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Issue: Vol. 4493 No. 1: 4 Oct. 2018
Type: Monograph
Published: 2018-10-03
DOI: 10.11646/zootaxa.4493.1.1
Page range: 1–69
Abstract views: 247
PDF downloaded: 5

The cicadas (Hemiptera: Cicadidae) of Panama including the description of six new species, three new combinations, one new synonymy, and nine new records

Department of Biology, Barry University, 11300 NE Second Avenue, Miami Shores, FL 33161–6695, USA.
Hemiptera taxonomy new species Neotropics Central America

Abstract

The known cicada fauna of Panama is identified. Procollina quadrimaculata n. sp., P. stigmosa n. sp., Guyalna woldai n. sp., Herrera nigratorquata n. sp., H. sigillata n. sp. and Conibosa megalopercula n. sp. are described as new. Proarna germari Distant, 1905 n. syn. is shown to be a junior synonym of Proarna invaria (Walker, 1850). Pacarina championi (Distant, 1881) is returned to Proarna Stål, 1864 to become Proarna championi Distant, 1881 n. comb. again. The first records of Proarna invaria (Walker, 1850), Guyalna bogotana (Distant, 1892), Dorisiana cachla (Distant, 1899), Ollanta modesta (Distant, 1881), Pacarina schumanni Distant, 1905, Majeorona truncata Goding, 1925, Herrera lugubrina lugubrina (Stål, 1864), Calyria cuna (Walker, 1850), and Calyria telifera (Walker, 1858) are provided. The records for Ollanta modesta (Distant, 1881) are the first records of the genus Ollanta Distant, 1905 for Panama. The records for Calyria cuna (Walker, 1850), and Calyria telifera (Walker, 1858) are the first record of the genus Calyria Stål, 1862 and the Tribe Parnisini Distant, 1905 in Panama. Previous records of Dorisiana metcalfi Sanborn & Heath, 2014 (= Cicada viridis Olivier, 1790), Carineta fasciculata (Germar, 1830), and Selymbria stigmatica (Germar, 1834) are considered to be misidentifications of G. bogotana, C. maculosa (Torres, 1948), and S. pluvialis (Ramos & Wolda 1985) respectively so that D. metcalfi, C. fasciculata, and S. stigmatica are removed from the cicada fauna of Panama. The Panamanian record of Dorisiana semilata (Walker, 1850) is shown to be a mistake and the species is removed from the faunal list as well. The currently known Panamanian cicada fauna is comprised of 54 described species along with six species from the literature that remain undetermined from 22 genera, seven tribes and three subfamilies. Bergalna xanthospila (Germar, 1830) is reassigned to Dorisiana Metcalf, 1952 to become Dorisiana xanthospila (Germar, 1830) n. comb. Fidicinoides flavibasalis (Distant, 1905) is reassigned to Guyalna Boulard & Martinelli, 1996 to become Guyalna flavibasalis (Distant, 1905) n. comb.

 

References

  1. Amyot, C.J.B. & Audinet-Serville, A. (1843) Histoire Naturelle des Insectes. Hemiptères. Deuxime partie. Homoptères. Homoptera Latr. Librairie encyclopédique de Roret, Paris, 675 pp.

    Apolinar, M. (1946) Vocabulario de términas vulgares en historia natural colombiana. Revista de la Academia Colombiana de Ciencias Exactos, Físicas y Naturales, 6, 467–472.

    Audouin, J.V. & Brullé, G.A. (1835) Histoire naturelle des insectes, traitant de leur organization et de leurs moeurs en general, et comprenant leur classification et la description des espèces. Le tout accompagné de planches gravées sur acier, d’après des peintures executes pour cette edition sur la collection du Muséum de Paris. F.D. Pillot, Paris, 56 pls.

    Bartholomew, G.A. & Barnhart, M.C. (1984) Tracheal gases, respiratory gas exchange, body temperature and flight in some tropical cicadas. Journal of Experimental Biology, 111, 131–144.

    Berg, C. (1879) Hemiptera Argentina (Continuacion.) Hemiptera Homoptera Latr. Anales de la Sociedad Cientifica Argentina, 8, 135–144.

    Blanchard, E. (1840) Homoptères. Histoire naturelle des insectes Orthoptères, Névroptères, Hémiptères, Lépidoptères et Diptères, 3, 1–672, 71 pls.

    Bonaccorso, F.J. (1979) Foraging and reproductive ecology in a Panamanian bat community. Bulletin of the Florida State Museum, Biological Sciences, 24, 359–408.

