiommata epimenides var. Epaminondas Staudinger, 1887 (Lepidoptera: Satyridae). Amurian zoological Journal, 6 (3), 282–283 [in Russian]

    Korb, S.K. (2014d) New data to the systematics and nomenclature of satyrid genus Karanasa Moore, 1893 (Lepidoptera: Satyridae). Eversmannia, 39, 29–34 [in Russian]

    Korb, S.K. (2014e) New and little known blues (Lepidoptera, Lycaenidae) from Middle Asia and Kazakhstan. Eversmannia, 40, 11–17 [in Russian]

    Korb, S.K. (2014f) To the systematics and nomenclature of Satyridae (Lepidoptera) of Central Asia. Caucasian entomological bulletin, 10 (2), 271–273 [in Russian]

    Korb, S.K. (2015a) Studies of Palaearctic Parnassiinae (Lepidoptera: Papilionidae): the status of Parnassius bremeri amgunensis Sheljuzhko, 1928. Entomologist’s Gazette, 66.

    Korb, S.K. (2015b) The lectotype designation of Anthocharis belia var. pulverata Christoph, 1884 and its comparison to the lectotype of Anthocharis daphalis Moore, 1865 (Lepidoptera, Pieridae). Eversmannia, 42, 14–16 [in Russian]

    Korb, S.K. (2015c) New data concerning the nomenclature, systematics and distribution of butterflies of Middle Asia (Lepidoptera: Nymphalidae, Lycaenidae). Caucasian entomological Bulletin, 11 (1), 189–193 [in Russian]

    Korb, S.K. (2015d) On the oldest available name for the taxon known as Limenitis reducta Staudinger, 1901, on the taxa Papilio rivularis Scopoli, 1763 and Papilio sappho Pallas, 1771 and designation of their primary types (Lepidoptera: Nymphalidae). Eversmannia, 41, 23–27 [in Russian]

    Korb, S.K. (2015e) The butterflies of Inner Tian-Shan (Lepidoptera: Papilionoformes). Eversmannia, Suppl. 6, 1–84 [in Russian]

    Korb, S.K. & Bolshakov, L.V. (2011a) A catalogue of butterflies (Lepidoptera: Papilionoformes) of ex-USSR. Second edition, reformatted and updated. Eversmannia, Suppl. 2, 1–124 [in Russian].

    Korb, S.K. & Bolshakov, L.V. (2011b) Taxonomic notes on the genus Euphydryas Scudder, 1872 (Lepidoptera: Nymphalidae). Eversmannia, 25-26, 25–33 [in Russian].

    Korb, S.K. & Bolshakov, L.V. (2011с) Corrections to the second edition of the «Catalogue of butterflies of the former USSR». Eversmannia, 27-28, 3–6 [in Russian]

    Korb, S.K. & Bolshakov, L.V. (2011d) To the knowledge of systematics of рalaearctic satyrids from the genus Coenonympha Hübner, [1819] (Lepidoptera: Satyridae). Eversmannia, 27-28, 7–21 [in Russian]

    Korb, S.K. & Bolshakov, L.V. (2013) Is Euphydryas aurinia cf. emba (Fruhstorfer, [1917]) a synonym of Euphydryas aurinia sareptana (Staudinger, 1871)? (Lepidoptera: Nymphalidae). Eversmannia, 36, 6–7 [in Russian]

    Korb, S.K. & Bolshakov, L.V. (2014) Nomenclatural notes on the representatives of the genus Euphydryas Scudder, 1871 (Lepidoptera: Nymphalidae) of Russian Far East. Eversmannia, 38, 3–5 [in Russian]

    Korb, S.K. & Rybalkin, S. (2014) Discovery of Hyponephele przhewalskyi Dubatolov, Sergeev & Zhdanko, 1994 (= Hyponephele przhewalskyi bosteriana Korb, 2011) from Boro-Khoro Shan in Kazakhstan (Lepidoptera: Nymphalidae, Satyrinae). Entomologist’s Gazette, 65, 257–259.

