Abstract
Phacosoma basilewskyi Balthasar, 1960 was described for a peculiar East African deltochiline dung beetle species. Years later, Paulian (1975) erected the deltochiline genus Madaphacosoma Paulian, 1975 for two new Malagasy species, namely Madaphacosoma humberti Paulian, 1975, as the type species, and M. betschi Paulian, 1975. Phacosoma basilewskyi was transferred to Madaphacosoma in that same publication. Later, Paulian (1991) added Madaphacosoma major Paulian, 1991 to the genus for a total of three Madagascan and one mainland African species. Recently, Madaphacosoma together with three other genera occurring in Madagascar, namely Aleiantus Olsoufieff, 1947, Phacosomoides Martinez & Pereira, 1959 and Sikorantus Paulian, 1976, were synonymized with Epactoides Olsoufieff, 1947 by Wirta & Montreuil (2008). Although Madaphacosoma basilewskyi (Balthasar, 1960) is not specifically mentioned as a new combination with three other species, M. major, M. humberti and M. betschi, catalogued on pages 661–662 in Wirta & Montreuil (2008), and the genus is treated as valid by Tarasov & Dimitrov (2016) and Tarasov (2017), it is currently lumped with the other Malagasy species in the genus Epactoides. The current accepted name for the species is thus Epactoides basilewskyi (Balthasar, 1960).
References
Balthasar, V. (1960) Mission zoologique de l’I.R.S.A.C. en Afrique orientale (P. Basilewsky et N. Leleup, 1957). XLI. Coleoptera Scarabaeidae: Scarabaeini, Coprini, Onitini et Oniticellini. Annales du Musée Royal de l’Afrique Centrale, Série 8°, Sciences Zoologiques, 88, 55–66.
Mlambo, S., Sole, C.L. & Scholtz, C.H. (2014) Affinities of the Canthonini dung beetles of the Eastern Arc Mountains. Organisms, Diversity and Evolution, 14, 115–120.
https://doi.org/10.1007/s13127-013-0158-y
Monaghan, M.T., Inward, D.J.G., Hunt, T. & Vogler, A.P. (2007) A molecular phylogenetic analysis of the Scarabaeinae (dung beetles). Molecular Phylogenetics and Evolution, 45, 674–692.
https://doi.org/10.1016/j.ympev.2007.06.009
Paulian, R. (1975) Sur quelques Canthonina (Coléoptères Scarabaéides) montagnards de Madagascar. Annales de la Société Entomologique de France, 11, 221–252.
Paulian, R. (1991) Les Scarabaeoidea obtenus d’excréments de Propithecus diadema à Madagascar. Bulletin de la Société Entomologique de France, 96 (4), 355–359.
Scholtz, C.H. & Howden, H.F. (1987) A revision of the African Cathonina [sic] (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of the Entomological Society of Southern Africa, 50, 75–119.
Tarasov, S. & Dimitrov, D. (2016) Multigene phylogenetic analysis redefines dung beetles relationships and classification. BMC Evolutionary Biology, 16, 1–19.
https://doi.org/10.1186/s12862-016-0822-x
Tarasov, S. (2017) A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data. Zootaxa, 4329 (2), 101–149.
https://doi.org/10.11646/zootaxa.4329.2.1
Wirta, H. & Montreuil, O. (2008) Evolution of the Canthonini Longitarsi (Scarabaeidae) in Madagascar. Zoologica Scripta, 37, 651–663.
https://doi.org/10.1111/j.1463-6409.2008.00352.x