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Type: Article
Published: 2018-08-20
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Unraveling Ocyptamus and the Baccha legacy (Diptera: Syrphidae): redefinition of groups and new species descriptions

Zoologisches Forschungsmuseum Alexander Koenig, Leibniz-Institut für Biodiversität der Tiere, Adenauerallee 160, D–53113 Bonn, Germany.
Laboratório de Entomologia Sistemática, Urbana e Forense, Coordenação de Biodiversidade, Instituto Nacional de Pesquisas da Amazônia, Avenida André Araujo 2936, Bairro Petrópolis, 69060-060 Manaus, Brazil.
Department of Entomology, National Museum of Natural History, Smithsonian Institution, 10th & Constitution NW, Washington, D.C. 20560-0169, USA.
Diptera hoverflies flower flies species groups systematics

Abstract

The systematics of Ocyptamus sensu lato has been reviewed, its species groups redefined, and some taxa resurrected. These and other related taxa are diagnosed to aid in distinguishing between them. The status of the following genera is revised: Calostigma Shannon stat. rev., Hermesomyia Vockeroth stat. rev., Hybobathus Enderlein stat. rev., Mimocalla Hull stat. rev., Orphnabaccha Hull stat. rev., Pipunculosyrphus Hull stat. rev., Pseudoscaeva Vockeroth stat. rev., and Styxia Hull stat. rev. New species of the Neotropical genus Ocyptamus are described from Costa Rica and Venezuela: Ocyptamus maximus Thompson sp. nov., Ocyptamus myiophagus Thompson sp. nov., and Ocyptamus megafemur Thompson sp. nov. In addition, an identification key for the genera and species groups of Ocyptamus sensu lato is provided, as well as a redescription of Syrphus sargoides Macquart.

Lectotypes are designated for Doros disjuncta Sack, Baccha placiva Williston, Baccha pulla Sack, Ocyptamus funebris Macquart, Syrphus sargoides, and Syrphus immaculatus Macquart. Moreover, Syrphus immaculatus Macquart is considered to be a species of the genus Toxomerus, and new species synonyms are also given: Baccha atypica Curran is a junior synonym of Syrphus conjunctus Wiedemann; Baccha flata Hull and Doros disjuncta Sack are jun. syn. of Syrphus sargoides Macquart; Baccha bromleyi Curran is a jun. syn. of Ocyptamus funebris Macquart; Baccha sepia Hull, Baccha danaida Hull, Baccha violacea Hull, and Baccha ursula Hull are all synonyms of Ocyptamus pullus (Sack); Baccha nerissa Hull is a jun. syn. of Pelecinobaccha hiantha (Hull); and Baccha panamensis Curran is a jun. syn. of Pelecinobaccha transatlantica (Schiner).

 

References

  1. Arcaya, E., Mengual, X., Pérez-Bañón, C. & Rojo, S. (2013) Reports and distribution of predatory syrphids (Diptera: Syrphidae: Syrphinae) in Lara State, Venezuela. Bioagro, 25, 143–148.

    Austen, E.E. (1893) Descriptions of new species of dipterous insects of the family Syrphidae in the collection of the British Museum, with notes on species described by the late Francis Walker - Part I. Bacchini and Brachyopini. Proceedings of the Zoological Society of London, 1893, 132–164.

    Bigot, J.M.F. (1884a) Diptères nouveaux ou peu connus, 22e partie (1), XXXII: Syrphidi (2e partie). Espèces nouvelles, No 1er. Annales de la Société entomologique de France, Series 6, 3, 315–356.

    Bigot, J.M.F. (1884b) Diptères nouveaux ou peu connus, 24e partie (1), XXXII: Syrphidi (2e partie). Espèces nouvelles, No III (2). Annales de la Société entomologique de France, Series 6, 4, 73–80, 81–116.

    Bréthes, J. (1905) Insectos de Tucumán. Anales del Museo Nacional de Buenos Aires, Series 3, 4, 329–347.

