Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2019-02-28
Page range: 41–66
Abstract views: 117
PDF downloaded: 3

Provisional checklist of the Tettigonioidea (Insecta: Orthoptera) from São Tomé & Príncipe with taxonomic remarks, bioacoustical data and the description of new taxa

Grillenstieg 18, 39120 Magdeburg, Germany.
Orthoptera Corycoides Catopropteryx Adapantus Conocephalus Ruspolia Brachyamytta calling song

Abstract

The state São Tomé and Príncipe consists of two separated oceanic archipelagos situated in the Gulf of Guinea (West Central Africa). Except of a first survey on São Tomé in the 19th century, its Orthoptera fauna has never been studied systematically. Here a list of all tettigonioid species which have been recorded from the islands is presented together with information about the bioacoustics and biology of some species. Each archipelago harbours nine species, but only two species are found in both parts of the country. Two species and one subgenus are described as new: Adapantus (Adapantus) excavatus sp. nov., Brachyamytta bombomensis sp. nov., Neoadapantus subg. nov. The following nomenclatural changes had to be done: Corycoides greeffi (Krauss, 1890) becomes new synonym of Corycoides paradoxus (Bolivar, 1890), Xiphidion guineensis Redtenbacher, 1891, Xiphidion africanum Redtenbacher, 1891 and Conocephalus inequalis Uvarov, 1928 become new synonyms of Conocephalus iris Serville, 1838. The species list of Conocephalus, subgenus Conocephalus, is revised. Additionally the known tettigonioids of Annobón (Equatorial Guinea) are listed (4 species).

 

References

  1. Ando, Y. & Hartley, J.C. (1982) Occurrence and biology of a long-winged form of Conocephalus discolor. Entomologia experimentalis et applicata, 32, 238–241.

    https://doi.org/10.1111/j.1570-7458.1982.tb03212.x

    Bailey, W.J. (1975) A review of the African species of the genus Ruspolia Schulthess (Orthoptera Tettigonioidea). Bulletin de l' Institut Fondamental d' Afrique Noire, Série A 37 A, 171–226.

    Bailey, W.J. (1976) Species isolation and song types of the genus Ruspolia Schulthess in Uganda. Journal of Natural History, 10, 511–528.

    https://doi.org/10.1080/00222937600770421

    Bazelet, C. & Naskrecki, P. (2014) Conocephalus iris. The IUCN Red List of Threatened Species 2014, e.T20633843A56178684.

    https://doi.org/10.2305/IUCN.UK.2014-3.RLTS.T20633843A56178684.en.

    Beier, M. (1954) Revision der Pseudophyllinen 1. Hälfte. Trabajos del Instituto Espanol de Entomologia, Madrid. 479 pp.

    Beier, M. (1962) Orthoptera Tettigoniidae (Pseudophyllinae I). Das Tierreich 73, Walter de Gruyter & Co., Berlin, 468 pp.

    Bland, R.G., Gangwere, S.K. & Morales M.M. (1996) An annotated list of the Orthoptera (sens. lat.) of the Canary Islands. Journal of Orthoptera Research, 5, 159–173.

    https://doi.org/10.2307/3503589

    Bolívar, I. (1890) Ortópteros de Africa del Museo de Lisboa. Jornal de Sciencias da Academia de Lisboa, 1 (2), 211–232.

    Bolívar, I. (1905a) Nueva especie de Xiphidium de las Azores. Boletin de la Real Sociedad Española de Historia Natural, 5, 180–181.

    Bolívar, I. (1905b) Conocéphalides de la Nouvelle-Guinée appartenant au Musée de Budapest. Annales Historico-Naturales Musei Nationalis Hungarici, Pars Zoologica, 3, 388–395.

    Bolívar, I. (1906) Fasgonuridos de la Guinea española. Memorias de la Real Sociedad Española de Historia Natura, 1, 327– 377.