    Korb, S.K., Bolshakov, L.V., Fric, Z.F. & Bartonova, A. (2016a) Cluster biodiversity as a multidimensional structure evolution strategy: checkerspot butterflies of the group Euphydryas aurinia (Rottemburg, 1775) (Lepidoptera: Nymphalidae). Systematic Entomology, 41, 441–457.

    Korb, S.K., Fric, Z.F. & Bartonova A. (2016b) On the status and position of Tian-Shanian populations of Paralasa cf. kusnezovi (Avinoff, 1910) (Lepidoptera: Satyridae). Russian entomological Journal, 25, 75–78.

    Korshunov, Y.P. (1972) A catalogue of butterflies (Lepidoptera, Rhopalocera) of the USSR fauna. Entomologicheskoe obozreniye, 51 (1), 136–154; 51 (2), 352–367 [in Russian]

    Korshunov, Y.P. (1988) New butterflies from Khakassia, Tuva and Yakutia. Novye i maloizvestnye vidy fauny Sibiri. Nauka Press, Novosibirsk, 65–80 [in Russian].

    Korshunov, Y.P. (1990) New genera of Parnassiinae. Chlenistonogie i gelminty. Nauka Press, Novosibirsk, 99–105 [in Russian]

    Korshunov, Y.P. & Dubatolov, V.V. (1987) New for the USSR fauna species and subspecies of butterflies. Vestnik zoologii, 1986 (3), 87 [in Russian]

    Korshunov, Y.P. & Ivonin, V.V. (1990) To the systematics of blue butterflies of genera Glaucopsyche and Celastrina. Taksonomiya nasekomykh i gelmintov. Nauka Press, Novosibirsk, 68–79 [in Russian]

    Korshunov, Y.P. & Nikolaev, S.L. (2002) New data to the knowledge of butterflies of the genus Oeneis Hübner, 1819 (Lepidoptera: Satyridae) of North Asia. Euroasian entomological Journal, 1 (2), 147–172 [in Russian]

    Koshkin, E.S. (2014) New records of butterflies (Lepidoptera, Papilionoidea) from Middle Amur region of Evreiskaya Avtonomnaya Oblast, Russia. Euroasian entomological Journal, 13 (1), 74–78 [in Russian]

    Kozlov, M.V. & Jalava, J. (1994) Lepidoptera of the Kola Peninsula, northwestern Russia. Entomologica Fennica, 5, 65–85.

    Kozlov, M.V. & Kullberg, J. (2010) New and interesting records of Lepidoptera from the Kola Peninsula, Northwestern Russia, in 2000–2009. Entomologica Fennica, 21, 254–272.

    Kozlov, M. V., Kullberg, J. & Zverev V. E. (2014) Lepidoptera of Arkhangelsk oblast of Russia: a regional checklist. Entomologica Fennica, 25, 113–141.

    Kreuzberg, A.V.-A. (1985) Papilionids of the groups delphius, charltonius, simo (Lepidoptera, Papilionidae) of the fauna od USSR. Issledovaniya flory i fauny Sredney Asii. Fan, Tashkent, 25–65 [in Russian]

    Kreuzberg, A.V.-A. (1998) Distribution and biology of the rare Nymphalid species Melitaea acraeina Staudinger, 1886 (Lepidoptera, Nymphalidae). Bulleten Moskovskogo Obstshestva Ispytateley Prirody, Otdelenie biologii, 103 (1), 38–40 [in Russian]

    Kudrna, O. (1975) A revision of the genus Gonepteryx Leach (Lep., Pieridae). Entomologist's Gazette, 6 (1): 3–37.

    Kudrna, O. (1977) A revision of the genus Hipparchia Fabricius. Oxon, Farington. 204 pp.

    Kudrna, O. (1981) An annotated list of the butterflies named by Colin W. Wyatt (Lepidoptera: Papilionoidea, Hesperioidea). Bonner Zoologischen Beiträge, 32, 221–236.

    Kudrna, O. (1983) An annotated catalogue of the butterflies (Lepidoptera: Papilionoidea) named by Roger Verity. Journal of Research on the Lepidoptera, 21 (1), 1–106.

    Kudrna, O. & Belicek, J. (2005) The ‘Wiener Verzeichnis, its authorship, publication date and some names proposed for butterflies therein. Oedippus, 23, 1–32.

