Abstract
The genus Speluncarius Reitter, 1886 (Harpalinae, Pterostichini) is made up by anophthalmous or almost blind species. The genus is related to Tapinopterus Schaum, 1858, which contains numerous species from the Balkan-Anatolian region and have more or less developed eyes. Speluncarius is distributed in the Western Palaearctic region; 26 species and one subspecies are known from Italy, the Balkan Peninsula and Turkey. The genus is classified in five subgenera, Elasmopterus Kraatz, 1886 (6 spp.); Hypogearius Jeannel, 1953 (3 spp.); Hypogium Tschitscherine, 1900 (1 sp.); Pontotapinus Guéorguiev & Lohaj, 2008 (1 sp.) and Speluncarius (s.str.) Reitter, 1886 (15 spp.) (Sciaky, 1982; Bousquet, 2003; Guéorguiev & Lohaj, 2008; Casale et al., 2013).
References
Bousquet, Y. (2003) Harpaliinae, Pterostichini. In: Löbl, I. & Smetana, A. (Eds.), Catalogue of the Palearctic Coleoptera. Vol. 1. Apollo Books, Stenstrup, pp. 469–521.
Casale, A., Giachino, P.M. & Vailati, D. (2013) Tre nuove specie di Coleotteri sotterranei di Grecia (Coleoptera: Carabidae e Cholevidae). Bollettino della Società Entomologica Italiana, 145 (1), 9–25.
http://dx.doi.org/10.4081/BollettinoSEI.2013.9Guéorguiev, B.V. & Lohaj, R. (2008) Studies on genus Speluncarius, with description of a new subgenus and notes on the systematic position of S. (Hypogium) albanicus (Coleoptera, Carabidae, Pterostichini). In: Penev, L., Erwin, T. & Assman, T. (Eds.), Back to the Roots and Back to the Future. Towards a New Synthesis amongst Taxonomic, Ecological and Biogeographical Approaches in Carabidology. Proceedings of the XIII European Carabidologists Meeting, Blagoevgrad, 20–24 August 2007, pp. 125–141.
Sciaky, R. (1982) Le attuali conoscenze sul genere Speluncarius Reitt. (VII contributo alla conoscenza dei Coleoptera Carabidae). Giornale Italiano di Entomologia, 1, 15–33.