Abstract
A new species of the genus Nigrobaetis Novikova & Kluge, 1987 is described based on larvae from the Caucasus Mountains. Nigrobaetis (Takobia) katerynae Martynov, sp. nov., is easily distinguishable from other Takobia species by the following combination of characters: the shapes of frontal suture, mandibles, labial palp, paraproctal prolongation, gills; the number of setal rows along the external margins of femora; the absence of gills on tergum I; and the colour pattern of body.
References
Bengtsson, S. (1912) Neue Ephemeriden aus Schweden. Entomologisk Tidskrift, 33 (1–2), 107–117.
Bogoescu, C. & Tabacaru, I. (1957) Contributii la studiul sistematic al nimfelor de ephemeroptere din R.P.R. Genul Baetis Leach. Buletin Stiintific, 9 (3), 241–284.
Braasch, D. & Soldán, T. (1983) Baetidae in Mittelasien III (Ephemeroptera). Entomologische Nachrichten und Berichte, 27 (6), 266–271.
Dubey, O.P. (1971) Torrenticole insects of the Himalaya. VI. Description of nine new species of Ephemerida from the Northwest Himalaya. Oriental Insects, 5, 521–548.
https://doi.org/10.1080/00305316.1971.10434039Fujitani, T. (2006) Japanese Baetidae (Ephemeroptera): keys to seven genera with information on taxonomy, distribution and habitat. Japanese Journal of Limnology, 67, 185–207 [in Japanese, with English summary]
https://doi.org/10.3739/rikusui.67.185Gattolliat, J.-L. (2011) A new species of Alainites (Ephemeroptera: Baetidae) from Borneo (East Kalimantan, Indonesia). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 84 (3–4), 185–192.
Godunko, R.J., Palatov, D.M. & Martynov, A.V. (2015) Mayflies of the Caucasus Mountains. III. A new representative of the subgenus Rhodobaetis Jacob, 2003 (Baetidae: Baetis) from the South-Western Caucasus. Zootaxa, 3948 (2), 182–202. http://dx.doi.org/10.11646/zootaxa.3948.2.2
Gose, K. (1980) The mayflies of Japanese. Part 6. Aquabiology 2 (1), 76–79. [in Japanese]
Hubbard, M.D. (1990) Mayflies of the World: A Catalog of the Family and Genus Group Taxa (Insecta: Ephemeroptera). Sandhill Crane Press, Gainesville, Florida (Flora & Fauna Handbook, No. 8), 119 pp.
Illies, J. (1978) Limnofauna Europaea. Second edition. Gustav Fischer, Stuttgart, 532 pp.
Imanishi, K. (1937) Mayflies from Japanese torrents. VIII. Notes on the genera Paraleptophlebia and Baetis. Annotationes Zoologicae Japonenses, 16 (4), 330–339.
Kakauridze, T.G. (1946) Raspredelenie benticheskoi fauny v gornoi rechke Dabakhana [Distribution of benthic fauna in mountain river Dabakhana]. Trudy Zoologicheskogo instituta AN Gruzinskoi SSR, VI, 16. [in Russian]
Kang, S.C., Chang, H.C. & Yang, C.T. (1994) A revision of the genus Baetis in Taiwan (Ephemeroptera, Baetidae). Journal of Taiwan Museum, 47 (2), 9–44.
Kasymov, A.G. (1972) Presnovodnaya fauna Kavkaza [Freshwater fauna of the Caucasus]. Elm, Baku, 287 pp. [in Russian]
Kazanci, N. & Thomas, A.G.B. (1989) Complements et corrections a la faune des Ephemeropteres du Proche-Orient: 2. Baetis kars n.sp. de Turquie. Mitteilungen Schweizerischen Entomologischen Geselschaft, 62, 323–327.
Kazlauskas, R.S. (1972) Neues über das System der Eintagsfliegen der Familie Baetidae (Ephemeroptera). Proc. XIII Int. Congr. Entomol., Moscow, 2–9 August 1968, 3, 337–338.
Kluge, N.J. & Novikova, E.A. (2014) Systematics of Indobaetis Müller-Liebenau & Morihara 1982, and related implications for some other Baetidae genera (Ephemeroptera). Zootaxa, 3835 (2), 209–236.
https://doi.org/10.11646/zootaxa.3835.2.3Kluge, N.J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, 456 pp.
http://dx.doi.org/10.1007/978-94-007-0872-3Leach, W.E. (1815) Entomology. Brewster's Edinburg Encyclopedia, Ed.1, 9 (1), 57–172.
Linnaeus, C. (1758) Systema Naturae per regna tria Naturae, secundum classes, ordines, genera, species, cum Characteribus, Differentiis, Synonymis, Locis. Ed.10. Vol.1. Typis Ioannis Thomae, Vindobonae [Vienna], 824 pp.
Linnaeus, C. (1761) Fauna Svecica sistens Animalia Sveciae regni: mammalia, aves, amphibia, pisces, insecta, vermes. Distributa per classes & ordines, genera & species, cum differentiis specierum, synonymis auctorum, nominibus incolarum, locis natalium, descriptionibus insectorum. Editio altera, auctior. Stockholmiae: Sumtu & Literis Direct. Laurentii Salvii, 578 p.