    Kudrna, O., Harpke, A., Lux, K., Pennerstorfer, J., Schweiger, O., Settele, J. & Wiemers, M. (2011) Distribution atlas of butterflies in Europe. Gesellschaft für Schmetterlingschutz, Halle, 576 pp.

    Kusnetsov, V.I. & Stekolnikov, A.A. (2001a) New methods to the system of Lepidoptera of the world fauna (based on the functional morphology of abdomen). Nauka, Saint-Petersburg, 462 pp. [in Russian]

    Kusnetsov, V.I. & Stekolnikov, A.A. (2001b) Comparative morphology of the male genitalia in the subfamilies Morphinae and Brassolinae and the resurrection of the family Satyridae, status resurr. (Lepidoptera) in new limits. Entomologicheskoe obozrenie, 80 (1), 121–136. [in Russian]

    Kuznetsov, G.I., Stradomsky, B.V. & Fomina, E.S. (2013) Record of population of Callophrys chalybeitincta Sovynski, 1905 (Lepidoptera: Lycaenidae) in Volgograd Region (Russia). Caucasian entomological Bulletin, 9 (2), 293–294 [in Russian]

    Kuznetsov, G.I., Stradomsky, B.V. & Fomina, E.S. (2014) Two forms of Melitaea trivia ([Denis et Schiffermüller], 1775) (Lepidoptera: Nymphalidae) from Volgograd Region (Russia): the systematic position on the basis of molecular genetic studies. Caucasian entomological bulletin, 10 (1), 119–120 [in Russian]

    Latreille, P.A. (1810) Considérations générales sour l'ordre naturel des animaux composant les classes des Crustacés, des Arachnides et des insectes. Schoell, Paris, 444 pp.

    Lindsey, A.W. (1925) The types of Hesperioid genera. Annales of the Entomological Society of America, 18, 75–106.
    http://dx.doi.org/10.1093/aesa/18.1.75

    Lorkovič, Z. (1930) Verwandtschefliche Beziehungen in der morsei-major-sinapis Gruppe des Gen. Leptidea (Kritischer Beitrag zur Auffassung des Speziesbegriffes). Zeitschrift des Österreichischen Entomologie-Vereins, 15 (6), 61–67.

    Lvovsky, A.L. & Morgun, D.V. (2007) Butterflies of East Europe. KMK, Moscow, 443 pp. [in Russian]

    Lukhtanov, V.A. (1984) Review of the palaearctic satyrids of the genus Oeneis Hübner (Lepidoptera, Satyridae). I. O. hora Gr.-Gr. species group. Entomologicheskoe Obozreniye, 63 (4), 776–789 [in Russian].

    Lukhtanov, V.A. (1987) A review of palaearctical butterflies of the genus Oeneis Hübner (Lepidoptera, Satyridae). II. O. jutta species group. Entomologicheskoe obozreniye, 66 (2), 142–158 [in Russian]

    Lukhtanov, V.A. (1989) A review of the palaearctic satyrids of the group Oeneis norna (Lepidoptera, Satyridae). Vestnik zoologii, 1989 (2), 29–36; 1989 (4), 28–34 [in Russian]

    Lukhtanov, V.A. (1990) New taxa of Satyrids and Lycaenids (Lepidoptera, Satyridae, Lycaenidae) from Kazakhstan and Middle Asia. Vestnik zoologii, 1990 (6), 13–23 [in Russian]

    Lukhtanov, V.A. (1995) Eine Übersicht über die Arten der Untergattung Satyrium (Superflua Strand, 1910) mit der Beschreibung einer neuen Art aus Südwestgissar, Usbekistan (Lepidoptera: Lycaenidae, Theclinae). Nachrichten des entomologischen Vereins Apollo, N.F., 16, 47–58.

    Lukhtanov, V.A. (1999) Neue Taxa und Synonyma zentralasiatischer Tagfalter (Lepidoptera, Papilionoidea). Atalanta, 30 (1/4), 135–150.

    Lukhtanov, V.A. (2000) Zur Systematik und Verbreitung der Taxa der Athamanthia dimorpha-Gruppe (Lepidoptera, Lycaenidae). Atalanta, 31 (1/2), 179–192.