    Bruner, L. (1891) Ten new species of Orthoptera from Nebraska—Notes on habits, wing variation, etc. Canadian Entomologist, 23, 36–40, 56–59, 70–73.

    Bruner, L. (1915) Notes on tropical American Tettigonoidea (Locustodea). Annals of the Carnegie Museum, 9, 284–404.

    Bruner, L. (1920) Orthoptera from Africa, being a report upon some Saltatoria mainly from Cameroon contained in the Carnegie Museum. Annals of the Carnegie Museum 13 (1919), 92–142.

    Brunner von Wattenwyl, C. (1878) Monographie der Phaneropteriden. Brockhaus, Wien, 401 pp.

    Burr, M. (1927) Sao Thome and Principe. Entomologist's Record and Journal of Variation, 39, 117–120.

    Brühl, C. (1993) Überblick über Geographie, Geologie, Klima und Vegetationsformationen der Insel São Tomé (Golf von Guinea). Faunistische Abhandlungen, Staatliches Museum für Tierkunde Dresden, 19 (2), 5–16.

    Buxton, P.A. & Uvarov, B.P. (1923) A contribution to our knowledge of Orthoptera of Palestine. Bulletin de la Societé entomolgique d' Egypt, 1923, 167–214.

    Charpentier, T. de (1825) De Orthopteris Europaeis. Horae entomologicae, 4, 61–181.

    Chopard, L. (1945) Orthoptéroïdes recueillis dans les montagnes du Cameroun par la mission Lepesme, Paulian, Villiers. Revue Française d' Entomologie, 11, 156–178.

    Chopard L. (1954) La réserve naturelle intégrale du Mont Nimba, fascicule 2. III. Orthoptéroides Ensifères. Mémoires de l'Institut Français d'Afrique Noire, 40, 25–98.

    Chopard, L. (1955) Orthoptera Ensifera. In: Hanstrom, B., Brinck, P. & Rudebeck, G. (Eds.), South African Animal Life (Results of the Lund Univ. Exp., 1950–51), 2. Almqvist & Wiksell, Stockholm, pp. 266–300.

    Chopard, L. (1958) Mission du Muséum dans les îles du Golfe de Guinée. Entom. 6. Orthoptéroides. Bulletin de la Société Entomologique de France, 63, 73–85.

    Cigliano, M.M., Braun, H., Eades, D.C. & Otte, D. (2018) Orthoptera Species File. Version 5.0/5.0. Available from: http://Orthoptera.SpeciesFile.org/ (accessed 12 June 2018).

    Copete, J.L. (2016) A scops-owl (Otus) discovered in the forests of Príncipe Island (Gulf of Guinea). Available from: https://www.hbw.com/news/scops-owl-otus-discovered-forests-principe-island-gulf-guinea/ (accessed 3 October 2018)

    Ebner, R. (1927) Gryllacriden und Tettigoniiden von Nordost-Sumatra (Vorlaeufige Mitteilung). Miscellanea Zoologica Sumatrana, 20, 1–3.

    Fox, H. (1912) Two apparently hitherto undescribed species of Xiphidium from the Salt Marshes of the Atlantic Coast of the United States (Orthop.). Entomological News, 23, 111–119.

    Gerstaecker, A. (1869) Beitrag zur Insekten–Fauna von Zanzibar. No. II. Orthoptera et Neuroptera. Archiv für Naturgeschichte (Zeitschrift für systematische Zoologie), 35 (1), 201–223.

    https://doi.org/10.5962/bhl.part.12294

    Giglio-Tos, E. (1907) Ortotteri Africani. Parte 2. Blattodea, Mantodea, Phasmodea, Locustodea, Gryllodea. Bollettino del Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino, 22 (563), 1–26.

    Greef, R. (1882) Die Insel Rolas. Globus. Illustrierte Zeitschrift für Länder und Völkerkunde mit besonderer Berücksichtigung der Anthropologie und Ethnologie, 41, I, 110-112, II, 122-125, III, 135–138.