    Brown, R.W. (1956) Composition of scientific words a manual of methods and a lexicon of materials for the practice of logotechnics. Published by the author, Baltimore, 882 pp.

    Coquillet, D.W. (1910) The type-species of the North American genera of Diptera. Proceedings of the United States National Museum, 37, 499–647.
    https://doi.org/10.5479/si.00963801.37-1719.499

    Cumming, J.M. & Wood, D.M. (2009) Adult morphology and terminology. In: Brown, B. (Eds.), Manual of Central American Diptera. Vol. 1. NRC Press, Ottawa, pp. 9–50.

    Curran, C.H. (1927) New Neotropical and Oriental Diptera in the American Museum of Natural History. American Museum Novitates, 245, 1–9.

    Curran, C.H. (1928) The Syrphidae of the Malay Peninsula. Journal of the Federated Malay States Museums, 14, 141–324.

    Curran, C.H. (1929) New Syrphidae and Tachinidae. Annals of the Entomological Society of America, 22, 489–510.
    https://doi.org/10.1093/aesa/22.3.489

    Curran, C.H. (1930a) New species of Diptera belonging to the genus Baccha Fabricius (Syrphidae). American Museum Novitates, 403, 1–16.

    Curran, C.H. (1930b) New Syrphidae from Central America and the West Indies. American Museum Novitates, 416, 1–11.

    Curran, C.H. (1930c) New Diptera from North and Central America. American Museum Novitates, 415, 1–16.

    Curran, C.H. (1934) The Diptera of Kartabo, Bartica District, British Guiana, with descriptions of new species from other British Guiana localities. Bulletin of the American Museum of Natural History, 66, 287–532.

    Curran, C.H. (1938) New American Diptera. American Museum Novitates, 975, 1–7.

    Curran, C.H. (1939) New Neotropical Baccha Fabricius (Syrphidae: Diptera). American Museum Novitates, 1041, 1–12.

    Curran, C.H. (1941b) New American Syrphidae. Bulletin of the American Museum of Natural History, 78, 243–304.

    Doesburg, P.H. van (1962) Preliminary list of Syrphidae known from Suriname and British and French Guiana. Studies on the fauna of Suriname and other Guyanas [Natuurwetenschappelijke Studiekring voor Suriname en de Nederlandse Antillen], 5 (28), 1–33.

    Doesburg, P.H. van (1966) Syrphidae from Suriname. Additional records and descriptions. Studies on the fauna of Suriname and other Guyanas [Natuurwetenschappelijke Studiekring voor Suriname en de Nederlandse Antillen], 9 (35), 61–107.

    Enderlein, G. (1938) Beitrag zur Kenntnis der Syrphiden. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1937, 192–237.

    Fabricius, J.C. (1775) Systema entomologiae: sistens insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. Kortii, Flensbvrgi et Lipsiae [= Flensburg & Leipzig], [32] + 832 pp.

    Fabricius, J.C. (1781) Species insectorum exhibentes eorum differentias specificas, synonyma auctorum, loca natalia, metamorphosin, adjectis observationibus, descriptionibus. Tome II. C.E. Bohnii, Hamburgi et Kilonii [= Hamburg and Kiel], 494 pp.

    Fabricius, J.C. (1805) Systema antliatorum: secundum ordines, genera, species adiectis synonymis, locis, observationibus, descriptionibus. C. Reichard, Brunsvigae [=Brunswick], 312 pp. (pp. xiv + 15–372) + [1] + 30 pp.

    Fluke, C.L. Jr. (1936) New Syrphidae (Diptera) from Brazil and Cuba. Journal of the Kansas Entomological Society, 9, 59–65.

    Fluke, C.L. Jr. (1942) Revision of the Neotropical Syrphini related to Syrphus (Diptera, Syrphidae). American Museum Novitates, 1201, 1–24.

    Fluke, C.L. Jr. (1950a) The male genitalia of Syrphus, Epistrophe and related genera (Diptera, Syrphidae). Transactions of the Wisconsin Academy of Sciences, Arts and Letters, 40, 115–148, 10 pls.

