Abstract
The female bears a pair of highly distinctive postocellar horns similar to those of three other species endemic to Costa Rica, E. corniculatus, E. echeverrii, and E. verticicornis. The males of two additional new species, E. morrisoni and E. wagneri, are described from premontane wet forest on the Pacific slope near Monteverde, resulting in a total of 13 species recorded from Costa Rica.
References
Belle, J. (1970) Studies on South American Gomphidae (Odonata). Studies on the Fauna of Suriname and other Guyanas, 11 (43), 1–-158.
Belle, J. (1980) A new species of Epigomphus from Guatemala (Odonata: Gomphidae). Entomologische Berichten, 40 (9), 136–138.
Belle, J. (1981) A new species of Epigomphus from Mexico (Odonata: Gomphidae). Entomologische Berichten, 41 (4), 61–63.
Belle, J. (1988) Epigomphus gibberosus, a new species from Peru, with lectotype designations for the eligible species of the genus Epigomphus (Odonata: Gomphidae). Tijdschrift voor Entomologie, 131, 135–140.
Belle, J. (1989) Epigomphus corniculatus, a new dragonfly from Costa Rica. Tijdschrift voor Entomologie, 132, 158–160.
Belle, J. (1993) Epigomphus jannyae spec. nov., a new dragonfly from Panama (Anisoptera: Gomphidae). Odonatologica, 22 (2), 187–189.
Belle, J. (1994) Three new neotropical Gomphidae from the genera Archaeogomphus Williamson, Cyanogomphus Selys and Epigomphus Hagen (Anisoptera). Odonatologica, 23 (1), 45–50.
Brooks, S.J. (1989) New dragonflies (Odonata) from Costa Rica. Tijdschrift voor Entomologie, 132 (2), 163–176.
Calvert, P.P. (1903) On Some American Gomphinae (Odonata). Entomological News, 14, 183–192.
Calvert, P.P. (1905) Insecta Neuroptera. Odonata. Biologia Centrali Americana, 1905, 145–212.
Calvert, P.P. (1908) Odonata. Biologia Centrali Americana, 1908, 405–420.
Calvert, P.P (1920) The Costa Rican species of Epigomphus and their mutual mating adaptations (Odonata). Transactions of the American Entomological Society, 46, 323–354.
Donnelly, T.W. (1986) Epigomphus westfalli spec. nov., a new dragonfly from Nicaragua (Anisoptera: Gomphidae). Odonatologica, 15 (1), 37–41.
Donnelly, T.W. (1987) Structural variation of Ophiogomphus mainensis: Description of a new subspecies and relationship to sibling species (Odonata: Gomphidae), Proceedings of the Entomological Society of Washington, 89, 205–214.
Donnelly, T.W. (1989) Three new species of Epigomphus from Belize and Mexico (Odonata: Gomphidae) Florida Entomologist, 72 (3), 428–435.
Garrison, R.W., von Ellenrieder, N. & Louton, J.A. (2006) Dragonfly Genera of the New World. The Johns Hopkins University Press, Baltimore, 368 pp.
González-Soriano, E. & Cook, C. (1988) Una nueva especie de Epigomphus Selys 1854, del estado de Veracruz (Odonata: Gomphidae). Folia Entomológica Mexicana, 74, 5–13.
Esquivel, C. (2006) Libélulas de Mesoamérica y el Caribe. Dragonflies and Damselflies of Middle America and the Caribbean. Editorial INBio, Santo Domingo de Heredia, 319 pp.
Förster, S. (2001) The Dragonflies of Central America Exclusive of Mexico and the West Indies. 2nd Edition. Gunnar Rehfeldt, Braunschweig, 141 pp.
Fraser, F.C. (1947) Notes on the Selysian types of Epigomphus paludosus Hagen in Selys. Annals of the Entomological Society of America, 40 (4), 672–676.
https://doi.org/10.1093/aesa/40.4.672Kennedy, C.H. (1936) Epigomphus crepidus, a new dragonfly (Odonata: Gomphidae) from Nayarit, Mexico, with notes on the genus. Annals of the Entomological Society of America, 29 (1), 126–135.
https://doi.org/10.1093/aesa/29.1.126Kennedy, C.H. (1946) Epigomphus subquadrices, a new dragonfly (Odonata: Gomphidae) from Panama, with notes on E. quadrices and Eugomphus n. subgen. Annals of the Entomological Society of America, 39 (4), 662–666.
https://doi.org/10.1093/aesa/39.4.662Paulson, D.R. (1998) Variation in head spines in female Ophiogomphus, with a possible example of reproductive character displacement (Anisoptera: Gomphidae). Bulletin of American Odonatology, 5, 55–58.
Paulson, D.R. (2016a) Web site: Biodiversity Resources, Dragonflies. Slater Museum of Natural History, University of Puget Sound. Available from: http://www.pugetsound.edu/academics/academic-resources/slater-museum/biodiversity-resources/dragonflies/ (accessed 26 November 2016)
Paulson, D.R. (2016b) IUCN List of Threatened Species.
https://doi.org/10.2305/IUCN.UK.2009-2.RLTS.T165007A5959644.enRamírez, A., Paulson, D.R. & Esquivel, C. (2000) Odonata of Costa Rica: Diversity and Checklist of Species. Revista de Biología Tropical, 48, 245–252.
Ris, F. (1918) Libellen (Odonaten) aus der Region der amerikanischen Kordilleren von Costarica bis Catamarca. Archiv für Naturgeschichte, Abteilung A, 82 (9), 1–197. [1916]
Selys-Longchamps, E. de (1854) Synopsis des Gomphines. Bulletin de l’Académie royale de Belgique, (2), 21, 23–112.
Selys-Longchamps, E. de (1869) Secundes additions au synopsis des Gomphines. Bulletin de l'Académie Royale de Belgique, (2), 28, 168–208 (1–45 reprint). [exact ref from Garrison et al. 2006 – E. obtusus]
Selys-Longchamps, E. de (1878) Quatriémes additions au synopsis des Gomphines. Bulletin de l'Académie Royale de Belgique, (2), 46, 408–698 (1–106 reprint). [exact ref from Garrison et al. 2006 – E. subobtusus]