    Greef, R. (1884) Ueber die Fauna der Guinea-Inseln S. Thome und Rolas. Sitzungsberichte der Gesellschaft zur Beförderung der gesammten Naturwissenschaften zu Marburg, 2, 41–79.

    Griffini, A. (1906) Ortotteri raccolti da Leonardo Fea nell'Africa occidentale. 1. Hetrodidi, Conocephalidi, Meconemidi, Pseudophyllidi, Mecopodidi e Fanerotteridi. Annali del Museo Civico di Storia Naturale 'Giacomo Doria‘, Genova, (3) 2, 358–397.

    Griffini, A. (1908) Phasgonuridae africane del R. Museo di Storia Naturale di Bruxelles, 4. Meconemidae et Conocephalidae. Mémoires de la Societé Entomologique de Belgique, 15, 65–73. Available from: https://www.biodiversitylibrary.org/item/41740#page/215/mode/1up (Accessed 27 Feb. 2019)

    Griffini, A. (1909). Note sopra alcune Phasgonuridae del Congo. Annales de la Société entomologique de Belgique, 53 [1908], 9–28. Available from: https://www.biodiversitylibrary.org/item/46155#page/17/mode/1up (Accessed 27 Feb. 2019)

    Haan, W. de (1843) Bijdragen tot de kennis der Orthoptera. In: Temminck (Ed.), Verhandelingen over de Natuurlijke Geschiedenis der Nederlansche Overzeesche Bezittingen, de Leden der Natuurkundige Commissie in Indië en andere Schrijvers. 19/20, 165–228.

    Harz, K. (1966) Orthopterlogische Beitraege VI. Nachrichtenblatt der Bayerischen Entomologen, 15, 24–29.

    Harz, K. (1967) Eine neue europäische Conocephalus-Art (Orthoptera, Ensifera). Nachrichtenblatt der Bayerischen Entomologen, 16, 98–100.

    Harz, K. (1969a) Vorläufige Revision der Gattung Conocephalus im Bereich der Paläarktis. Berichte Naturwissenschaftliche Gesellschaft Bayreuth, 13, 97–118.

    Harz, K. (1969b) Die Orthopteren Europas 1. Dr. W. Junk N.V., The Hague, 749 pp.

    https://doi.org/10.1007/978-94-017-2511-8

    Harz, K. (1970) New and little known Tettigoniidae (Orthoptera) from Israel and Africa. Israel Journal of Zoology, 19, 191–201.

    Hebard, M. (1933) The Dermaptera and Orthoptera of the Marquesas Islands. Bulletin of the Bernice P. Bishop Museum, Honolulu, 114, 105–140.

    Heller, K.-G. (1988) Bioakustik der europäischen Laubheuschrecken. Ökologie in Forschung und Anwendung, Bd. 1. Verlag J. Margraf, Weikersheim, 358 pp.

    Heller, K.-G., Hemp,, C., Liu,C.-X. & Volleth, M. (2014) Taxonomic, bioacoustic and faunistic data on a collection of Tettigonioidea from Eastern Congo (Insecta: Orthoptera). Zootaxa, 3785 (3), 343–376.
    http://dx.doi.org/10.11646/zootaxa.3785.3.2

    Heller, K.-G., Volleth, M., Vedenina, V., Maryańska-Nadachowska, A. & Warchałowska-Śliwa, E. (2019) A perfect duet? The acoustic behaviour of Anaulacomera almadaenis n. sp. (Orthoptera, Tettigonioidea, Phaneropterinae), a species with an unusual chromosome complement, discovered in the footsteps of the explorers Spix and Martius in Brazil. Spixiana. [in press)]

    Hemp, C. & Heller, K.-G. (2017) Revision of the genus Amytta (Orthoptera: Tettigoniidae, Meconematinae) and new species from East Africa. Zootaxa, 4263 (2), 295–317.
    http://dx.doi.org/10.11646/zootaxa.4263.2.5

