Abstract
Iran has a complex dragonfly fauna influenced by contacts and overlaps of different geographical zones. Its fauna is dominated by Eurosiberian taxa. However, the SE Province Sistãn-va-Baluchestãn is rich in oriental species, many of which having their western distribution limit in Iran. In NE-Iran, Irano-Turanian elements live and in the S- and SW-Iran African species are found. The Iranian Odonata fauna seems well studied, however, a closer look reveals many uncertainties and confusion, some records coming clearly from misidentification whereas other, which were missing proofs of correct identification remains to be confirmed. Even today, every new collecting trip reveals species new for Iran whereas species new for science are still detected, although rarely. In this checklist we include seven taxa new for Iran: Stylurus ubadschii (although a male of uncertain origin is recorded in Schmidt (1954), Ischnura senegalensis (although two uncertain records were published by Martin (1912) and Schmidt (1954), Coenagrion ponticum, C. lunulatum, C. pulchellum, Lestes macrostigma, and Calopteryx splendens tschaldirica. We critically checked all available data, including all published records. Fourteen taxa have been rejected, or placed in the category for need of confirmation. Till the end of 2017, 100 autochthonous taxa of dragonflies and two migratory species could be confirmed to be or have been present in Iran. We provide distribution maps, created after evaluation of published data and containing our own data from 12 trips to Iran, travelling over 35000 km in the country. Over 200 new localities are integrated. Taxonomic confusion is reduced by rejecting the following taxa for Iran: Calopteryx splendens mingrelica, C. hyalina, Erythromma najas, Aeshna cyanea, Gomphus davidi, and Sympetrum sinaiticum. We regard the taxa Gomphus amseli and G. schneiderii transcaspicus as synonyms of G. schneiderii, and Onychogomphus forcipatus lucidostriatus as a synonym of O. f. albotibialis.
References
Ahmadi, R., Mohebbi, F., Hagigi, P., Esmailly, L. & Salmanzadeh, R. (2011) Macro-invertebrates in the wetlands of the Zarrineh estuary at the south of Urmia Lake (Iran). International Journal of Environmental Research, 5 (4), 1047–1052.
Ahmadi, R., Aliyev, A., Seidgar, M., Bayramov, A. & Ganji, S. (2012) Macroinvertebrate Communities Differences on Riverine Parts and Reservoirs of Zarrineh River. American Journal of Agricultural and Biological Sciences, 7 (1), 71–75.
https://doi.org/10.3844/ajabssp.2012.71.75
Akramowski, N.N. (I948) Dragonfly fauna of Soviet Armenia. Zoologicheskii Sbornik, Akademya Nauk Armyanskoi SSR, 5, 1I7–188. [in Russian]
Asahina, S. (1963) Odonata taken by Japanese expeditions to Karakoram, Afghanistan, Iran and Western Pakistan. In: Ueno, M. (Ed.), Insect Fauna of Afghanistan and Hindukush 4. Committee of the Kyoto University Scientific Expedition to the Karakoram and Hindukush, Kyoto University, Japan, pp. 45–49.
Asahina, S. (1973) The odonata of Iraq. Japanese Journal of Zoology, 17, 17–36.
Bakhshi, Y. & Sadeghi, S. (2014) Odonata fauna of Khuzestan province (Iran) with a note on an Oriental species Trithemis aurora (Burmeister, 1839) as a new record for the south-west of Iran. Zoology in the Middle East, 59, 280–282.
https://doi.org/10.1080/09397140.2014.966522
Ball, L. (2014) An investigation of odonate communities within Wadi Sayq, Dhofar province, Oman (Insecta: Odonata). Check List, 10, 857–863.