    Bonarelli, G. (1893) Osservazioni sul Toarciano e l’Aleniano dell’ Appennino centrale. Bolletino della Società geologica Italiana, 12 (2), 195–254.

    Boulard, M. (1976) Sur une deuxime cigale africaine depourvue d'appareil sonore (Homoptera). Bulletin de l'Institute Fondamental d'Afrique Noire, Séries A, Sciences Naturelles, 37, 629–636.

    Boulard, M. (1998) Aperçu de cicadologie antillo-guyanaise. 123e Congrès national des sociétés historiques et scientifiques, Antilles-Guyane, 1998, Histoire naturelle, 1998, 75–94.

    Boulard, M. (1999) Postures de cymbalisation, cymbalisations et cartes d’identité acoustique des cigales 3.—Espèces tropicales des savanes et milieux ouverts (Cicadoidea, Cicadidae et Tibicinidae). Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea, 11/12, 77–116.

    Boulard, M. (2008) Statuts et éthologies sonores de quatre cigales malgaches, trois sont nouvelles et deux représentent un nouveau genre et une nouvelle sous-tribu (Rhynchota, Auchenorhyncha Cicadidae). Lambillionea, CVIII, 161–178.

    Boulard, M. & Martinelli, N.M. (1996) Révision des Fidicini, nouveau statut de la tribu, espèces connues et nouvelles espèces (Cicadomorpha, Cicadidae, Cicadinae). Premiere partie: Sous-tribu nouvelle des Fidicinina. Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea, 9, 11–81.

    Boulard, M. & Martinelli, N.M. (2011) Nouvelles cigales néotropicales de la sous-tribu des Guyalinina (Rhynchota, Cicadidae, Cicadinae, Fidicinini). Lambillionea, 61, 219–232.

    Buckton, G.B. (1889) Note on the classification of Cicadae. Entomologist, 22, 269–270.

    Davis, W.T. (1923) Notes on North American cicadas with descriptions of new species. Journal of the New York Entomological Society, 31, 1–15.

    Davis, W.T. (1938) New North American cicadas with notes on described species. Journal of the New York Entomological Society, 46, 291–310.

    Davis, W.T. (1939) New cicadas from North Carolina and the West Indies. Journal of the New York Entomological Society, 47, 287–302.

    Delétang, L.F. (1919) Contribución al estudio de los Cicádidos (Cicadidae) argentinos (Hemiptera-Homoptera) ensayo filogenético. Anales de la Sociedad Cientifica Argentina, 88, 25–94.

    Distant, W.L. (1881) Rhynchota: Homoptera. Biologia Centrali-Americana; contributions to the knowledge of the fauna and flora of Mexico and Central America, Part 15 (1), 1–16.

    Distant, W.L. (1883) Rhynchota: Homoptera. Biologia Centrali-Americana; contributions to the knowledge of the fauna and flora of Mexico and Central America, Part 15 (1), 17–24.

    Distant, W.L. (1892) On some undescribed Cicadidae, with synonymical notes. Annals and Magazine of Natural History, Series 6, 10, 54–67.
    https://doi.org/10.1080/00222939208677373

    Distant, W.L. (1899) Some apparently undescribed Neotropical Homoptera. Annals and Magazine of Natural History, Series 7, 4, 112–113.
    https://doi.org/10.1080/00222939908678169

    Distant, W.L. (1900) Rhynchota: Homoptera. Biologia Centrali-Americana; contributions to the knowledge of the fauna and flora of Mexico and Central America, Part 15 (1), 41–43.

    Distant, W.L. (1903) Additons to the Rhynchotal fauna of Central America. Transaction of the Royal Entomological Society of London, 1903, 525–527.

    Distant, W.L. (1905a) Cicadidae and Fulgoridae. Biologia Centrali-Americana; contributions to the knowledge of the fauna and flora of Mexico and Central America, Part 15 (1), 140–146.