    Hemp, C., Heller, K.-G., Warchalowska-Sliwa, E., Grzywacz,B. & Hemp, A. (2017) Review of the East African species of the phaneropterine genus Parapyrrhicia Brunner von Wattenwyl, 1891 (Insecta: Orthoptera): secret communication of a forest-bound taxon. Organisms Diversity & Evolution, 17, 231–250.

    https://doi.org/10.1007/s13127-016-0303-5

    Higaki, M. & Ando, Y. (2003) Effects of crowding and photoperiod on wing morph and egg production in Eobiana engelhardti subtropica (Orthoptera: Tettigoniidae). Applied Entomology and Zoology, 38 (3), 321–325.

    https://doi.org/10.1303/aez.2003.321

    Holstein, J. & Ingrisch, S. (2004) The Orthoptera type material (Insecta) in the collections of the Staatliches Museum für Naturkunde Stuttgart - Part 1: Ensifera. Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie), 665,1–33.

    Hugel, S. (2009) Gryllacrididae and Tettigoniidae (Insecta, Orthoptera, Ensifera) from Espiritu Santo, Vanuatu. Zoosystema, 31 (3), 525–576.

    https://doi.org/10.5252/z2009n3a10

    Hugel, S. (2010) New and little known predatory katydids from Mascarene islands (Ensifera: Meconematinae and Hexacentrinae). Zootaxa, 2543, 1–30.

    https://doi.org/10.11646/zootaxa.2543.1.1

    Hugel, S. (2012) New and little known Phisidini from Madagascar, Comoros and Seychelles (Orthoptera, Ensifera, Meconematinae). Zoosystema 34 (3), 525–552.

    https://doi.org/10.5252/z2012n3a3

    Huxley, J. (1970) A revision of the genus Catoptropteryx Karsch (Orthoptera: Tettigoniidae). Bulletin of the British Museum (Natural History), Entomology Series, 24, 129–170.

    https://doi.org/10.5962/bhl.part.1520

    Ingrisch, S. (1996) Evidence of an embryonic diapause in a tropical Phaneropterinae (Insecta Ensifera Tettigonioidea). Tropical Zoology, 9, 431–439.

    https://doi.org/10.1080/03946975.1996.10539321

    Ingrisch, S. (1990) Zur Laubheuschrecken-Fauna von Thailand (Insecta: Saltatoria: Tettigoniidae). Senckenbergiana Biologica, 70 (1989), 89–138.

    Ingrisch, S. & Köhler, G. (1998). Die Heuschrecken Mitteleuropas. Die Neue Brehm-Bücherei 629, Westarp Wissenschaften, Magdeburg, 460 pp.

    Juste, B.J. & Ibañez, C. (1994) Bats of the Gulf of Guinea islands: faunal composition and origins. Biodiversity and Conservation, 3, 837-850.

    https://doi.org/10.1007/BF00129662

    Karny, H.H. (1907) Revisio Conocephalidarum. Abhandlungen der zoologisch-botanischen Gesellschaft Wien, 4, 1–114.

    Karny, H.H. (1912) Orthoptera Fam. Locustidae Subfam. Conocephalinae. Genera Insectorum, 135, 1–17.

    Karny, H.H. (1920) Dodecas Conocephalidarum novarum. Verhandlungen der Zoologisch-Botanischen Gesellschaft Wien,70, 21–33.

    Karsch, F. (1889) Orthopterologische Beiträge. III. Berliner Entomologische Zeitschrift, 32 [1888], 415–464.

    Karsch, F. (1890). Neue westafrikanische, durch Herrn Premierlieutenant Morgen von Kribi eingesendete Orthopteren. Entomologische Nachrichten, Berlin, 16, 257–276.