https://doi.org/10.15560/10.4.857
Bartenev, A.N. (1912) The Palaearctic and East Asian species and subspecies of the genus Calopteryx Leach (Odonata, Calopterygidae). Raboty laboratorii Zoologicheskago kabineta Imperatorskago Varshavskago Universiteta, I, 63–257. [In Russian]
Bartenev, A.N. (1915) Sympetrum vulgatum flavum subsp. nova. In Insectes pseudoneuropteres (Insecta Pseudoneuroptera). Vol 1 Libellulidae. In Faune de la Russie et des pays limitrophes (Ed. Nasonov, N.V.), Académie impériale des Sciences, Petrograd, pp. 317–332. [in Russian]
Bartenev, A.N. (1916) Contributions à la faune des Odonates du Nord de Perse. Revue russe d’ Entomologie, 16, 38–45. [In Russian]
Bartenev, A.N. (1929) Donnéecs nouvelles sur les Odonates de la Transcaucasie, de la Perse et du Turkestan. Revue russe d’ Entomologie, 23, 124–131. [In Russian]
Belyshev, B. (1964) Strekozy Sibiri. Trudy Vostochno-Sibirskogo Filiala AN SSSR. Seriya biologocheskaya, 40, 61. [In Russian]
Blom, W.L. (1982) List of Odonata collected during various lepidopterological trips in Iran (1971-1974). Notulae Odonatologicae, 1, 150–151.
Borisov, S.N. (2014) Distribution and ecology of Ischnura forcipata Morton, 1907 (Odonata, Coenagrionidae) in Tien-Shan and Pamir-Alai. Euroasian Entomological Journal, 13, 323–328.
Borisov, S.N. & Haritonov, A.Y. (2007) The Dragonflies (Odonata) of Middle Asia. Part 1. Caloptera, Zygoptera. Euroasian Entomological Journal, 6, 343–360.
Boudot, J.-P. & Kalkman, V.J. (Eds.), (2015) Atlas of the European dragonflies and damselflies. KNNV publishers, the Netherlands, 381 pp.
Boudot, J.-P., Jacquemin, G. & Dumont, H.J. (1990) Revision of the subspecies of Onychogomphus forcipatus (Linnaeus, 1758) in Europe and Asia Minor, and the true distribution of Onychogomphus forcipatus unguiculatus (Vander Linden, 1823) (Odonata, Gomphidae). Bulletin & Annales de la Société Royale Belge d'Entomologie, 126, 95–111.
De Knijf, G., Sparrow, D.J., Demetriou, A., Kent, R., Kent, H., Siedle, K. & Crossley, L. (2016) Distribution, ecology and status of a threatened species Ischnura intermedia (Insecta: Odonata), new for Europe. International Journal of Odonatology, 19, 257–274.
https://doi.org/10.1080/13887890.2016.1259662
Dumont, H.J. (1977) A review of the dragonfly fauna of Turkey and adjacent mediterranean islands (Insecta Odonata). Bulletin et Annales de la Société Royale Belge d’Entomologie, 113, 119–171.
Dumont, H.J. (2004) Dragonflies from Azerbaijan. Zoology in the Middle East, 31, 87–92.
https://doi.org/10.1080/09397140.2004.10638027
Dumont, H.J. & Borisov, S.N. (1995) Status and range of the species-pair Ischnura forcipata Morton, 1907 and Ischnura intermedia Dumont, 1974 (insecta: Odonata: Coenagrionidae). Biologisch jaarboek Dodonaea, 62, 157–163.
Dumont, H.J. & Heidari, H. (1996) On a collection of spring odonata from Iran, with the description of Coenagrion australocaspicum n. sp. Bulletin et Annales de la Société Royale Belge d’Entomologie, 132, 63–78.
Dumont, H.J., Demirsoy, A. & Verschuren, D. (1987) Breaking the Calopteryx-bottleneck: taxonomy and range of Calopteryx splpendens waterstoni Schneider, 1984 and of C. splendens tschaldirica, 1909 (Zygoptera: Calopterygidae). Odonatologica, 16, 239–247.
Dumont, H.J., Schneider, W., Verschuren, D. & Pavesi, M. (1995) Coenagrion lindenii zernyi (Schmidt): an east mediterranean taxon threatened by extinction by introgression from the west (Zygoptera: Coenagrionidae). Odonatologica, 24, 451–459.