    Fluke, C.L. Jr. (1950b) Some new tropical syrphid flies with notes on others. Acta Zoologica Lilloana, 9, 439–454.

    Fluke, C.L. Jr. (1956) Catalogue of the family Syrphidae in the Neotropical Region. Revista Brasileira de Entomologia, 6, 193–268.

    Frey, R. (1946) Ubersicht der Gattungen der Syrphiden-Unterfamilie Syrphinae (Syrphine + Bacchinae). Notulae Entomologicae, 25, 152–172. [1945]

    Giglio-Tos, E. (1892) Diagnosi di nuove specie di Ditteri. VII. Sirfidi del Messico (1). Bollettino dei musei di zoologia ed anatomia comparata della R. Università di Torino, 7 (132), 1–5.

    Goot, V.S. van der (1964) Fluke's catalogue of Neotropical Syrphidae (Insects, Diptera), a critical study with an appendix on new names in Syrphidae. Beaufortia, 10, 212–221.

    Hine, J.S. (1914) Diptera of Middle America. Family Syrphidae. The Ohio Naturalist, 14, 333–343.

    Hull, F.M. (1930) Some new species of Syrphidae (Diptera) from North and South America. Transactions of the American Entomological Society, Philadelphia, 56, 139–148, 1 pl.

    Hull, F.M. (1937) New species of exotic syrphid flies. Psyche, 44, 12–32, pl. 2.

    Hull, F.M. (1940) Some new species of Syrphidae. Journal of the Washington Academy of Sciences, 30, 432–434.

    Hull, F.M. (1941a) Some new syrphid flies from North and South America (Diptera). Entomological News, 52, 157–163.

    Hull, F.M. (1941b) Descriptions of some new species of Syrphidae. Psyche, 48, 149–165.

    Hull, F.M. (1941c) Some species of the genus Baccha from the New World. Proceedings of the Entomological Society of Washington, 43, 181–183.

    Hull, F.M. (1942a) New species of Syrphidae from the Neotropical region. Psyche, 49, 84–107.

    Hull, F.M. (1942b) Some flies of the genus Mesogramma. Proceedings of the New England Zoölogical Club, 30, 17–24.

    Hull, F.M. (1942c) Some new species of Baccha and Mesogramma (Dipt. Syrphidae). Revista de Entomologia, Rio de Janeiro, 13, 44–49.

    Hull, F.M. (1943a) New species of American syrphid flies. Bulletin of the Brooklyn Entomological Society, 38, 48–53.

    Hull, F.M. (1943b) The New World species of the genus Baccha. Entomologica Americana, 23, 42‑99, 10 pls.

    Hull, F.M. (1943c) New species of the genera Baccha and Rhinoprosopa (Syrphidae). Journal of the Washington Academy of Sciences, 33, 214–216.

    Hull, F.M. (1943d) Some undescribed species of the genus Baccha (Syrphidae). Journal of the Washington Academy of Sciences, 33, 72–74.

    Hull, F.M. (1943e) New species of Baccha and related flies. Entomological News, 54, 135–140.

    Hull, F.M. (1943f) New species of syrphid flies in the National Museum. Journal of the Washington Academy of Sciences, 33, 39–43.

    Hull, F.M. (1943g) New species of Syrphidae of the genus Baccha and Mesogramma. Entomological News, 54, 89–91.

    Hull, F.M. (1944a) A study of some syrphid flies of South America. Revista de Entomologia, 15, 34–54.

    Hull, F.M. (1944b) Additional species of the genus Baccha from the New World. Bulletin of the Brooklyn Entomological Society, 39, 56–64.

    Hull, F.M. (1944c) Studies of flower flies (Syrphidae) in the Vienna Museum (Natural History). Journal of the Washington Academy of Sciences, 34, 398–404.

    Hull, F.M. (1944d) Some flies of the family Syrphidae in the British Museum (Natural History). The Annals and Magazine of Natural History, Series 11, 11, 21–61.

    Hull, F.M. (1945) Some undescribed syrphid flies. Proceedings of the New England Zoölogical Club, 23, 71–78.