    Lukhtanov, V. & Lukhtanov, A. (1994) Die Tagfalter Nordwestasiens (Lepidoptera, Diurna). Herbipoliana, 3, 1–440.

    Lukhtanov, V.A. & Tikhonov, V.V. (2015) Chromosomal and molecular evidence for presence of Polyommatus (Agrodiaetus) poseidon (Lepidoptera, Lycaenidae) in Caucasus region. Comparative Cytogenetics, 9 (2), 249–255.
    http://dx.doi.org/10.3897/CompCytogen.v9i2.5020

    Masui, A., Bozano, G.C. & Floriani, A. (2011) Guide to the butterflies of the palaearctic region. Nymphalidae part IV. Subfamily Apaturinae. Omnes Artes, Milano, 82 pp.

    Matsuda, S. (1996) Taxonomic status of Thecla orientalis var. cognata Staudinger (Lepidoptera, Lycaenidae), with some notes on its related species. Tyô to Ga, 47 (3), 174–184.

    Mazel, R. (2005) Éléments de phylogénie dans le genre Leptidea Billberg, 1820 (Lepidoptera, Pieridae, Dismorphinae). Revue de l´Association Roussillonnaise d´ Entomologie, 14 (3), 98–111.

    Mazel, R. & Leestmans, R. (2003) Analýse et discussion de quelques types ifraspécifiques dans le genre Leptidea Billberg, 1820 (Lepidoptera: Pieridae). I: types asiatiques. Revue de l´ Association Roussillonnaise d´ Entomologie, 19 (4), 179–192.

    Michel, F., Rebourg, C., Cosson, E. & Descimon, H. (2008) Molecular phylogeny of Parnassiinae butterflies (Lepidoptera: Papilionidae) based on the sequences of four mitochondrial DNA segments. Annales de la Société. entomologique de France (N.S.), 44, 1–36.

    Moore, F. ([1893 - 1896]) Rhopalocera. Family Nymphalidae. Sub-Family Satyridae. Lepidoptera Indica, 2, I–VII, 1–274.

    Morgun, D.V. (2013) Notes on the Paleophilotes (Inderskia) panope (Eversmann, 1851) (Lepidoptera: Lycaenidae), a rare and little known lycaenid taxon for Europe. Zootaxa, 3619 (5), 595–600.
    http://dx.doi.org/10.11646/zootaxa.3619.5.9

    Morgun, D.V. & Tikhonov, V.V. (2009) New records of butterfly fauna (Lepidoptera: Hesperioidea, Papilionoidea) from Daghestan. Caucasian entomological bulletin, 5 (2), 273–274. [in Russia]

    Nazari, V. & Sperling, F.A. (2007) Mitochondrial DNA divergence and phylogeography in western palaearctic Parnassiinae (Lepidoptera: papilionidae): How many species are there? Insect Systematics and Evolution. 38, 121–138.
    http://dx.doi.org/10.1163/187631207788783996

    Nazari, V., Zakharov, E.V. & Sperling, F.A.H. (2007) Phylogeny, historical biogeography, and taxonomic ranking of Parnassiinae (Lepidoptera, Papilionidae) based on morphology and seven genes. Molecular phylogenetics and evolution. 42, 131–156.
    http://dx.doi.org/10.1016/j.ympev.2006.06.022

    Nazari, V., ten Hagen, W. & Bozano, G.C. (2009) Molecular systematic and phylogeny of the ‘Marbled Whites’ (Lepidoptera: Nymphalidae, Satyrinae, Melanargia Meigen). Systematic Entomology, 35 (1), 132–147.
    http://dx.doi.org/10.1111/j.1365-3113.2009.00493.x

    Nekrutenko, Y.P. (1968) Phylogeny and distribution of Gonepteryx (Lepidoptera, Pieridae). Naukova Dumka, Kiyv, 152 pp. [in Russian]

    Nekrutenko, Y.P. (1980) Revisional notes on lycaenid butterfly species assigned to Ultraaricia Beuret (Lycaenidae). Nota lepidopterologica, 3 (1–2), 55–68.