Dumont, H.J., Heidari, H. & Atamuradov, K.I. (1997) Hybridisation in Calopteryx orientalis (Selys) East of the shores of the Caspian lake (Zygoptera: Calopterygidae). Odonatologica, 26, 205–213.
Dumont, H.J., Kiany, M. & Sadeghi, S. (2011) First record of Rhodischnura nursei (Morton) from Iran (Zygoptera: Coenagrionidae). Odonatologica, 40, 251–254.
Dumont, H.J., Ikemeyer, D. & Schneider, T. (2017) Lestes concinnus and L. pallidus: two non-metallic species with wide, complementary ranges (Odonata: Lestidae). Odonatologica, 46, 99–110.
Ebrahimi, A., Madjdzadeh, S.M. & Mohammadian, H. (2009) Dragonflies (Odonata) from South-Eastern Iran. Caspian Journal of Environmental Science, 7, 107–112.
Ebrahimi, A., Mohammadian, H. & Madjdzadeh, S.M. (2014) A note on libellulid dragonflies (Odonata) of Khabr National Park (Kerman province, South-Eastern Iran). International Dragonfly Fund—Report, Journal of the International Dragonfly Fund, 69, 1–9.
Eslami, Z., Rad, S.P. & Dumont, H.J. (2014) Sympetrum flaveolum (Odonata: Libellulidae) a new species record for Iran. Journal of Entomological Society of Iran, 34, 71–73.
Eslami, Z., Rad, S.P. & Dumont, H.J. (2015) Dragonflies (Odonata) found in the central of Iran (the Northwest of Isfahan province). International Journal of Zoology and Research, 5, 25–32.
Fraser, F.C. (1929) A revision of the Fissilabioidae (Cordulegasteridae, Petaliidae, and Petaluridae) (Order Odonata). Part 1. Cordulegasteridae. Memoirs of the Indian Museum, 9, 69–167. plus plates IX–XII.
Fraser, F.C. (1934) The fauna of British India, including Ceylon and Burma. Odonata. Volume 2. Ariidae, Gomphidae. Taylor and Francis London, Great Britain, 398 pp, 120 figures, 1 coloured plate.
Ghahari, H. & Thipaksorn, A. (2014) A preliminary checklist of Odonata (Insecta) from Thearasbaran Biosphere Reserve and vicinity, Northwestern Iran. The Journal of Tropical Asian Entomology, 3, 48–54.
Ghahari, H., Tabari, M., Sakenin, H., Ostovan, H. & Imani, S. (2009) Odonata (Insecta) from Northern Iran, with comments on their presence in rice fields. Munis Entomology & Zoology, 4, 148–154.
Ghahari, H., Thipaksorn, H., Naderian, H., Sakenin, H. & Tajali, A.A. (2012) A faunistic study on the Odonata (Insecta) from Kurdistan province and vicinity, western Iran. Linzer biologische Beiträge, 44, 1079–1085.
Gitashenasi (2009) Map of administrative divisions of Islamic Republic of Iran, No. 296. Gitashenasi Geographical & Cartographic Institut, Teheran. ISBN 978-964-342-075-8.
Haritonov, A.Y. (1988) Strekosi roda Ischnura Charp. (Insecta: Odonata) fauny SSSR. In: Zolotarenko, G.S (Ed.), Taksonomiya zhivotnyh Sibiri, Nauka, Novosibirsk, pp. 32–46.
Heidari, H. & Dumont, H.J. (2002) An annotated check-list of the Odonata of Iran. Zoology in the Middle East, 26, 133–150.
https://doi.org/10.1080/09397140.2002.10637929
Holuša, O. (2015) Description of the female of Cordulegaster vanbrinkae (Lohmann, 1993) (Odonata: Anisoptera: Cordulegastridae). Zootaxa, 2, 229–238.
https://doi.org/10.11646/zootaxa.3949.2.5
Holuša, O., Dalecký, V. & Imanpour Namin, J. (2015) Habitat choice of Cordulegaster vanbrinkae in Iran (Odonata: Cordulegastridae). Odonatologica, 44, 11–20.