    Distant, W.L. (1905b) Rhynchotal notes XXXI. Annals and Magazine of Natural History, Series 7, 15, 379–387.
    https://doi.org/10.1080/03745480509443064

    Distant, W.L. (1905c) Rhynchotal notes XXXII. Annals and Magazine of Natural History, Series 7, 15, 478–486.
    https://doi.org/10.1080/03745480509442837

    Distant, W.L. (1905d) Rhynchotal notes XXX. Annals and Magazine of Natural History, Series 7, 15, 304–319.
    https://doi.org/10.1080/03745480509443047

    Distant, W.L. (1905e) Rhynchotal notes XXXIV. Annals and Magazine of Natural History, Series 7, 16, 203–216.
    https://doi.org/10.1080/03745480509443671

    Distant, W.L. (1905f) Rhynchotal notes XXXV. Annals and Magazine of Natural History, Series 7, 16, 265–281.
    https://doi.org/10.1080/03745480509442862

    Distant, W.L. (1905g) Rhynchotal notes XXIX. Annals and Magazine of Natural History, Series 7, 15, 58–70.
    https://doi.org/10.1080/03745480509443637

    Distant, W.L. (1906) A synonymic catalogue of Homoptera. Part I. Cicadidae. Trustees of the British Museum, London, 207 pp.

    Dorval, A., Filho, O.P., Costa, R.B. da & de Souza, M.D. (2011) Diversidade de cigarras (Hemiptera-Homoptera) em ambientes florestais no município de Cotriguaçu, estado de Mato Grosso. Diversity of cicadas (Hemiptera-Homoptera) in forest environments in the municipality of Cotriguaçu, state of Mato Grosso. Multitemas, Campo Grande, MS, 40, 7–19.

    Duffels, J.P. & van der Laan, P.A. (1985) Catalogue of the Cicadoidea (Homoptera, Auchenorhyncha) 1956–1980. In: Series Entomologica 34. Dr. W. Junk Publishers, Dordrect, pp. 1–414.

    Fabricius, J.C. (1775) Systema entomologiae, sistens insectorum classes, ordines, genera, species, adiectis, synonymis, locis, descriptionibus, observationibus. In Officina Libraria Kortii, Flensbvrgi et Lipsiae, 832 pp.

    Fabricius, J.C. (1803) Systema Rhyngotorum secundum ordines, genera, species, adiectis synonymis, locis, observationibus, descriptionibus. Reichard, Brunsvigae, 314 pp.

    Germar, E.F. (1821) Bemerkungen über einige Gattungen der Cicadarien. Magazin der Entomologie, 4, 1–106.

    Germar, E.F. (1830) Species Cicadarium enumeratae et sub genera distributae. Thon's Entomologisches Archiv, 2, 37–57.

    Germar, E.F. (1834) Observations sur plusieurs especes du genre Cicada, Latr. Revue Entomologique Publiée par Gustav Silbermann, 2, 49–82.

    Goding, F.W. (1925) Synopsis of the Cicadidae of Ecuador. Revista del Colegio Nacional Rocafuerte, 7, 1–34.

    Goemans, G. (2010) A historical overview of the classification of the Neotropical tribe Zammarini (Hemiptera, Cicadidae) with a key to the genera. ZooKeys, 43, 1–13.
    https://doi.org/10.3897/zookeys.43.386

    Goemans, G. (2016) The classification and phylogeny of the Neotropical cicada tribe Zammarini (Hemiptera, Cicadidae) and a revision of its type genus Zammara Amyot & Audinet-Serville, 1843 and its sister genus Zammaralna Boulard & Sueur, 1996. Ph.D. dissertation, University of Connecticut, Storrs, 206 pp.

    Gogala, M., Šporar, K., Sanborn, A.F. & Maccagnan, D.H.B. (2015) New cicada species of the genus Guyalna (Hemiptera: Cicadidae) from Brazil. Acta Entomologica Slovenica, 23, 105–116.

    Handlirsch, A. (1925) Ordung: Homoptera (Latr.) Westw. (Homopteren). In: Systematische Übersicht (Schluss.). Schröder’s Handbuch der Entomologie, 3, pp. 1102–1126.

    Hill, K.B.R., Marshall, D.C., Moulds, M.S. & Simon, C. (2015) Molecular phylogenetics, diversification, and systematics of Tibicen Latreille 1825 and allied cicadas of the tribe Cryptotympanini, with three new genera and emphasis on species from the USA and Canada (Hemiptera: Auchenorrhyncha: Cicadidae). Zootaxa, 3985 (2), 219–251.
    https://doi.org/10.11646/zootaxa.3985.2.3

    International Commission on Zoological Nomenclature. (1999) International Code of Zoological Nomenclature. 4th Edition. International Trust for Zoological Nomenclature, London, 306 pp.