    Hull, F.M. (1947a) More flies of the genus Baccha (Diptera, Syrphidae). Revista de Entomologia, Rio de Janeiro, 18, 395–410.

    Hull, F.M. (1947b) Some American syrphid flies. Psyche, 54, 230–240.

    Hull, F.M. (1948) Some Neotropical species of syrphids. Entomological News, 59, 1–12.

    Hull, F.M. (1949a) The genus Baccha from the New World. Entomologica Americana, 27 [1947], 89–285, 47 pls.

    Hull, F.M. (1949b) The morphology and inter-relationship of the genera of syrphid flies, recent and fossil. Transactions of the Linnean Society of London, 26, 257–408.

    Hull, F.M. (1949c) Studies upon Diptera in the British Museum. The Annals and Magazine of Natural History, Series 12, 2, 727–746.

    Hull, F.M. (1949d) New species of New World Baccha. The Ohio Journal of Science, 49, 244–246.

    Hull, F.M. (1949e) Some unusual syrphid flies. Entomological News, 60, 225–233.

    Hull, F.M. (1950) New South American syrphid flies. Revista de Entomologia, Brazil, 21, 225–236.

    Hull, F.M. (1957) Some undescribed species of the genus Baccha Fabricius (Diptera: Syrphidae). Journal of the New York Entomological Society, 65, 187–190.

    Hull, F.M. (1958) Three new species of the genus Baccha Fabricius (Diptera: Syrphidae). Studia Entomologica, 1, 427–432.

    Hull, F.M. (1960) New species of Syrphidae and Asilidae (Diptera). The Pan-Pacific Entomologist, 36, 69–74.

    Inouye, D., Larson, B.M.H., Ssymank, A. & Kevan, P.G. (2015) Flies and flowers III: Ecology of foraging and pollination. Journal of Pollination Ecology, 16 (16), 115–133.

    International Code of Zoological Nomenclature (1999) International Code of Zoological Nomenclature. 4th Edtion. The International Trust for Zoological Nomenclature, London, 306 pp.

    Kassebeer, C.F. (2000) Zur Gattung Pseudodoros Becker, 1903 (Diptera: Syrphidae). Dipteron, 3 (1), 73–92.

    Knab, F. (1914) Two North American Syrphidae. Insecutor inscitiae menstruus, 2, 151–153.
    https://doi.org/10.5962/bhl.part.9590

    Larson, B.M.H., Kevan, P.G. & Inouye, D.W. (2001) Flies and flowers: taxonomic diversity of anthophiles and pollinators. Canadian Entomologist, 133, 439–465.
    https://doi.org/10.4039/Ent133439-4

    Loew, H. (1861) Diptera aliquot in insula Cuba collecta. Wiener entomologische Monatschrift, 5, 33–43.

    Loew, H. (1863) Diptera Americae septentrionalis indigena. Centuria tertia. Berliner entomologische Zeitschrift, 7, 1–55.
    https://doi.org/10.1002/mmnd.18630070104

    Loew, H. (1866) Diptera Americae septentrionalis indigena. Centuria septima. Berliner entomologische Zeitschrift, 10, 1–54.
    https://doi.org/10.1002/mmnd.18660100103

    Löhr, B., Varela, A.M. & Santos, B. (1990) Exploration for natural enemies of the cassava mealybug, Phenacoccus manihoti (Homoptera: Pseudococcidae), in South America, for the biological control of this introduced pest in Africa. Bulletin of Entomological Research, 80, 417–425.
    https://doi.org/10.1017/S0007485300050677

    Lynch Arribálzaga, F. (1891–1892). Dipterología Argentina, Syrphidae. Anales de la Sociedad Científica Argentina, 32–34, 80–99 + 118–131 + 194–202 + 247–256 + 307–314, 51–58 + 111–121 + 189–199 + 236–253, 33–46 + 173–192 + 242–280.

    Macquart, P.J.M. (1834) Histoire naturelle des insectes. Diptères. Tome premiere. Roret, Paris, 578 + 8 pp., 12 pls.