Ikemeyer, D., Schneider, E., Schneider, J. & Schneider, T. (2015) Records of Odonata in North-and North-East Iran including Sympecma gobica (Förster, 1900) as a new species for Iran. Entomologische Zeitschrift, 125, 147–152.
Jeziorski, P. (2013) First record of Sympecma gobica (Odonata: Lestidae) from Iran. Klapalekiana, 49, 39–42.
Jödicke, R., Borisov S.N., Haritonov, A.Y. & Popova, O. (2000) Additions to the knowledge of Sympetrum sinaiticum (Odonata: Libellulidae). International Journal of Odonatology, 3, 131–140.
https://doi.org/10.1080/13887890.2000.9748144
Kalkman, V.J. (2006) Key to the dragonflies of Turkey. Brachytron, 10, 3–82.
Kalkman, V.J. & van Pelt, G.J. (2006) The distribution and flight period of the dragonflies of Turkey. Brachythron, 10, 83–153.
Kalkman, V.J., van Pelt, G.J., Dumont, H.J., Haritonov, A.Y. & Tailly, M. (2004) Critical species of Odonata in Turkey, Iran and the Caucasus. International Journal of Odonatology, 7, 325–339.
https://doi.org/10.1080/13887890.2004.9748220
Kalkman, V.J., Aagaard, K. & Dolmen, D. (2015) Pyrrhosoma nymphula (Sulzer, 1776). In: Boudot, J.-P. & Kalkman, V.J. (Eds.), Atlas of the European dragonflies and damselflies, KNNV publishing, the Netherlands, 143-144
Kalkman, V.J., Iversen, L.L. & Nielsen, E. (2015) Aeshna isoceles (Müller, 1767). In: Boudot, J.-P. & V.J. Kalkman (Eds.), Atlas of the European dragonflies and damselflies, KNNV publishing, the Netherlands, pp. 157–158.
Kiany, M. & Sadeghi, S. (2016) Odonata from southwest of central desert of Iran with occurrence notes on Ischnura intermedia Dumont, 1974. Iranian Journal of Animal Biosystematics, 2, 67–76.
Kiany, M., Sadeghi, S. & Ehteshami, F. (2016) New record of Platycnemis kervillei and Lestes dryas (Odonata: Platycnemididae & Lestidae) from Iran. 19th National and 7th International Congress of Biology, 30. Aug–1.Sep 2016, University of Tabriz, Iran, 94 pp. [no official publication, congress data without peer review]
Kimmins, D.E. (1950) The 3th Danish Expedition to Central Asia. Zoological Results 4. Odonata, Ephemeroptera and Neuroptera (Insecta) from Afghanistan. Videnskabelige Meddelelser Dansk Naturhistorisk Forening, 112, 235–241.
Kosterin, O. & Solovyev, V. (2017) Odonata found in mid-summer 2015 and 2016 at the north-westernmost Black Sea Coast of the Caucasus, with the first record of Cordulegaster picta Selys, 1854 in Russian Federation. International Dragonfly Fund—Report, Journal of the International Dragonfly Fund, 107, 1–43.
Lambret, P., Boudot, J.-P., Chelmick, D., De Knijf, G., Durand, É., Judas, J. & Stoquert, A. (2017) Odonata surveys 2010–2016 in the United Arab Emirates and the Sultanate of Oman, with emphasis on some regional heritage species. Odonatologica, 46, 153–205.
Lohmann, H. (1990) Anax immaculifrons Rambur, 1842 in Iran (Odonata: Aeshnidae). Opuscula Zoologica Fluminensia, 52, 9–10.
Lohmann, H. (1992) Gomphus kinzelbachi Schneider in Iran. Notulae odonatologicae, 3, 169.
Lohmann, H. (1993a) Coenagrion vanbrinkae spec. nov. and C. persicum spec. nov. aus Vorderasien (Zygoptera: Coenagrion). Odonatologica, 22, 203–211.