    Jacobi, A. (1904) Neue Cicadiden und Fulgoriden Brasiliens. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin, 1904, 155–164.
    https://doi.org/10.5962/bhl.part.8959

    Jacobi, A. (1907) Homoptera Andina. Die Zikaden des kordillerengebietes von Südamerika nach systematik und Verbreitung. I. Cicadidae. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch-Enthnographischen Museums zu Dresden, 11, 1–28.

    Johnson, L.K. & Foster, R.B. (1986) Association of large Homoptera (Fulgoridae and Cicadidae) and the trees in a tropical forest. Journal of the Kansas Entomological Society, 59, 415–422.

    Kato, M. (1932) Monograph of Cicadidae. Sanseido, Tokyo, 450 pp.

    Kirkaldy, G.W. (1909) Hemiptera, old and new. No. 2. Canadian Entomologist, 41, 388–392.
    https://doi.org/10.4039/Ent41388-11

    de Laporte, F.L. (1832) Mémoire sur cinquante espèces nouvelles ou peu connues d'insectes. Annales de la Société Entomologique de France, 1, 386–415.

    Latreille, P.A. (1802) Histoire naturelle, générale et particulière des crustacés et des insectes. Ouvrage faisant suite à l’histoire naturelle générale et particulière, composée par Leclerc de Buffon, et rédigée par C.S. Sonnini, membre de plusieurs sociétés savantes. Familles naturelles des genres. Tome Troisième. F. Dufast, Paris, 467 pp. [pp. i–xii + 13–467]

    Leeuwen, W.J.D. van, Hartfield, K., Miranda, M. & Meza, F.J. (2013) Trends and ENSO/AAO Driven variability in NDVI derived productivity and phenology alongside the Andes Mountains. Remote Sensing, 5, 1177–1203.
    https://doi.org/10.3390/rs5031177

    Linnaeus, C. (1758) Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tomus I. Edito duodecima, reformata. Laurentius Salvius, Holmiae, 824 pp.

    Linné, C. (1767) Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tomus I. Pars II. Editio duodecima reformata. Laurentius Salvius, Holmiae, 795 pp. [pp. 533–1327]
    https://doi.org/10.5962/bhl.title.37256

    MacArthur, R.H. & Wilson, E.O. (1967) The theory of island biogeography. Princeton University Press, Princeton, 203 pp.

    Maccagnan, D.H.B. & Martinelli, N.M. (2011) Description and key to the fifth-instars of some cicadas (Hemiptera: Cicadidae) associated with coffee plants in Brazil. Neotropical Entomology, 40, 445–451.
    https://doi.org/10.1590/S1519-566X2011000400006

    Maccagnan, D.H.B., Martinelli, N.M., Goemans, G. & Guilherme, E. (2011) Zammara smaragdula Walker, 1858 (Hemiptera: Cicadidae): first record for Brazil. Check List, 7, 563–564.
    https://doi.org/10.15560/7.4.563

    Maccagnan, D.H.B., Matuo, T.K. & Martinelli, N.M. (2017) Oviposition of Quesada gigas in weed no hostess: implication in pest management. Ciências Rural, 47 (09), e20101752, 1–4.
    https://doi.org/10.1590/0103-8478cr20170152

    Maccagnan, D.H.B., Pitta, R.M., Lunz, A.M., Behling, M. & Martinelli, N.M. (2014) Primeiro registro de cigarra em reflorestamentos com paricá no Estado de Mato Grosso Brasil. Revista de Ciências Agrárias, 57, 451–454.
    https://doi.org/10.4322/rca.1387

    Maes, J.-M., Moulds, M. & Sanborn, A.F. (2012) Cicadidae (Homoptera) de Nicaragua: catálogo ilustrado, incluyendo especies exóticas del Museo Entomólogico de Léon. Revista Nicaragüense Entomología, 72 (Suplemento 2), 1–138.

    Marshall, D.C., Moulds, M.S., Hill, K.B.R., Price, B.W., Wade, E.J., Owen, C.J., Goemans, G., Marathe, K., Sarkar, V., Cooley, J.R., Sanborn, A.F., Kunte, K., Villet, M.H. & Simon, C. (2018) A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily level classification. Zootaxa, 4424 (1), 1–64.
    https://doi.org/10.11646/zootaxa.4424.1.1

    Martinelli, N.M. & Zucchi, R.O. (1997) Cigarras (Hemiptera: Cicadidae: Tibicinidae) associadas ao cafeeiro: Distribuição, hospedeiros e chave para as espécies. Anais da Sociedade Entomológica do Brasil, 26, 133–143.
    https://doi.org/10.1590/S0301-80591997000100018

    Metcalf, Z.P. (1952) New names in the Homoptera. Journal of the Washington Academy of Sciences, 42, 226–231.