    Macquart, P.J.M. (1842) Diptères exotiques nouveaux ou peu connus. Tome deuxième, 2.e partie. Mémoires de la Société Royal des sciences, de l'agriculture et des arts de Lille, 1841 (1), 65–200, 22 pls.

    Macquart, P.J.M. (1846) Diptères exotiques nouveaux ou peu connus. Supplément [1]. Mémoires de la Société Royal des sciences, de l'agriculture et des arts de Lille, 1844, 133–364, 20 pls.

    Macquart, P.J.M. (1850) Diptères exotiques nouveaux ou peu connus. 4.e supplément. Mémoires de la Société Royal des sciences, de l'agriculture et des arts de Lille, 1849, 309–479, pls. 1–14.

    Macquart, P.J.M. (1855) Diptères exotiques nouveaux ou peu connus. 5.e supplément. Mémoires de la Société Royal des sciences, de l'agriculture et des arts de Lille, Series 2, 1, 25–156, 7 pls.

    Matsumura, S. & Adachi, J. (1917) Synopsis of the economic Syrphidae of Japan. (Pt II). The Entomological Magazine, Kyoto, 2 (4), 133–156, pl. 6.

    Meijere, J.C.H. de (1908) Studien über südostasiatische Dipteren. III. Tijdschrift voor entomologie, 51, 191–332, pls. 7–8.

    Mengual, X. & Thompson, F.C. (2011) Carmine cochineal killers: the flower fly genus Eosalpingogaster Hull (Diptera: Syrphidae) revised. Systematic Entomology, 36, 713–731.
    https://doi.org/10.1111/j.1365-3113.2011.00588.x

    Mengual, X. (2011) Black-tie dress code: two new species of the genus Toxomerus (Diptera, Syrphidae). ZooKeys, 140, 1–26.

    Mengual, X. (2015a) The systematic position and phylogenetic relationships of Asiobaccha Violovitsh (Diptera, Syrphidae). Journal of Asia-Pacific Entomology, 18, 397–408.

    Mengual, X. (2015b) Revision of the genus Rhinoprosopa (Diptera: Syrphidae). The Canadian Entomologist, 147, 1–22.

    Mengual, X., Ståhls, G. & Rojo, S. (2008a) First phylogeny of predatory hoverflies (Diptera, Syrphidae, Syphinae) using mitochondrial COI and nuclear 28S rRNA genes: conflict and congruence with the current tribal classification. Cladistics, 24, 543–562.
    https://doi.org/10.1111/j.1096-0031.2008.00200.x

    Mengual, X., Ståhls, G. & Rojo, S. (2008b) Molecular phylogeny of Allograpta (Diptera, Syrphidae) reveals diversity of lineages and non-monophyly of phytophagous taxa. Molecular Phylogenetics and Evolution, 49, 715–727.
    https://doi.org/10.1016/j.ympev.2008.09.011

    Mengual, X., Ståhls, G. & Rojo, S. (2012) Is the mega-diverse genus Ocyptamus (Diptera, Syrphidae) monophyletic? Evidence from molecular characters including the secondary structure of 28S rRNA. Molecular Phylogenetics and Evolution, 62, 191–205.
    https://doi.org/10.1016/j.ympev.2011.09.014

    Mengual, X., Ståhls, G. & Rojo, S. (2015) Phylogenetic relationships and taxonomic ranking of pipizine flower flies (Diptera: Syrphidae) with implications for the evolution of aphidophagy. Cladistics, 31, 491–508.
    https://doi.org/10.1111/cla.12105

    Mengual, X., Ståhls, G., Láska, P., Mazánek, L. & Rojo, R. (2018) Molecular phylogenetics of the predatory lineage of flower flies Eupeodes-Scaeva (Diptera: Syrphidae), with the description of the Neotropical genus Austroscaeva gen. nov. Journal of Zoological Systematics and Evolutionary Research, 56, 148–169.
    https://doi.org/10.1111/jzs.12212