Lohmann, H. (1993b) Revision der Cordulegastridae. 2. Beschreibung neuer Arten in den Gattungen Cordulegaster, Anotogaster, Neallogaster und Sonjagaster (Anisoptera). Odonatologica, 22, 273–294.
Malikova, E.I. (1997) The fauna of dragonflies (Insecta, Odonata) of the Amurskaya oblast. Problemy ecologii verkhnego Priamurya. Vypusk 3. Blagovestshensk, pp. 109–116. [In Russian]
Martin, R. (1912) Les odonates. In: de Morgan, J. (Ed.), Annales d´Histoire Naturelle, Delegation en Perse, Tome II Entomologie. Ernest Leroux, Paris, 2, pp. 5–9, liste des localités avec altitude, pp. 1-2.
Morton, K.J. (1915-1916) Some Palaearctic species of Cordulegaster. The Transactions of the Entomological Society of London 1915, 273–290, plates 34-37..
Morton, K.J. (1919) Odonata from Mesopotamia. Entomologists’ Monthly Magazine, 3, 143–151, 183–196.
Morton, K.J. (1920) Odonata collected in North-Western Persia by Captain P.A. Buxton. Entomologists’ Monthly Magazine, 56, 82–87.
Morton, K.J. (1921) Neuroptera. Mecoptera and Odonata from Mesopotamia and Persia. Entomologists’ Monthly Magazine, 57, 213–225.
https://doi.org/10.5962/bhl.part.22331
Morton, K.J. (1924) The dragonflies (Odonata) of Palestine, based primarily on collections made by Dr. P. A. Buxton, with notes on the species of the adjacent regions. Transactions Entomological Society London, 73, 25–44.
Nasirian, H. & Irvine, K.N. (2017) Odonata larvae as a bioindicator of metal contamination in aquatic environments: application to ecologically important wetlands in Iran. Environmental Monitoring and Assessment, 189, 436.
https://doi.org/10.1007/s10661-017-6145-6
Okudaira, M., Sugimura, M., Ishida, S., Kojima, K., Ishida, K. & Aoki, T. (2001) The dragonflies of the Japanese Archipelago in color. Hokkaido University press, Sapporo, Japan, 641 pp.
Porter, R.F. (2016) Odonata from Iraq, with three new records. Notulae odonatologicae, 8, 247–318.
Radde, G. (1899) Orthoptera caucasica. In: Radde, G. (Ed.), Die Sammlungen des Kaukasischen Museums, Band 1 Zoologie, Typographie der Kanzlei des Landeschefs. Tiflis, p. 454.
Rastegar, J., Havaskary, M., Khodaparast, S. & Rafeii, A. (2013) A contribution to the knowledge of Odonata (Insecta) from West Azarbaijan province, northwestern Iran. Entomofauna, 34, 369–375.
Ris, F. (1928) Zur Erforschung des Persischen Golfes. (Beitrag Nr. 8): Libellen (Odonata). Wiener Entomologische Zeitung, 44, 155–164.
Ris, F. in von Bodemeyer, B. (1930) Über meine Entomologischen Reisen nach Iran, das Ebursgebirge (1914). Bd. 4: Iran, das Elbursgebirge, Stuttgart. Odonata, 91–92.
Sadeghi, S. & Dumont, H.J. (2004) First record of Libellula fulva pontica Selys, 1887 (Odonata, Anisoptera) from Iran. Zoology in the Middle East, 32, 116–117.
https://doi.org/10.1080/09397140.2004.10638055
Sadeghi, S. & Mohammadalizadeh, J. (2009) Addition to the Odonata Fauna of Iran. Iranian Journal of Science & Technology, Transaction A, 33, 255–359.
Sage, B.L. (1960) Notes on the Odonata of Iraq. Entomologist, 93, 118–125.
Schmidt, E. (1939) Odonata aus Syrien und Palästina. Sitzungsberichte der österreichischen Akademie der Wissenschaften, Wien, Kl. 1, 147, 135–150.
Schmidt, E. (1954) Die Libellen Irans. Sitzungsberichte der österreichischen Akademie der Wissenschaften, Wien, Kl. 1, 163, 223–260.