    Metcalf, Z.P. (1955) New names in the Homoptera. Journal of the Washington Academy of Sciences, 45, 262–267.

    Metcalf, Z.P. (1963a) General catalogue of the Homoptera, Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section I. Tibiceninae. North Carolina State College Contribution, 1502, i–vii + 1–585.

    Metcalf, Z.P. (1963b) General catalogue of the Homoptera. Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section II. Gaeninae and Cicadinae. North Carolina State College Contribution, 1502, 587–919.

    Metcalf, Z.P. (1963c) General catalogue of the Homoptera, Fascicle VIII. Cicadoidea. Part 2. Tibicinidae. North Carolina State College Contribution, 1564, i–vi + 1–492.

    Monteiro, O.M., Lunz, A.M., Azevedo, R. de, Júnior, M.M. & Batista, T.F.V. (2014) Avaliação de inseticida para controle da cigarra Quesada gigas em plantios de paricá. Pesquisa Florestal Brasileira, 34, 169–172.
    https://doi.org/10.4336/2014.pfb.34.78.502

    Möschler, H.B. (1883) Beiträge zur Schmetterlings-Fauna von Surinam. V. (Supplement.). Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, 32, 303–362, pls. XVII–XVIII.

    Moulds, M.S. (2005) An appraisal of the higher classification of cicadas (Hemiptera: Cicadoidea) with special reference to the Australian fauna. Records of the Australian Museum, 57, 375–446.
    https://doi.org/10.3853/j.0067-1975.57.2005.1447

    Moulds, M.S. (2012) A review of the genera of Australian cicada (Hemiptera: Cicadoidea). Zootaxa, 3287, 1–262.

    Oliveira, R.R., Caramori, S.S. & Maccagnan, D.H.B. (2017) Could Fidicina mannifera (Hemiptera: Cicadoidea: Fidicinini) promote a resource pulse in two Brazilian Cerrado vegetation classes? Brazilian Journal of Biology, 77, 673–679.
    https://doi.org/10.1590/1519-6984.13715

    Olivier, G.A. (1790) Cigale, Cicada. Encyclopedie Méthodique Histoire Naturelles Insectes, 5, 735–760.

    Ramos, J.A. & Wolda, H. (1985) Description and distribution of two new species of Selymbria from Panama (Homoptera, Cicadoidea, Tibicinidae). Caribbean Journal of Science, 21, 177–185.

    Reis, P.R., Rebelles, P.P.R., Pereira, M.C., Liska, G.R. & Morais, A.R. de (2015) Eficácia do enxofre aplicado via solo no controle da cigarra Quesada gigas (Olivier) em cafeeiro. Coffee Science, Lavras, 10, 527–536.

    Ruschel, T.P. (2017) Seven new species of the cicada genus Guyalna Boulard & Martinelli, 1996 (Hemiptera: Cicadidae: Fidicinini) with a re-description of the type species. Zootaxa, 4281 (1), 246–279.
    https://doi.org/10.11646/zootaxa.4281.1.25

    Sanborn, A.F. (2001) A first contribution to a knowledge of the cicada fauna of El Salvador (Homoptera: Cicadoidea). Florida Entomologist, 84, 449–450.
    https://doi.org/10.2307/3496510

    Sanborn, A.F. (2004) Two new cicada Zammara species from South America (Hemiptera: Cicadoidea: Cicadidae). Florida Entomologist, 87, 365–371.
    https://doi.org/10.2307/0015-4040(2004)087[0365:TNZSFS]2.0.CO;2

    Sanborn, A.F. (2005) Fidicina variegata, a new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae). Annals of the Entomological Society of America, 98, 187–190.
    https://doi.org/10.1603/0013-8746(2005)098[0187:FVANCS]2.0.CO;2

    Sanborn, A.F. (2006a) New records for the cicada fauna from four Central American countries (Hemiptera: Cicadoidea: Cicadidae). Florida Entomologist, 89, 75–79.
    https://doi.org/10.1653/0015-4040(2006)89[75:NRFTCF]2.0.CO;2

    Sanborn, A.F. (2006b) New records of cicadas from Mexico (Hemiptera: Cicadoidea: Cicadidae). Southwestern Naturalist, 51, 255–257.
    https://doi.org/10.1894/0038-4909(2006)51[255:NROCFM]2.0.CO;2

    Sanborn, A.F. (2007a) New species, new records and checklist of cicadas from Mexico (Hemiptera: Cicadoidea: Cicadidae). Zootaxa, 1651, 1–42.