    Karsch, F. (1893) Die Insecten der Berglandschaft Adeli im Hinterlande von Togo (Westafrika) nach dem von den Herren Hauptmann Eugen Kling (1888 und 1889) und Dr. Richard Büttner (1890 und 1891) gesammelten Materiale bearbeitet. Berliner Entomologische Zeitschrift, 38, 1–266.

    Krauss, H.A. (1890a) Beitrag zur Kenntnis westafrikanischer Orthopteren. 1. Das Genus Corycus Sauss. Zoologische Jahrbücher, Abteilung für Systematik, Oekologie und Geographie der Tiere, 5, 344–360.

    Krauss, H.A. (1890b) Beitrag zur Kenntnis westafrikanischer Orthopteren. 2. Orthopteren der Guinea-Inseln São Thomé und Rolas, gesammelt von Prof. Dr. Richard Greef. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere, 5 (4), 647–667.

    Le Guillou, E.J.F. (1841) Description de 23 espèces nouvelles d‘Orthoptères, recueillies pendant son Voyage autour du Monde sur la Zelée. Revue Zoologique par la société Cuvierienne, 4, 291–295.

    Linnaeus, C. (1767) Systema Naturae, per Regna tria Naturae, secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. (12th ed.). Laurentius Salvius, Holmiae [Stockholm], 533–1327 pp.

    Liu, X.-W. & Zhang, D.-J. (2007) A study of the genus Conocephalus Thunberg and descriptions of two new species from China (Orthoptera, Conocephalidae). Acta Zootaxonomica Sinica, 32 (2), 438–444.

    Loiseau, C., Melo, M., Lee, Y., Pereira, H., Hanemaaijer, M.J., Lanzaro, G.C. & Cornel, A.J. (2019) High endemism of mosquitoes on São Tomé and Prıncipe Islands: evaluating the general dynamic model in a worldwide island comparison. Insect Conservation and Diversity, 12, 69–79.

    https://doi.org/10.1111/icad.12308

    Massa, B. (2013) Diversity of leaf katydids (Orthoptera: Tettigoniidae: Phaneropterinae) of Dzanga-Ndoki National Park, Central African Republic, with selected records from other African countries. Journal of Orthoptera Research, 22 (2), 125–152.

    https://doi.org/10.1665/034.022.0201

    Massa, B. (2016) The identity of the tropical African Polichne mukonja Griffini, 1908 (Orthoptera, Tettigoniidae, Phaneropterinae). ZooKeys, 621, 37–44.

    https://doi.org/10.3897/zookeys.621.9725

    Massa, B. (2017a) New data and taxa for Orthoptera Tettigoniidae and Acrididae from tropical Africa. Annales de la Société entomologique de France (N.S.), 53, 236–255.

    https://doi.org/10.1080/00379271.2017.1334586

    Massa, B. (2017b) New genera, species and records of Afrotropical Phaneropterinae (Orthoptera, Tettigoniidae) preserved at the Royal Belgian Institute of Natural Sciences, Bruxelles. Zootaxa, 4358 (3), 401–429.

    https://doi.org/10.11646/zootaxa.4358.3.1

    Massa, B., Buzzetti, F.M. & Fontana, P. (2010) Order Orthoptera. In: Van Harten, A. (Ed.), Arthropod Fauna of the United Arab Emirates. Volume 3. Dar Al Ummah Printing, Publishing, Distribution & Advertising, Abu Dhabi, pp. 70–112.

    Massa, B., Fontana, P., Buzzetti F.M., Kleukers, R. & Odé, B. (2012) Orthoptera. Fauna d’Italia 48. Edagricole-New Business Media, Milano, 563 pp.

    Matyot, P. (1998) The orthopteroids of the Seychelles: a threatened island fauna. Journal of Insect Conservation, 2, 235–246.

    https://doi.org/10.1023/A:1009647930445

    McNeill, J. (1901) Papers from the Hopkins Stanford Galapagos Expedition 1898-1899, IV. Entomological results (4): Orthoptera. Proceedings of the Washington Academy of Sciences, 3, 487–506.