    Nekrutenko, Y.P. (1983) Revision of the genus Hyrcanana (Lepidoptera, Lycaenidae). Vestnik zoologii, 1983 (3), 7–16 [in Russian]

    Nekrutenko, Y.P. (1984a) A revision of the type specimens of the group Lycaena phoenicurus (Lepidoptera, Lycaenidae). Vestnik zoologii, 1984 (6), 43–49 [in Russian]

    Nekrutenko, Y.P. (1984b) The blue butterflies of the genus Chilades (Lepidoptera, Lycaenidae) of the fauna of USSR. Vestnik zoologii, 1984 (3), 29–40 [in Russian]

    Nekrutenko, Y.P. (1985) The butterflies of Crimea. Naukova Dumka, Kiev, 152 pp. [in Russian]

    Nekrutenko, Y.P. (1990) Butterflies of Caucasus. Naukova Dumka, Kiev, 216 pp. [in Russian]

    Nekrutenko, Y.P. (1998) A catalogue of the type specimens of Riodinidae and Lycaenidae deposited in the collection of Zoologisches Forschungsinstitut und Museum Alexander Koenig (Bonn). Nota lepidopterologica, 21 (2), 119–148.

    Nekrutenko, Y.P. (2000) A catalogue of the type specimens of Palaearctic Riodinidae and Lycaenidae (Lepidoptera, Rhopalocera) deposited in the collection of the Museum für Naturkunde der Humboldt Universität zu Berlin. Nota lepidopterologica, 23 (3/4), 192–352.

    Nekrutenko, Y.P. & Effendi, R.M.E. (1983) A review of the blue butterflies of the group Lycaena phoenicurus Ld. (Lepidoptera, Lycaenidae) with description of a new species from Azerbaidzhan. Vestnik zoologii, 1983 (4), 8–15 [in Russian]

    Nekrutenko, Y.P. & Kerzhner, I.M. (1986) On the species and varieties of Parnassius (Lepidoptera, Papilionidae), established by E. Menetries in the book of Y.Siemashko «Russian fauna». Entomologicheskoe obozreniye, 65 (4), 769–779 [in Russian]

    Nekrutenko, Y.P. & Tshikolovets, V.V. (2005) Butterflies of Ukraine. Rayevsky Press, Kiev, 232 pp. [in Ukrainian]

    Nikolaev, S.L. (2004) Neotype of Erebia ligea uralensis Goltz, 1932 (Lepidoptera, Satyridae). Euroasian entomological Journal, 3 (3), 225–226.

    Novomodny, E.V. & Fonova, E.A. (2010) Butterflies (Insecta, Lepidoptera: Hesperioidea, Papilionoidea) of Ayan-Maysky district of Khabarovsk Province. Amurian zoological Journal, 2 (4), 322–337 [in Russian]

    Olivier, A., van der Poorten, D. & de Prins, W. (1998) Rhopalocera and Grypocera of Turkey 16. Taxonomic notes on Plebeius christophi and Plebeius idas in northeastern Turkey (Lepidoptera: Lycaenidae). Phegea, 26 (3), 87–102.

    Omoto, K., Katoh, T., Chichvarkin,, A. & Yagi, T. (2004) Molecular systematics and evolution of the “Apollo» butterflies of the genus Parnassius (Lepidoptera: Papilionidae) based on mitochondrial DNA sequence data. Gene, 326, 141–147.
    http://dx.doi.org/10.1016/j.gene.2003.10.020

    Omoto, K., Yonezawa, T. & Shinkawa, T. (2008) Molecular systematics and evolution of the recently discovered "Parnassian" butterfly (Parnassius davydovi Churkin, 2006) and its allied species (Lepidoptera, Papilionidae). Gene, 330, 1–9.