    Sanborn, A.F. (2007b) Additions to the cicada fauna of Venezuela with the description of a new species and checklist of the Venezuelan cicada fauna (Hemiptera: Cicadoidea: Cicadidae). Zootaxa, 1503, 21–32.

    Sanborn, A.F. (2009) Checklist, new species and key to the cicadas of Cuba (Hemiptera: Cicadoidea: Cicadidae). Mitteilungen aus dem Museum für Naturkunde in Berlin—Deutsche Entomologische Zeitschrift, 59, 85–92.
    https://doi.org/10.1002/mmnd.200900007

    Sanborn, A.F. (2010a) Two new species and new records of cicadas from Central America (Hemiptera: Cicadoidea: Cicadidae). Studies on Neotropical Fauna and Environment, 45, 67–76.
    https://doi.org/10.1080/01650521003778814

    Sanborn, A.F. (2010b) The cicadas of Colombia including new records and the description of a new species (Hemiptera: Cicadidae). Journal of Natural History, 44, 1577–1607.
    https://doi.org/10.1080/00222931003760046

    Sanborn, A.F. (2011a) Checklist of the cicadas of French Guiana including new records and the description of nine new species (Insecta, Hemiptera, Cicadoidea, Cicadidae). Zoosystema, 33, 377–418.
    https://doi.org/10.5252/z2011n3a6

    Sanborn, A.F. (2011b) Checklist of the cicadas (Insecta: Hemiptera: Cicadidae) of Paraguay including new records for six species. Check List, 7, 465–467.
    https://doi.org/10.15560/7.4.465

    Sanborn, A.F. (2013) Catalogue of the Cicadoidea (Hemiptera: Cicadoidea). With contributions to the bibliography by Martin H. Villet. Elsevier/Academic Press, San Diego, 1001 pp.

    Sanborn, A.F. (2014) Checklist of the cicadas (Insecta: Hemiptera: Cicadidae) of Costa Rica including new records for fourteen species. Check List, 10, 246–252.
    https://doi.org/10.15560/10.2.246

    Sanborn, A.F. (2016a) The cicada genus Guyalna Boulard & Martinelli, 1996 (Hemiptera: Cicadidae: Cicadinae: Fidicinini): generic description, twelve new combinations, and a key to species. Zootaxa, 4105 (5), 430–454.
    https://doi.org/10.11646/zootaxa.4105.5.2

    Sanborn, A.F. (2016b) Five new species, a new genus and a new record of cicadas from French Guiana with four new combinations and three new synonymies (Insecta, Hemiptera, Cicadoidea, Cicadidae). Zoosystema, 38, 177–199.
    https://doi.org/10.5252/z2016n2a2

    Sanborn, A.F. (2017a) Generic redescription, seven new species and a key to the Taphura Stål, 1862 (Hemiptera: Cicadidae: Cicadettinae: Taphurini). Zootaxa, 4324 (3), 451–481.
    https://doi.org/10.11646/zootaxa.4324.3.3

    Sanborn, A.F. (2017b) A new species of Carineta Amyot & Audinet-Serville, 1843 (Hemiptera: Cicadoidea: Cicadidae) from Martinique. Proceedings of the Entomological Society of Washington, 119, 629–636.
    https://doi.org/10.4289/0013-8797.119.4.629

    Sanborn, A.F. (2018a) The cicada genus Procollina Metcalf, 1952 (Hemiptera: Cicadidae): redescription including fourteen new species, with a key to the species of the subtribe Dazina Kato, 1932 rev. stat., the description of the Aragualnini n. tribe, and one new combination. Zootaxa, 4389 (1), 1–65.
    https://doi.org/10.11646/zootaxa.4389.1.1

    Sanborn, A.F. (2018b) Two new genera and two new species of cicadas from Central America (Hemiptera: Cicadidae: Cicadinae). Zootaxa, 4418 (2), 148–160.
    https://doi.org/10.11646/zootaxa.4418.2.4

    Sanborn, A.F. & Heath, M.S. (2012) Catalogue of the cicadas (Hemiptera: Cicadoidea: Cicadidae) of continental North America North of Mexico. Thomas Say Publications in Entomology: Monographs. Entomological Society of America, Lanham, MD, 227 pp.