    Miranda, G.F.G. (2017) Revision of the Hybobathus arx and Pelecinobaccha summa species groups (Diptera: Syrphidae). Zootaxa, 4338 (1), 001–043.
    https://doi.org/10.11646/zootaxa.4338.1.1

    Miranda, G.F.G., Marshall, S.A. & Skevington, J.H. (2014) Revision of the genus Pelecinobaccha Shannon, description of Relictanum gen. nov. and redescription of Atylobaccha flukiella (Curran, 1941). Zootaxa, 3819 (1), 1–154.
    https://doi.org/10.11646/zootaxa.3819.1.1

    Miranda, G.F.G., Skevington, J.H., Marshall, S.A. & Kelso, S. (2016) The genus Ocyptamus Macquart (Diptera: Syrphidae): A molecular phylogenetic analysis. Arthropod Systematics and Phylogeny, 74, 161–176.

    Montoya, A.L. & Pérez, S.P. (2009) Primer registro de Ocyptamus arx Fluke 1936 (Diptera: Syrphidae: Syrphinae) depredando ninfas de Antianthe expansa Germar 1835 [sic] (Hemiptera: Membracidae). In: Bustillo Pardy, A.E. (Ed.), XXXVI Congreso de la Sociedad Colombiana de Entomología. Resúmenes. Sociedad Colombiana de Entomología, Medellín, pp. 130.

    NRC [=National Research Council of the National Academies] (2007) Status of Pollination in North America. The National Academies Press, Washington, D.C., 322 pp.

    Osten Sacken, C.R. (1877) Western Diptera: Descriptions of new genera and species of Diptera from the region west of the Mississippi and especially from California. Bulletin of the United States Geological and Geographical Survey of the Territories, 3, 189–354.
    https://doi.org/10.5962/bhl.title.57939

    Pape, T. & Thompson, F.C. (2013) Systema Dipterorum. Version 1.5. Available from: http://www.diptera.org (accessed 8 March 2017)

    Pérez, D. & Iannacone, J. (2009) Fluctuación y distribución espacio-temporal de Tuthillia cognata (Hemiptera, Psyllidae) y de Ocyptamus persimilis (Diptera, Syrphidae) en el cultivo de camu-camu Myrciaria dubia (Myrtaceae) en Ucayali, Perú. Revista Brasileira de Entomologia, 53, 635–642.
    https://doi.org/10.1590/S0085-56262009000400014

    Pérez-Bañón, C., Arcaya, E., Mengual, X. & Rojo, S. (2013) Preimaginal morphology of the genera Salpingogaster Schiner, 1868 and Eosalpingogaster Hull, 1949 (Diptera: Syrphidae), with its systematic implications. Zootaxa, 3599 (4), 361–370.

    Pérez-Bañón, C., Juan, A., Petanidou, T., Marcos-García, M.A. & Crespo, M.B. (2003) The reproductive ecology of Medicago citrina (Font Quer) Greuter (Leguminosae): a bee-pollinated plant in Mediterranean islands where bees are absent. Plant Systematics and Evolution, 241, 29–46.

    Philippi, R.A. (1865) Aufzählung der chilenischen Dipteren. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, 15, 595–782.

    Prado, E., Santa-Cecília, L.V.C., Reis, P.R. & Mosca, E. (2015) The Coffee Red Mealybug, Nipaecoccus coffeae (Hempel) (Hemiptera: Pseudococcidae), an uncommon mealybug attacking coffee plants in Brazil. Check List, 11 (4), 1694.
    https://doi.org/10.15560/11.4.1694

    Reemer, M. (2010) A second survey of Surinam Syrphidae (Diptera): introduction and Syrphinae. Tijdschrift voor Entomologie, 153, 163–196.
    https://doi.org/10.1163/22119434-900000295

    Reemer, M. (2012) Natural history of Microdontinae (Diptera: Syrphidae): a review. In: Reemer, M. (Ed), Unravelling a hotchpotch: phylogeny and classification of the Microdontinae (Diptera: Syrphidae). PhD dissertation, Universiteit Leiden, Leiden, 384 pp. Avaliable from: https://openaccess.leidenuniv.nl/bitstream/handle/1887/18582/02.pdf?sequence=7 (Accessed 17 Aug. 2018)

    Röder, V. von (1885) Dipteren von der Insel Portorico. Stettiner Entomologische Zeitung, 46, 337–349.

