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Type: Monograph
Published: 2018-10-01
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Names of hydroids (Cnidaria, Hydrozoa) established by Charles McLean Fraser (1872–1946), excluding those from Allan Hancock Expeditions

Department of Natural History, Royal Ontario Museum, 100 Queen’s Park, Toronto, Ontario, Canada M5S 2C6. Research Associate, Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, Canada V8W 9W2.
Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, Canada V8W 9W2. Research Associate, Department of Natural History, Royal Ontario Museum, 100 Queen’s Park, Toronto, Ontario, Canada M5S 2C6.
Protist bibliography Hydroidolina invertebrate zoology marine biology Medusozoa museum collections natural history taxonomy zoological nomenclature

Abstract

An account is given of the names of families, genera, and species of hydroids established by C.M. Fraser of Canada, excluding those from Allan Hancock Pacific Expeditions and the Allan Hancock Caribbean Sea Expedition. The names of four families, 11 genera, and 101 species are attributed to him in this work, complementing those of another two genera and 164 species described by Fraser in various Allan Hancock publications. Information is provided on type localities of his new species, on locations and kinds of type material in existence, where known, and on the current taxonomic status of families, genera, and species described by him in works reviewed herein. Two replacement names (Thuiaria geniculata Fraser, 1918a and Salacia fraseri Calder, 1991) exist for Thuiaria distans Fraser, 1914a (not Thuiaria distans Allman, 1877). The widely overlooked senior synonym, T. geniculata, is recognized as the valid name of the species. Diphasia alta nom. nov. is proposed as a new replacement name for the permanently invalid junior primary homonym Diphasia robusta Fraser, 1943a (not D. attenuata robusta Billard, 1924). In addition, Cryptolaria crassa nom. nov. is proposed as a new replacement name for the junior secondary homonym Cryptolaria rigida (Fraser, 1948) [not C. rigida (Fraser, 1940a)]. Lectotypes have been designated for 47 of the species to establish objective standards for application of their names. Corymorpha adventitia Fraser, 1941b, from the Pacific coast of Panama, is assigned to Ralpharia Watson, 1980, as R. adventitia. No type specimens are known to exist for seven of the species considered, including one holotype (of Hebella eximia Fraser, 1944a) missing from its bottle. Particular attention has been paid to dating and chronology of the 51 publications of Fraser covered in this work. Earlier bibliographic errors are corrected, most notably establishing that the book Distribution and relationship in American hydroids was published in early 1947 rather than 1946 as per the title page.

 

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    Fraser, C.M. (1922a) A new Hydractinia and other west coast hydroids. Contributions to Canadian Biology, New Series, 1, 95–100. [24 July 1922: noted in Contents (page iii) of the journal]

    Fraser, C.M. (1922b) Hydroids of the Canadian Arctic Expedition, 1913–1918. Report of the Canadian Arctic Expedition 1913–1918, 8(I), 1–5. [24 August 1922: recorded on cover of article]

    Fraser, C.M. (1924a) Canadian Atlantic fauna. 3a. Hydroida. Hydroids. Publications of the Biological Board of Canada, Ottawa, 3–46. [This article, a reprint of parts of the paper above by Fraser (1921), is dated “May 13, 1924” in copy at the Gerstein Science Information Centre, University of Toronto]

    Fraser, C.M. (1924b) Acaulis primarius Stimpson. Transactions of the Royal Society of Canada, Series 3, Section 5, 18, 167–173. [Read May 1924; 1924]

    Fraser, C.M. (1925) Some new and some previously unreported hydroids, mainly from the Californian coast. University of California Publications in Zoology, 28, 167–172. [16 December 1925: noted in publication data at beginning of article]

    Fraser, C.M. (1926) Hydroids of the Miramichi estuary collected in 1918. Transactions of the Royal Society of Canada, Series 3, Section 5, 20, 209–214. [Read May 1926; 1926]

    Fraser, C.M. (1927) The hydroids of the Cheticamp Expedition of 1917. Contributions to Canadian Biology and Fisheries, new series, 3, 325–329. [20 October 1927: noted in Contents of the journal]