    Pazhenkova, E.A., Zakharov, E.V. & Lukhtanov, V.A. (2015) DNA barcoding reveals twelve lineages with properties of phylogenetic and biological species within Melitaea didyma sensu lato (Lepidoptera, Nymphalidae). ZooKeys, 538, 35–46.
    http://dx.doi.org/10.3897/zookeys.538.6605

    Peña, C., Witthauer, H., Klečková, I., Fric, Z. & Wahlberg N. (2015) Adaptive radiations in butterflies: evolutionary history of the genus Erebia (Nymphalidae: Satyrinae). Biological Journal of the Linnean Society, 116 (2), 449–467.
    http://dx.doi.org/10.1111/bij.12597

    Pisanets, E.M. & Tshikolovets, V.V. (2001) A catalogue of the types specimens deposited in Zoological Museum of Academy of Sciences of Ukraine. Part 1. Monogenoidea, Trematoda, Cestoda, Nematoda, Hirudinea, Arachnida, Crustacea, Gastropoda, Insecta, Chondrichthyes, Actinopterygii, Amphibia, Reptilia, Mammalia. Zoological Museum of Academy of Sciences of Ukraine, Kiev, 140 pp. [in Russian]

    Prins, W.O. de & van der Poorten, D. (1995) Rhopalocera and Grypocera of Turkey 14. Taxonomic revision of the Pyrgus alveus (Hübner, [1803]) complex from Greece to West China, with description of two new species from southern Turkey (Lepidoptera: Hesperiidae). Phegea, 23 (1), 1–44.

    Reissinger, E. (1971) Die geographisch-subspecifische Gliederung von Colias alfacariensis Ribbe unter Berücksichtigung der Migrationsverhältnisse (Lepidoptera Pieridae). Atalanta, 3 (4), 145–176.

    Renner, F. (1991) Neue Untersuchungsergebnisse aus der Pyrgus alveus Hübner Gruppe in der Palaearktis unter besonderer Berücksichtigung von Süddeutschland (Lepidoptera: Hesperidae [sic!]). Neue entomologische Nachrichten, 28, 1–157.

    Rose, K. (1995) Zur Unterarten-Inflation in der Gattung Parnassius (Lepidoptera, Papilionidae). Nechrichten des entomologischen Vereins Apollo, N.F, 16 (2/3), 243–252.

    Rothschild, L. (1919) List of types of Lepidoptera in the Tring Museum. Novitates Zoologicae, 26, 193–251.

    Rubin, N.I. & Yakovlev, R.V. (2013) Checklist of the butterflies (Papilionoidea) of the Saur Mountains and adjacent territories (Kazakhstan), including systematic notes about the Erebia callias group. Nota lepidopterologica, 36 (2), 137–170.

    Sachkov, S.A. (1991) Butterflies described by P.S.Pallas from Samara and its environs. Samarskaya Luka, Bulletin, 1, 108–110 [in Russian].

    Samodurov, G.D., Tschikolowez, W.W. & Korolew, W.A. (1995) Eine Übersicht über die Satyriden der Gattung Hyponephele Muschamp, 1915. I. Die Arten Hyponephele haberhaueri (Staudinger, 1886), H. germana (Staudinger, 1887), H. maureri (Staudinger, 1886), H. rueckbeili (Staudinger, 1887) und H. interposita (Erschoff, 1874) (Lepidoptera, Satyridae). Atalanta, 25 (1/2), 157–195.

    Samodurov, G.D., Korolev, V.A. & Tshikolovez, V.V. (1996) Eine Übersicht über die Satyriden der Gattung Hyponephele Muschamp, 1915. II. Die Arten Hyponephele dysdora (Lederer, 1870), H. tristis (Grum-Grshimailo, 1899), H. prasolovi Lukhtanov, 1990, H. murzini Dubatolov, 1989 und H. jasavi Lukhtanov, 1990 (Lepidoptera, Satyridae). Atalanta, 27 (1/2), 223–252.

    Samodurov, G.D., Korolev, V.A. & Tshikolovez, V.V. (1997) Eine Übersicht über die Satyriden der Gattung Hyponephele Muschamp, 1915. III. Die Arten Hyponephele cadusia (Lederer, 1869), H. cadusina (Staudinger, 1881), H. laeta (Staudinger, 1886), H. pamira Lukhtanov, 1990, H. kirghisa (Alpheraky, 1881), H. sheljuzhkoi Samodurov et Tshikolovets, 1996, H. rubriceps Herz, 1900 und H. pseudokirgisa J.J.Stshetkin, 1984 (Lepidoptera, Satyridae). Atalanta, 28 (1/2), 49–96.