Martynov, A.V. Godunko, R.J. & Palatov, D.M. (2016) Kintrishi State Nature Reserve — a hotspot for mayfly (Insecta: Ephemeroptera) diversity in Adjara (Georgia). Zoosymposia, 11, 168–173.
http://dx.doi.org/10.11646/zoosymposia.11.1.18
Müller-Liebenau, I. (1969) Revision der europäischen Arten der Gattung Baetis Leach, 1815 (Insecta, Ephemeroptera). Gewässer und Abwässer, 28/29, 1–214.
Müller-Liebenau, I. (1974) Baetidae aus Südfrankreich, Spanien und Portugal (Insecta, Ephemeroptera). Gewässer und Abwässer, 53/54, 7–42.
Müller-Liebenau, I. (1984) New genera and species of the family Baetidae from West Malaysia (River Gombak) (Insecta: Ephemeroptera). Spixiana, 7 (3), 253–284.
Novikova, E.A. & Kluge, N.J. (1987) Sistematika roda Baetis (Ephemeroptera, Baetidae) s opisaniem novogo vida iz Sredney Asii [Systematics of the genus Baetis (Ephemeroptera, Baetidae), with description of new species from Middle Asia]. Vestnik zoologii, 4, 8–19. [in Russian, with English summary]
Novikova, E.A. & Kluge, N.J. (1994) Podenki podroda Nigrobaetis (Ephemeroptera, Baetidae, Baetis Leach, 1815) [Mayflies of the subgenus Nigrobaetis (Ephemeroptera, Baetidae, Baetis Leach, 1815)]. Entomologicheskoe Obozrenie, 73 (3), 623–644. [in Russian, with English summary]
Palatov, D.M. (2013) Novye dannye o faune i rasprostranenii podenok semejstva Baetidae (Ephemeroptera) na territoriii Kavkaza i Zakavkaz'ja [New data on the fauna and distribution of mayflies family Baetidae (Ephemeroptera) of the Caucasus and Transcaucasia]. In: Hydroentomology in Russia and adjacent countries: Materials of the Fifth All-Russia Symposium on Amphibiotic and Aquatic Insects / Papanin Institute for Biology of Inland Waters, Russian Academy of Sciences.—Yaroslavl: Filigran, pp. 107–114. [in Russian, with English summary]
Puthz, V. (1978) Ephemeroptera, pp. 256–263, l table, In: Illies, J. (Ed.), Limnofauna Europaea. Second edition. Gustav Fischer, Stuttgart.
Salur, A., Darilmaz, M.C. & Bauernfeind, E. (2016) An annotated catalogue of the mayfly fauna of Turkey (Insecta, Ephemeroptera). ZooKeys, 620, 67–118.
https://doi.org/10.3897/zookeys.620.9405Sartori, M. & Thomas, A.G.B. (1989) Contribution à la connaissance du genre Baetis Leach, 1815 en Corse (Ephemeroptera, Baetidae). B. albinatii nov. sp. du groupe muticus. Annls Limnol, 25 (2), 131–137.
https://doi.org/10.1051/limn/1989013Sroka, P. (2012) Systematics and phylogeny of the West Palaearctic representatives of subfamily Baetinae (Insecta: Ephemeroptera): combined analysis of mitochondrial DNA sequences and morphology. Aquatic Insects, 34 (1), 23–53.
https://doi.org/10.1080/01650424.2012.718081Thomas, A. & Gagneur, J. (1994) Compléments et corrections à la faune des Ephéméroptères d'Afrique du Nord. 6. Alainites sadati n. sp. d'Algérie (Ephemeroptera, Baetidae). Bulletin de la Société d'Histoire Naturelle de Toulouse, 130, 43–45.
Thomas, A.G.B., Bouzidi, A., Sartori, M., Assef, S. & Ajakane, A. (1992) Compléments et corrections à la faune des Ephéméroptères d'Afrique du Nord. 5. Baetis oukaimeden, n. sp. du Haut-Atlas marocain: description et écologie (Ephemeroptera, Baetidae). Mitt. Schweiz. Ent. Ges. Bulletin de la Société Entomologique Suisse, 65, 369–377.
Tong, X.-L. & Dudgeon, D. (2000) Two new species of Alainites (Ephemeroptera: Baetidae) from Hong Kong, China. Pan Pacific Entomologist, 46 (2), 115–120.
Waltz, R.D. & McCafferty, W.P. (1997) New generic synonymies in Baetidae (Ephemeroptera). Entomological News, 108 (2), 134–140.
Waltz, R.D., McCafferty, W.P. & Thomas, A. (1994) Systematics of Alainites n. gen., Diphetor, Indobaetis, Nigrobaetis n. stat., and Takobia n. stat. (Ephemeroptera, Baetidae). Bulletin de la Société d'Histoire Naturelle de Toulouse, 130, 33–36.
Zimmermann, W. (1981) Beitrag zur Kenntnis der Gattung Baetis Leach, 1815 (Insecta, Ephemeroptera) im Kaukasus und in Transkaukasien (UdSSR). Entomologische Nachrichten, 25 (7–8), 97–112.
Zrelli, S., Gattolliat, J.L., Boumaiza, M. & Thomas, A. (2012) First record of Alainites sadati Thomas, 1994 (Ephemeroptera: Baetidae) in Tunisia, description of the larval stage and ecology. Zootaxa, 3497, 60–68.