    Sanborn, A.F. & Heath, M.S. (2014) The cicadas of Argentina with new records, a new genus and fifteen new species (Hemiptera: Cicadoidea: Cicadidae). Zootaxa, 3883 (1), 1–94.
    https://doi.org/10.11646/zootaxa.3883.1.1

    Sanborn, A.F. & Heath, M.S. (2017) Catalogue of the cicadas (Hemiptera: Cicadoidea: Cicadidae) of continental North America North of Mexico, Second Edition. Thomas Say Publications in Entomology: Monographs. Entomological Society of America, Lanham, MD. 302 pp.

    Sanborn, A.F., Heath, M.S., Heath, J.E. & Noriega, F.G. (1995) Diurnal activity, temperature responses and endothermy in three South American cicadas (Homoptera: Cicadidae: Dorisiana bonaerensis, Quesada gigas, and Fidicina mannifera). Journal of Thermal Biology, 20, 451–460.
    https://doi.org/10.1016/0306-4565(95)00004-G

    Sanborn, A.F., Heath, M.S., Phillips, P.K. & Heath, J.E. (2011) The genus Cacama Distant, 1904 (Hemiptera: Cicadidae) with the description of three new species. Zootaxa, 2897, 35–50.

    Sanborn, A.F., Heath, M.S., Phillips, P.K. & Heath, J.E. (2012) The genus Pacarina Distant, 1905 (Hemiptera: Cicadidae) with the description of a new species. Journal of Natural History, 46, 923–941.
    https://doi.org/10.1080/00222933.2011.651647

    Sanborn, A.F., Heath, M.S., Sueur, J. & Phillips, P.K. (2005) The genus Neocicada Kato, 1932 (Hemiptera: Cicadomorpha: Cicadidae), with descriptions of three new species. Systematic Entomology, 30, 191–207.
    https://doi.org/10.1111/j.1365-3113.2005.00266.x

    Sanborn, A.F. & Maes, J.-M. (2012) Checklist of the cicadas (Insecta: Hemiptera: Cicadidae) of Nicaragua including new records for seventeen species. Check List, 8, 437–442.
    https://doi.org/10.15560/8.3.437

    Sanborn, A.F., Moore, T.E. & Young, A.M. (2008) Two new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae) with a key to Fidicinoides in Costa Rica. Zootaxa, 1846, 1–20.

    Santos, R.S., Martinelli, N.M., Maccagnan, D.H.B., Sanborn, A.F. & Ribeiro, R. (2010) Description of a new cicada species associated with the coffee plant and an identification key to the species of Fidicinoides (Hemiptera: Cicadidae) from Brazil. Zootaxa, 2602, 48–56.

    Santos-Cividanes, T.M., Seguino, E., Cividanes, F.J., Martinelli, N.M., Martins, A.M. & Perdoná, M.J. (2013) First record of Dorisiana viridis (Hemiptera: Cicadidae) on Macadamia Nut (Proteales: Proteacea) in Brazil. Florida Entomologist, 96, 1221–1223.
    https://doi.org/10.1653/024.096.0373

    Scopoli, J.A. (1763) Entomologia Carniolica exhibens Insecta Carnioliae indigena et distribute in ordines, genera, species, varietates. Methodo Linnaeana. Typis Ioannis Thomae Trattner, Caes. Reg. Avlae Typogr. Et Bibliop, Vindobonae, 420 pp.
    https://doi.org/10.5962/bhl.title.119976

    Stål, C. (1854) Nya Hemiptera. Öfversigt af Svenska Vetenskaps Akademien Förhandlingar, 11, 231–255.

    Stål, C. (1861) Genera nonnulla nova Cicadinorum. Annales de la Société Entomologique de France, Series 4, 1, 613–622.

    Stål, C. (1862) Bidrag till Rio Janeiro - traktens Hemipter-fauna. II. Handlingar. Kongliga Svenska Vetenskaps Akademien, Stockholm, 3, 1–75.

    Stål, C. (1864) Hemiptera mexicana enumeravit speciesque novas descripsit (continuatio). Stettiner Entomologische Zeitung. Herausgegeben von dem entomologischen Vereine zu Stettin, 25, 49–86.

    Stål, C. (1866) Analecta Hemipterologica. Berliner Entomologischer Zeitschrift, 10, 151–172.
    https://doi.org/10.1002/mmnd.18660100112

    Stål, C. (1870) Hemiptera insularum Phillippinarum. Bidrag till Philippinska öarnes Hemipter-fauna. Öfversigt af Svenska Vetenskaps Akademien Förhandlingar, 27, 607–776.