    Sañudo-Restrepo, C.P., Dincă, V., Talavera, G. & Vila, R. (2012) Biogeography and systematics of Aricia butterflies (Lepidoptera, Lycaenidae). Molecular Phylogenetics and Evolution.
    http://dx.doi.org/10.1016/j.ympev.2012.10.010

    Schurian, K.G. (1988) Neueinteilung des Subgenus Lysandra der Gattung Polyommatus Latreille (Lepidoptera: Lycaenidae). Entomologische Zeitschrift, 98, 129–144.

    Schurian, K.G. (1989a) Ergänzungen und Berichtigungen zu: «Neueinteilung des Subgenus Lysandra der Gattung Polyommatus Latreille (Lepidoptera: Lycaenidae)». Entomologische Zeitschrift, 99, 305–311.

    Schurian, K.G. (1989b) Revision der Lysandra-Gruppe des Genus Polyommatus Latr. (Lepidoptera: Lycaenidae). Neue entomologische Nachrichten, 24, 1–181.

    Scudder, S.H. (1872) A systematic revision of some of the American butterflies. Annual Reports of Peabody Academy of Sciences, 4, 24–83.

    Scudder, S.H. (1874) The species of the lepidopterous genus Pamphila. Memoires of the Boston Socirty of Natural History, 2, 341–353.

    Scudder, S.H. (1875) Historical sketch of the generic names proposed for butterflies. Naturalists Agency, Salem, 193 pp.

    Sheljuzhko, L. (1931) Einige Randbemerkungen zum «Supplement» zu den «Groß-Schmetterlingen der Erde», herausgegeben von Dr.A.Seitz. Papilionidae und Pieridae. Internationale entomologische Zeitschrift, 25 (7), 69–74.

    Shima, H. (1988) Phylogenetic relationships of the genus Ypthima Hübner (Lepidoptera, Satyridae). Special Bulletin of the lepidopterological Society of Japan, 6, 69–81.

    Sinev, S.Y. (2008) A catalogue of Lepidoptera of Russia / Ed. S.Y.Sinev. KMK, Saint-Petersburg-Moscow, 424 pp. [in Russian]

    Sokolov, G.N. (1929) Male genitalia structure in Parnassius Latr. Revue Russe d’Entomologique, 23 (1–2), 60–75.

    Staudinger, O. (1892) Lepidopteren des Kentei-Gebirges. Deutsche entomologische Zeitschrift Iris, 5, 360–393.

    Staudinger, O. & Wocke, M. (1871) Catalog der Lepidopteren des europaeischen Faunengebietes. Staudinger, Dresden, 426 pp.

    Stshurov, V.I., Gorbachev, P.Y. & Lukhtanov, V.A. (2007) [Lepidoptera: part]. Red Data Book of Krasnoyarsk Region (Animals), Edition 2, Krasnodar, 246–279 [in Russian]

    Stradomsky, B.V. (2005) Taxonomy of the closely related species Plebeius argyrognomon (Bergsträsser, [1779]), P. maracandicus (Erschoff, 1874) and P. caspicus (Forster, 1936) stat.n. (Lepidoptera: Lycaenidae). Caucasian entomological Bulletin, 1 (1), 81–84 [in Russian]

    Stradomsky, B.V. & Fomina, E.S. (2013) The comparison of taxa Polyommatus yurinekrutenko Koçak, 1996 and P. shamil (Dantchenko, 2000) (Lepidoptera: Lycaenidae). Caucasian entomological bulletin, 9 (1), 181–182 [in Russian]

    Stradomsky, B.V. & Korb, S.K. (2015) On the status of Polygonia c-album var. interposita Staudinger, 1881 (Lepidoptera: Nymphalidae). Caucasian entomological bulletin, 11 (2), 407–409.

    Stradomsky, B.V. & Vodolazhsky, D.I. (2011) Callophrys rubi (Linnaeus, 1758) and C. chalybeitincta Sovynski, 1905 (Lepidoptera: Lycaenidae): a compara