    Rojo, S., Gilbert, F., Marcos-García, M.A, Nieto, J.M. & Mier, M.P. (2003) A world review of predatory hoverflies (Diptera, Syrphidae: Syrphinae) and their prey. Centro Iberoamericano de la Biodiversidad (CIBIO), Alicante, 278 pp.

    Rotheray, G.E. & Gilbert F. (2011) The natural history of hoverflies. Forrest Text, Ceredigion, 333 pp.

    Rotheray, G.E., Zumbado, M.A., Hancock, E.G. & Thompson, F.C. (2000) Remarkable aquatic predators in the genus Ocyptamus (Diptera, Syrphidae). Studia Dipterologica, 7, 385–398.

    Sack, P. (1921) Dr. L. Zurcher’s Dipteren - Ausbeute aus Paraguay: Syrphiden. Archiv fur Naturgeschichte, Abteilung A, 87, 127–149.

    Say, T. (1823) Descriptions of dipterous insects of the United States. Journal of the Academy of Natural Sciences of Philadelphia, 3, 9–32, 33–64, 73–104.

    Say, T. (1829–1830) Descriptions of North American dipterous insects. Journal of the Academy of Natural Sciences of Philadelphia, 6, 149–178, 183–188.

    Schiner, I.R. (1868) Diptera. In: Wüllerstorf-Urbair, B. von (in charge), Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wüllerstorf-Urbair. Vol. 2. Band 1. Abtheilung B. Kaiserlich-königliche Hof- und Staatsdruckerei in Commission bei Karl Gerold's Sohn, Wien, pp. i–vi + 1–388, 4 pls.

    Shannon, R.C. (1927) A review of the South American two-winged flies of the family Syrphidae. Proceedings of the United States National Museum, 70 (9), No. 2658, 1–34.
    https://doi.org/10.5479/si.00963801.70-2658.1

    Ssymank, A. & Kearns, C. (2009) Flies-pollinators on two wings. In: Ssymank, A., Hamm, A. & Vischer-Leopold, M. (Eds.), Caring for pollinators—safeguarding agro-biodiversity and wild plant diversity. Bundesamt für Naturschutz, Bonn, pp. 39–52.

    Ssymank, A., Kearns, C.A., Pape, T. & Thompson, F.C. (2008) Pollinating flies (Diptera): A major contribution to plant diversity and agricultural production. Biodiversity, 9, 86–89.
    https://doi.org/10.1080/14888386.2008.9712892

    Stahl, A. (1882) Catálogo del gabinete zoológico del Dr. A. Stahl en Bayamón (Pto.-Rico) precedido de una clasificación sistemática de los animales que corresponden á esta fauna. Impresora del Boletín Mercantil, Puerto Rico, 248 pp.

    Telford, H.S. (1973) The Syrphidae of Puerto Rico. The Journal of Agriculture of the University of Puerto Rico, 57, 217–246.

    Thompson, F.C. & Rotheray, G. (1998) Family Syrphidae. In: Papp, L. & Darvas, B. (Eds.), Contributions to a manual of Palaearctic Diptera (with special reference to flies of economic importance), Vol. 3. Higher Brachycera. Science Herald, Budapest, pp. 81–139.

    Thompson, F.C. & Zumbado, M.A. (2000) Flower flies of the subgenus Ocyptamus (Mimocalla Hull) (Diptera: Syrphidae). Proceedings of the Entomological Society of Washington, 102, 773–793.

    Thompson, F.C. (1981) The flower flies of the West Indies (Diptera: Syrphidae). Memoirs of the Entomological Society of Washington, 9, 1–200.

    Thompson, F.C. (1999) A key to the genera of the flower flies of the Neotropical Region including the descriptions of genera and species and a glossary of taxonomic terms. Contributions on Entomology, International, 3, 319–378.