    Montealegre-Z, F., Ogden, J., Jonsson, T. & Soulsbury, C.D. (2017) Morphological determinants of signal carrier frequency in katydids (Orthoptera): a comparative analysis using biophysical evidence of wing vibration. Journal of Evolutionary Biology, 30 (11), 2068–2078.

    https://doi.org/10.1111/jeb.13179

    Morse, A.P. (1901) The Xiphidiini of the Pacific Coast. Canadian Entomologist, 33, 201–206.

    https://doi.org/10.4039/Ent33201-7

    Naskrecki, P. (2008a) Sylvan Katydids (Orthoptera: Tettigoniidae: Pseudophyllinae) of the Guinean Forests of West Africa Hotspot: an overview and descriptions of new species. Zootaxa, 1712, 1–41.

    Naskrecki, P. (2008b) New species of arboreal predatory katydids from West Africa (Orthoptera: Tettigoniidae: Meconematinae). Zootaxa, 1732, 1–28.

    NBSAP, II (2016) São Tomé and Principe National Biodiversity Strategy and Action Plan 2015-2020v.2. Ministry of Infrastructure, Natural Resources and Environment.

    OSFO (2018) = Cigliano et al.

    Otte, D. (1997) Orthoptera Species File 7. Tettigonioidea. Orthopterists' Society at the Academy of Natural Sciences of Philadelphia, Philadelphia, 373 pp.

    Paris, M. (1994) Catalogo de Tipos de Ortopteroides (Insecta) de Ignacio Bolivar, I: Blattaria, Mantodea, Phasmoptera y Orthoptera (Stenopelmatoidea, Rhaphidophoroidea, Tettigonioidea, Grylloidea, Tetrigoidea). Eos. Revista Española de Entomologia, 69 (1993), 143–264. Available from: http://hdl.handle.net/10261/170427 (Accessed 27 Feb. 2019)

    Peel, C.V.A., Austen, E.E., Dixey, F.A., Druce, H., Gahan, C.J., Arrow, G.J., McLachlan, R., Burr, M. & Pocock, R.I. (1900). On a collection of insects and Arachnids made in 1895 and 1897 by Mr. C.V.A. Peel. F.Z.S, in Somaliland, with descriptions of new species. Proceedings of the Zoological Society of London, 1900, 4–63. Available from: https://www.biodiversitylibrary.org/item/97409#page/30/mode/1up (Accessed 27 Feb. 2019)

    Pinhey, E. (1974) Odonata of the Northwest Cameroons and particularly of the islands stretching southwards from the Guinea Gulf. Bonner Zoologische Beiträge, 25, 179–212. Available from: https://www.biodiversitylibrary.org/Part/119825#/Summary (Accessed 27 Feb. 2019)

    Pitkin, L.M. (1977) A taxonomic study of the genus Thyridorhoptrum Rehn & Hebard (Orthoptera: Tettigoniidae), with the description of a new species. Journal of Natural History, 11, 645–659.

    Pitkin, L.M. (1980) A revision of the Pacific species of Conocephalus Thunberg (Orthoptera: Tettigoniidae). Bulletin of the British Museum (Natural History), Entomology Series, 41, 315–355.

    Ragge, D.R. (1956) A revision of the genera Phaneroptera Serville and Nephoptera Uvarov (Orthoptera: Tettigoniidae), with conclusions of zoogeographical and evolutionary interest. Proceedings of the Zoological Society of London, 127, 205–283.

    https://doi.org/10.1111/j.1096-3642.1956.tb00471.x

    Ragge, D.R. (1980) A review of the African Phaneropterinae with open tympana (Orthoptera: Tettigoniidae). Bulletin of the British Museum (Natural History), Entomology Series, 40, 67–192. Available from: https://www.biodiversitylibrary.org/part/26800#/summary (Accessed 27 Feb. 2019)

    Ragge, D.R. & Reynolds, W.J. (1998) The songs of the grasshoppers and crickets of western Europe. Harley Books, Colchester, Essex, 591 pp.