Schmidt, E. (1961) Ergebnisse der Deutschen Afghanistan-Expedition 1956 der Landessammlungen für Naturkunde Karlsruhe sowie der Expeditionen J. Klapperich, Bonn 1952-53 und Dr. K. Lindberg, Lund (Schweden) 1957–60. Beiträge naturkundlicher Forschung SW-Deutschland, 19, 399–435.
Schneider, T. & Dumont, H.J. (2015) Odonata records from southern Iran. Notulae odonatologicae, 8, 137–146.
Schneider, T. & Ikemeyer, D. (2016a) Notes on Odonata species in South-West Iran including Plactycnemis kervillei (Martin, 1909) as a new species for Iran. Entomologische Zeitschrift, 126, 3–8.
Schneider, T. & Ikemeyer, D. (2016b) Records of Odonata species from Khorãsãn-e-Razavi and Khorãzãn-e-Shomãli in Northeast-Iran. Entomologische Zeitschrift, 126, 211–216.
Schneider, T. & Ikemeyer, D. (2017) Late summer observations on Odonata from the Armenian Highland to the Talysh Mountains in north-west Iran. Entomologist’s Monthly Magazine, 153, 1–12.
Schneider, T. & Schneider, J. (2010) Occurrence, behaviour, and habitat preference of the Levant Pincertail Onychogomphus macrodon Selys, 1887 in Turkey (Insecta: Odonata). Zoology of the Middle East, 49, 79–88.
https://doi.org/10.1080/09397140.2010.10638393
Schneider, T., Schneider, E., Schneider, J. & Müller, O. (2014) Rediscovery of Cordulegaster vanbrinkae in Iran (Odonata: Cordulegastridae). Odonatologica, 43, 25–34.
Schneider, T., Ikemeyer, D. & Dumont, H.J. (2015a) New records of Dragonflies (Odonata) from Belutschistan-e-Sistan province Iran. Zoology of the Middle East, 62, 288–290.
https://doi.org/10.1080/09397140.2015.1069249
Schneider, T., Schneider, E., Schneider, J., Vierstraete, A. & Dumont, H.J. (2015b) Aeshna vercanica spec. nov. from Iran (Anisoptera: Aeshnidae) and a new insight into the Aeshna cyanea-group. Odonatologica, 44, 81–106.
Schneider, T., Ikemeyer D., Ferreira, S. & Müller, O. (2016) Rediscovery and redescription of Coenagrion persicum (Lohmann 1993) with description of the female, and some notes on habitat selection (Odonata: Coenagrionidae). Zootaxa, 6, 561–573.
https://doi.org/10.11646/zootaxa.4103.6.6
Schneider, T., Ikemeyer, D., Ferreira, S. & Müller, O. (2017a) Gomphus kinzelbachi Schneider, 1984 in Iran: identification, habitat and behaviour (Odonata: Gomphidae). Zootaxa, 4216 (6), 572–584.
https://doi.org/10.11646/zootaxa.4216.6.4
Schneider, T., Schneider, E., Ikemeyer, D., Seidenbusch, R. & Müller, O. (2017b)Description of last instar larva of Gomphus kinzelbachi Schneider, 1984 and new aspects on distribution and habitats in Iran (Odonata: Gomphinae). Zootaxa, 4365 (6), 455–466.
https://doi.org/10.11646/zootaxa.4216.6.4
Schneider, W. (1987) Die Verbreitung von Onychogomphus macrodon Selys, 1887, mit der Beschreibung des bisher unbekannten Weibchens und einer Wiederbeschreibung des Männchens (Odonata: Gomphidae). Opuscula Zoologica Fluminensia, 13, 1–12.
Schneider, W. & Dumont, H.J. (1997) The dragonflies and damselflies (Insecta: Odonata) of Oman. An updated and annotated checklist. Fauna of Saudi Arabia, 16, 89–110.
Schröter, A., Seehausen, M., Kunz, B., Günther, A., Schneider, T. & Jödicke, R. (2015) Update of the Odonata fauna of Georgia, southern Caucasus ecoregion. Odonatologica, 44, 279–342.