    Fraser, C.M. (1931) Biological and oceanographic conditions in Hudson Bay. 3. Hydroids of Hudson Bay and Hudson Strait. Contributions to Canadian Biology and Fisheries, new series, 6: 475–481. [A note records “Received for publication April 2, 1931”; no publication date recorded other than the year 1931; taken to be 31 December 1931 under ICZN Art. 21.3.2]

    Fraser, C.M. (1932) A comparison of the marine fauna of the Nanaimo region with that of the San Juan Archipelago. Transactions of the Royal Society of Canada, Series 3, Section 5, 26, 49–70. [Read May 1932; 1932]

    Fraser, C.M. (1933a) Some Greenland hydroids. Journal of the Washington Academy of Sciences, 23, 563–566. [15 December 1933: noted at top of alternate pages of issue 12, Vol. 23 of the journal]

    Fraser, C.M. (1933b) Hydroids as a food supply. Transactions of the Royal Society of Canada, Series 3, Section 5, 27, 259–264. [read 19 May 1933; 1933]

    Fraser, C.M. (1935a) Hydroids from the west coast of Vancouver Island. Canadian Field-Naturalist, 49, 143–145. [10 December 1935: issue date of Vol. 49, No. 9 marked on the journal]

    Fraser, C.M. (1935b) Some Japanese hydroids, mostly new. Transactions of the Royal Society of Canada, Series 3, Section 5, 29, 105–112. [read 23 May 1935; 1935]

    Fraser, C.M. (1936a) Hydroids from the Queen Charlotte Islands. Journal of the Biological Board of Canada, 1, 503–507. [13 February 1936, from entry in Contents page of the journal]
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    Fraser, C.M. (1936b) Hydroid distribution in the vicinity of the Queen Charlotte Islands. Canadian Field-Naturalist, 50, 122–126. [1 October 1936: issue date of Vol. 50, No. 7 marked on the journal]

    Fraser, C.M. (1936c) Some Japanese hydroids, mostly new. Π. Transactions of the Royal Society of Canada, Series 3, Section 5, 30, 49–54. [read May 1936; 1936]

    Fraser, C.M. (1937a) Hydroids of the Pacific coast of Canada and the United States. University of Toronto Press, Toronto, 207 pp. [a copy at the Gerstein Science Information Centre, University of Toronto, bears the hand-written date “14.9.37”]

    Fraser, C.M. (1937b) New species of hydroids from the Puerto Rican region. Smithsonian Miscellaneous Collections, 91 (28), 1–7. [10 November 1937: noted on title page]

    Fraser, C.M. (1937c) Distribution of marine organisms. Canadian Field-Naturalist, 51, 132–134. [8 December 1937: issue date of Vol. 51, No. 9 marked on the journal]

    Fraser, C.M. (1938a) Hydroids of the 1934 Allan Hancock Pacific Expedition. Allan Hancock Pacific Expeditions, 4 (1), 1–105. [August 1938: noted in publication data facing page 1]

    Fraser, C.M. (1938b) The relation of the marine fauna to the physiography of the west coast of the Queen Charlotte Islands. Canadian Field-Naturalist, 52, 88–93. [1 September 1938: issue date of Vol. 52, No. 6 marked on the journal]

    Fraser, C.M. (1938c) Hydroids of the 1936 and 1937 Allan Hancock Pacific Expeditions. Allan Hancock Pacific Expeditions, 4 (2), 107–127. [20 October 1938: noted in publication data facing page 107]

    Fraser, C.M. (1938d) Hydroids of the 1932, 1933, 1935, and 1938 Allan Hancock Pacific Expeditions. Allan Hancock Pacific Expeditions, 4 (3), 129–153. [20 October 1938: noted in publication data facing page 129]

    Fraser, C.M. (1938e) Hydroid distribution in the north-eastern Pacific. Transactions of the Royal Society of Canada, Series 3, Section 5, 32, 39–42. [read 26 May 1938; 1938]

    Fraser, C.M. (1939a) Distribution of the hydroids in the collections of the Allan Hancock Expeditions. Allan Hancock Pacific Expeditions, 4 (4), 155–178. [6 February 1939: noted in publication data facing page 155]