    Sueur, J. (2000) Une nouvelle espèce de cigale du Mexique (Los Tuxtlas, Veracruz), et étude de son émission sonore (Homoptera, Auchenorhyncha, Cicadoidea). Bulletin de la Société entomologique de France, 105, 217–222.

    Torres, B.A. (1948) Sobre seis nuevas especies del genero Carineta Amy. et Serv. (Homoptera-Cicadidae). Notas del Museo de La Plata, 13, 113–127.

    Torres, B.A. (1963) Desmembracion en los generos Proarna Stål y Tympanoterpes Stål. Creacion del nuevo genero: Prasinosoma (Homoptera, Cicadidae). Revista Sociedad Uruguaya Entomologica, 5, 13–23.

    Uhler, P.R. (1903) Enumeration of the Cicadidae of Brazil in the collection of Mr. Herbert H. Smith. Transactions of the Maryland Academy of Sciences, 2, 1–17.

    Walker, F. (1850) List of the Specimens of Homopterous Insects in the Collection of the British Museum. Part I. British Museum Trustees, London, 260 pp.

    Walker, F. (1858a) List of the specimens of Homopterous insects in the collection of the British Museum. Supplement. British Museum Trustees, London, 307 pp.

    Walker, F. (1858b) Insecta Saundersiana: or characters of undescribed insects in the collection of William Wilson Saunders, Esq. Homoptera. John van Voorts, London, 117 pp.
    https://doi.org/10.5962/bhl.title.5112

    Westwood, J.O. (1851) Descriptions of some new species of exotic homopterous insects. Annals and Magazine of Natural History, Series 2, 7, 207–210.

    White, A. (1845) Appendix (D.) Description and figures of four new species of Australian insects. In: Eyre, E.J. (Ed.), Journals of expeditions of discovery into Central Australia and overland from Adelaide to King George Sound, in the years 1840–1; sent by the colonists of South Australia, with the sanction and support of the government: including an account of the manners and customs of the Aborigines and the state of their relations with Europeans. T. and W. Boone, London, pp. 432–434, pl. 4.

    Wolda, H. (1977) Fluctuations in abundance of some Homoptera in a neotropical forest. Geo-Eco-Trop, 3, 229–257.

    Wolda, H. (1978) Seasonal fluctuations in rainfall, food, and abundance of tropical insects. Journal of Animal Ecology, 47, 369–381.
    https://doi.org/10.2307/3789

    Wolda, H. (1983a) "Longterm" stability of tropical insect populations. Researches on Population Ecology, Supplement, 3, 112–126.
    https://doi.org/10.1007/BF02539633

    Wolda, H. (1983b) Spatial and temporal variation in abundance in tropical animals. In: Sutton, S.L., Whitmore, T.C. & Chadwick, H.C. (Eds.), Tropical rain forest: Ecology and management. Blackwell Scientific Publishers, Oxford, pp. 93–105.

    Wolda, H. (1984) Diversity and seasonality of Panamanian cicadas. Mitterlungen du Schweizerischen Entomologischen Gesellschaft, 57, 451.

    Wolda, H. (1988) Insect seasonality: Why? Annual Review of Ecology and Systematics, 19, 1–18.
    https://doi.org/10.1146/annurev.es.19.110188.000245

    Wolda, H. (1989) Seasonal cues in tropical organisms: rainfall? Not necessarily. Oecologia, Berlin, 80, 437–442.
    https://doi.org/10.1007/BF00380064

    Wolda, H. (1993) Diel and seasonal patterns of mating calls in some neotropical cicadas. Acoustic interference? Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen. Biological, Chemical, Geological, Physical and Medical Sciences, 96, 369–381.

    Wolda, H. & Ramos, J.A. (1992) Cicadas in Panama, their distribution, seasonality and diversity. In: Quintero, D. & Aiello, A. (Eds.), The insects of Panama and Mesoamerica. Selected studies. Oxford University Press, New York, pp. 271–279.

    Young, A.M. & Sanborn, A.F. (2015) The seasonal and nocturnal attraction of Proarna and Pacarina cicadas to lights in the Caribbean floodplain of Costa Rica and in French Guiana (Hemiptera: Cicadidae). Entomologica Americana, 121, 13–17.
    https://doi.org/10.1664/15-RA-027.1