Seehausen, M., Schröter, A., Mumladze, L. & Grebe, B. (2016) Additional Odonata records from Georgia, southern Caucasus ecoregion, with the first record of Ischnura fountaineae (Odonata: Libellulidae). Notulae odonatologicae, 8, 266–283.
Seidenbusch, R. (1997a) Structural maginal features for discrimination in Gomphus vulgatissimus Linnaeus, 1758 and Gomphus schneideri selys, 1850. Sulzbach-Rosenberger Libellenrundbrief, 5, 5–8. [privately published]
Seidenbusch, R. (1997b) Gomphus amseli Schmidt, 1961 versus Gomphus schneideri amseli Schmidt, 1961—Gomphus schneiderii transcaspicus ssp. nov.—new results from the south-east Caspian territory. Sulzbach-Rosenberger Libellenrundbrief, 5, 5–12. [privately published]
Seidenbusch, R., Ikemeyer, D. & Schneider, T. (2015) Description of the last instar exuviae of Cordulegaster vanbrinkae from Iran and comparison with the last instar exuviae of Cordulegaster insignis nobilis (Odonata: Cordulegastridae). Sulzbach-Rosenberger Libellenrundbrief, 18, 1–25. [privately published]
Selys Longchamps, E. (de) (1887) Odonates de l’Asie mineure et revision des autres parties de la faune paléarctique (dite Européenne). Bulletin et Annales de la Société Royale Belge d’Entomologie, 31, 1–85.
Skvortsov, V.E. & Kuvaev A.V. (2010) Ischnura fountaineae Morton, Lindenia tetraphylla (Vander Linden) and Selysiothemis nigra (Vander Linden) new for European Russia (Zygoptera: Coenagrionidae; Anisoptera: Gomphidae, Libellulidae). Notulae odonatologicae 7, 49–51.
Skvortsov, V.E. & Snegovaya, N.Y. (2014) Additions to the knowledge of the Odonata fauna of Azerbaijan, with six new records. Notulae Odonatologicae, 8, 37–76.
Soufi, M. & Jafari, A. (2011) Impacts of habitat destruction on wetland biodiversity. World Applied Sciences Journal, 12, 1897–1902.
Tailly, M., Ananian, V. & Dumont, H.J. (2004) Recent dragonfly observations in Armenia with an updated checklist. Zoology in the Middle East, 31, 93–102.
https://doi.org/10.1080/09397140.2004.10638028
Tariq, M. (2010) Systematics of the dragonflies (Anisoptera: Odonata) of Pakistan. PhD thesis, Arid Agriculture University, Rawalpindi, Pakistan, pp. 1–190.
Valle, K.J. (1942) Odonaten aus dem Westlichen Zentralasien. Annales Entomologici fenici, 8, 114–126.
Viganò, M., Janni, O. & Corso, A. (2017) Tramea basilaris on Linosa Island, Italy: A new species for Europe and the Western Palaearctic (Odonata: Libellulidae). Odonatologica, 46, 55–66.
Wildermuth, H. (2008) Die Falkenlibellen Europas—Corduliidae. Die Neue Brehm-Bücherei Bd. 653—Westarp Wissenschaften Verlagsgesellschaft, Hohenwarsleben, 496 pp.
Yakubovich, V.S. (2010) First record of the dragonfly Orthetrum albistylum speciosum (Uhler, 1858) (Odonata: Libellulidae) from Khabarovskii krai. Far Eastern Entomologist, 219, 11–12.
Yakubovich, V.S. (2013) First record of the dragonfly Orthetrum albistylum speciosum (Uhler, 1858) (Odonata: Libellulidae) from Evreiskaya avtonomnaya oblast, Russian Far East. Far Eastern Entomologist, 262, 7–8.
Zafarnejad, Z. (2009) The contribution of dams to Iran’s desertification. International Journal of Environmental Studies, 66, 327–341.
https://doi.org/10.1080/00207230902798648