    Ragge, D.R. & Roy, R. (1961) VII. Orthoptera Tettigonioidea. Le parc national du Niokolo-Koba (deuxieme fascicule). Mémoires de l' Institut Français d' Afrique Noire, 62, 105–107.

    Redtenbacher, J. (1891) Monographie der Conocephaliden. Verhandlungen der Zoologisch-Botanischen Gesellschaft Wien, 41, 315–562.

    Redtenbacher, J. (1892) Monographische Uebersicht der Mecopodiden. Verhandlungen der Zoologisch-Botanischen Gesellschaft Wien, 42, 183–224.

    Rehn, J.A.G. & Hebard, M. (1915) Studies in American Tettigoniidae (Orthoptera). V. A synopsis of the species of the genus Conocephalus (Xiphidium of authors) found in America, north of Mexico. Transactions of the American Entomological Society, 41, 155–224.

    Röller, O. (2007) Zur Verbreitung und Ökologie der Vierpunkt-Sichelschrecke (Phaneroptera nana) im Siedlungsgebiet von Haßloch/Pfalz. Pollichia-Kurier 23 (4), 19–23.

    Saussure, H. de & Pictet, A. (1897–1899) Locustidae. In: Saussure, H. de, Zehntner, L. & Pictet, A. (Eds.), Biologia Centrali–Americana. Insecta Orthoptera 1, R.H.Porter, London, pp. 285–457.

    Schlüter, T. (2008) Geological Atlas of Africa. Springer, Berlin, Heidelberg. 308 pp.

    https://doi.org/10.1007/978-3-540-76373-4

    Scopoli, J.A. (1786) Deliciae faunae et florae insubricae. Monasterii S.Salvatoris, Ticini, 85 pp.

    Scudder, S.H. (1862) List of Orthoptera collected on trip from Assiniboia to Cumberland. The Canadian Naturalist and Geologist, and Proceedings of the Natural History Society of Montreal, 7, 283–288.

    Scudder, S.H. (1869) Descriptions of new species of Orthoptera in the collection of the American Entomological Society. Transactions of the American Entomological Society, 2, 305–307.

    Scudder, S.H. (1872) Notes on the Orthoptera collected by Dr. Hayden in Nebraska. In: Hayden, F.V. (Ed), Final Report of the U. S. Geological Survey of Nebraska and portions on adjacent territories made under the direction of the commissioner of the General Land Office, part III, United States. General Land Office, Washington, pp. 249–261.

    Scudder, S.H. (1875) Entomological Notes IV. A century of Orthoptera. Decade II. Locustariae. Proceedings of the Boston Society of Natural History, 17, 454–462.

    Serville, J.G.A. (1838) Histoire naturelle des Insectes. Orthoptères. Librairie Encyclopédique de Roret [Collection des suites a Buffon], Paris (Roret), 776 pp.

    Shi, F.-M. & Liang, G.-Q. (1997) Descriptions of two new species of the genus Conocephalus Thunberg (Orthoptera: Conocephalidae). Entomologia Sinica, 4, 211–214.

    Shi, F.-M., Wang, J.-F. & Fu, P. (2005) A review of the genus Conanalus Tinkham (Orthoptera, Tettigonioidea) from China. Acta Zootaxonomica Sinica, 30, 84–86.