    Fraser, C.M. (1939b) Hydroids of the western Canadian Arctic region, 1935–1937. Canadian Journal of Research, Section D, Zoological Sciences, 17 (3), 59–61. [March 1939: noted on the journal]
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    Fraser, C.M. (1939c) Hydroid notes. Perigonimus pugetensis Heath. Transactions of the Royal Society of Canada, Series 3, Section 5, 33, 23–26. [read 24 May 1939; 1939]

    Fraser, C.M. (1939d) Hydroid notes. A pelagic hydroid from Friday Harbor. Transactions of the Royal Society of Canada, Series 3, Section 5, 33, 26–28. [read 24 May 1939; 1939]

    Fraser, C.M. (1940a) Seven new species and one new genus of hydroids, mostly from the Atlantic Ocean. Proceedings of the United States National Museum, 88, 575–580. [13 September 1940: noted in Contents (page IV) of the journal]
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    Fraser, C.M. (1940b) Some hydroids from the California coast, collected in 1939. Transactions of the Royal Society of Canada, Series 3, Section 5, 34, 39–44. [read 21 May 1940; 1940]

    Fraser, C.M. (1941a) Hydroid distribution in the Pacific. Proceedings of the Sixth Pacific Science Congress, 3 (Oceanography and Marine Biology), 495–500. [listed as “1940” in Vervoort (1995) and as “1941” in Schmitt (1948). In Catalogue, University of California Press Publications, University of California Press, Berkeley and Los Angeles, 1940, 247 pp., an inserted and typewritten page numbered “10a” gives a date of “April 9, 1941” for Vol. III of these proceedings (and “January 18, 1941” for Vols. I and II)]

    Fraser, C.M. (1941b) New species of hydroids, mostly from the Atlantic Ocean, in the United States National Museum. Proceedings of the United States National Museum, 91, 77–89. [14 November 1941: noted in Contents page of the journal]
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    Fraser, C.M. (1943a) Distribution records of some hydroids in the collection of the Museum of Comparative Zoölogy at Harvard College, with description of new genera and new species. Proceedings of the New England Zoölogical Club, 22, 75–98. [19 July 1943: noted on separate in ROM library]

    Fraser, C.M. (1943b) Relationship in North American families of gymnoblastic hydroids. Transactions of the Royal Society of Canada, Series 3, Section 5, 37, 29–33. [read 25 May 1943; 1943]

    Fraser, C.M. (1944a) Hydroids of the Atlantic coast of North America. University of Toronto Press, Toronto, 451 pp. [published no later than 19 July 1944: W.J. Dunlop (University of Toronto Press) to S.J. Cook (National Research Council, Canada), letter dated June 8, 1944, “We have almost finished our work on Hydroids of the Atlantic coast. The sheets are in the bindery and we should begin shipping before long.”—C.M. Fraser to W.J. Dunlop, letter dated July 19, 1944, “The six copies of ‘Hydroids of the Atlantic coast of North America’ arrived safely.” Both letters in University of Toronto Archives and Records Management Services, University of Toronto Press Fonds, B1998-0025, Box 5]

    Fraser, C.M. (1944b) Nematophores in American hydroids. Transactions of the Royal Society of Canada, Series 3, Section 5, 38, 33–49. [read 31 May 1944; 1944]

    Fraser, C.M. (1945a) Notes on some recently collected hydroids in the United States National Museum, with descriptions of three new species. Journal of the Washington Academy of Sciences, 35, 21–23. [Vol. 35, No. 1 of the journal containing this article is marked “January 15, 1945,” but the actual publication date, given on a page following the title page, was 20 January 1945]

    Fraser, C.M. (1945b) On the hydroid Dahlgrenella farcta Miles. Transactions of the Royal Society of Canada, Series 3, Section 5, 39, 61–69. [read 23 May 1945; 1945]

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    Fraser, C.M. (1947c) The medusa Gonionemus. Canadian Field-Naturalist, 60, 119–122. [Vol. 60, No. 6 of the journal containing this article, for November-December 1946, was not issued until 1947: a copy on BHL from the MCZ is marked “Oct 14 1947”]

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