    Thompson, F.C. (2006) Primer Taller de Identificación Syrphidae del Neotrópico. Universidad del Valle, Cali, Colombia, 618 pp.

    Thompson, F.C. (2013) Syrphidae. Systema Dipterorum, version 1.5, 13354 records. Available from: http://www.diptera.org (accessed 8 March 2017)

    Thompson, F.C., Rotheray, G.E. & Zumbado, M. (2010) Family Syrphidae. In: Brown, B. (Eds.), Manual of Central American Diptera. Vol. 2. NRC Press, Ottawa, pp. 763–792.

    Thompson, F.C., Vockeroth, J.R. & Sedman, Y.S. (1976) Family Syrphidae. A catalogue of the Diptera of the American South of the United States, 46, 1–195.
    https://doi.org/10.5962/bhl.title.49898

    Ureña, O. & Hanson, P. (2010) A fly larva (Syrphidae: Ocyptamus) that preys on adult flies. Revista de Biologia Tropical, 58, 1157–1163.

    Violovitsh, N.A. (1976) Survey on species of genus Baccha Fabricius, 1805 (Diptera, Syrphidae) from the Palaearctic fauna. Novye i maloizvestnye vidy fauny Sibiri, 10, 130–154. [in Russian]

    Vockeroth, J.R. (1969) A revision of the genera of the Syrphini (Diptera: Syrphidae). Memoirs of the Entomological Society of Canada, 62, 1–176.
    https://doi.org/10.4039/entm10162fv

    Walker, F. (1836) Descriptions, & c. of the Diptera. In: Curtis, J., Haliday, A.H. & Walker, F. (Eds.), Descriptions, & c. of the Insects collected by Captain P. P. King, R. N., F. R. S., in the Survey of the Straits of Magellan. Transactions of the Linnean Society of London, 17, pp. 315–359.
    https://doi.org/10.1111/j.1095-8339.1834.tb00026.x

    Walker, F. (1849) List of the specimens of dipterous insects in the collection of the British Museum. Part II, III & IV. British Museum (Natural History), London, 949 pp. [Part II: [iii] + 231–484, Part III: [iii] + 485–687, Part IV: pp. [3] + 689–1172 + [2]]

    Walker, F. (1852) Diptera. Part III. In: Saunders, W.W. (Eds.), Insecta Saundersiana: or characters of undescribed insects in the collection of William Wilson Sauders, Esq., F.R.S., F.L.S., &c. Vol. 1. Van Voorst, London, pp. 157–252, pls. 5–6.

    Walker, F. (1857–1858) Characters of undescribed Diptera in the collection of W. W. Saunders, Esq., F.R.S., &c. Transactions of the Entomological Society of London, New Series, 4, 119–158, 190–235.

    Wiedemann, C.R.W. (1830) Aussereuropäische Zweiflügelige Insekten. Zweiter Theil. Schulz, Hamm, xii + 684 pp., 5 pls.

    Williston, S.W. (1888) Diptera Brasiliana, ab H. H. Smith collecta. Part I, Stratiomyidae, Syrphidae. Transactions of the American Entomological Society, 15, 243–292.
    https://doi.org/10.2307/25076505

    Williston, S.W. (1891–1892) Fam. Syrphidae. Biologia Centrali-Americana, Insecta: Diptera, III, 1–56 + 57–72 + 73–78, pls. 1 + 2, figs. 1–12.

    Wirth, W.W., Sedman, Y.S. & Weems, H.V. Jr. (1965) Family Syrphidae. In: Stone, A., Sabrosky, C.W., Wirth, W.W., Foote, R.H. & Coulson, J.R. (Eds.), A Catalog of the Diptera of America north of Mexico. United States Department of Agriculture, Washington D.C., pp. 557–625.

    Woodcock, T.S., Larson, B.M.H., Kevan, P.G., Inouye, D.W. & Lunau, K. (2014) Flies and flowers II: Floral rewards and attractants. Journal of Pollination Ecology, 12, 63–94.