    Shi, F.-M. & Zheng, Z.-M. (1994) Two new species of katydids from Sichuan (Orthoptera: Tettigonioidea). Journal of Shanxi Teacher's University Natural Science Edition, 8 (1), 44–46. [Chinese, English summary]

    Shi, F.-M. & Zheng, Z.-M. (1999) A new species of the genus Conocephalus Thunberg (Orthoptera: Conocephalidae) from Sichuan, China. Entomologia Sinica, 6, 219–221.

    https://doi.org/10.1111/j.1744-7917.1999.tb00114.x

    Shorthouse, D.P. (2010) SimpleMappr, an online tool to produce publication-quality point maps. Available from: http://www.simplemappr.net/ (accessed 05 October 2018)

    Sjöstedt, Y. (1926) Zoological results of the Swedish Expedition to Central Afrika 1921. Insecta 17. Phasgonurodea. Arkiv för Zoologi, 18A No.16, 1–10.

    Stål, C. (1857) Orthoptera cursoria och Locustina fran Cafferlandet. Öfversigt Kongliga Vetenskaps–akademiens Förhandlingar, Stockholm, 13 (1856), 165–170.

    Stål, C. (1861) Orthoptera species novas descripsit. In: Kongliga Svenska Fregatten Eugenies Resa, Omkring Jorden under befal af C.A. Virgin, aren 1851-1853, Vetenskapliga Lakttagelser Pa H. M. Konung Oscar der Forstes Befallning, vol. 2, Zoologi. 1. Insecta.—Utgifna of K. Svenska Vetanskaps Akademien 3 [1858–1868], P.A. Norstedt and Soner, Stockholm, pp. 299–350.

    Stål, C. (1873) Orthoptera nova descripsit. Öfversigt Kongliga Vetenskaps–akademiens Förhandlingar, Stockholm, 30, 39–53.

    Thunberg, C.P. (1789) Dissertatio entomologica novas insectorum species, sistens: cuius partem sextam. Cons. Exper. Facult. Med. Upsal., publice ventilandam exhibent, 85–106 pp.

    Uvarov, B.P. (1924) Some new and interesting Orthoptera in the collection of the Ministry of Agriculture, Cairo. Bulletin Ministry of Agriculture Egypt, Technical and Scientific Service, 41, 1–41.

    Uvarov, B.P. (1928) Notes on the types of Orthoptera described by Dr. L. Péringuey. Annals of the South African Museum, 25, 341–357.

    Uvarov, B.P. (1942) New and less known southern Palaearctic Orthoptera. Transactions of the American Entomological Society, 67 (1941), 303–361.

    Vahed, K. (1994) The evolution and function of the spermatophylax in bushcrickets (Orthoptera: Tettigoniidae). Ph.D Thesis. Nottingham University, Nottingham, 227 pp. Available from: http://eprints.nottingham.ac.uk/13737/ (Accessed 27 Feb. 2019)

    Walker, F. (1869) Catalogue of the specimens of Dermaptera Saltatoria in the collection of the British Museum. Part II. Catalogue of Locustidae. British Museum of Natural History, London, 225–423 pp.

    Weidner, H. (1966) Die Entomologischen Sammlungen des Zoologischen Staatsinstituts und Zoologischen Museums Hamburg. VI. Teil. Insecta III. (9.-13. Ordnung: Blattariae, Isoptera, Notoptera, Cheleutoptera, Ensifera). Mitteilungen Hamburger Zoologisches Museum und Institut, 63, 209–264.

    Willemse, C. (1942) On a collection of Indo-Australian, Melanesian and Micronesian Tettigoniidae. Natuurhistorisch Maandblad, 31, 86–88, 94–100, 107–112.

    Willemse, L.P.M., Hochkirch, A., Heller, K.-G., Papapavlou, K., Kati, V. & Tzirkalli, E. (2016) Conocephalus grebenchikovi. The IUCN Red List of Threatened Species 2016, e.T68281269A70624036.

    https://doi.org/10.2305/IUCN.UK.2016-1.RLTS.T68281269A70624036.en

    Willemse, L.P.M., Kleukers, R.M.J.C. & Odé, B. (2018) The Grasshoppers of Greece. EIS Kenniscentrum Insecten & Naturalis Biodiversity Center, Leiden, 439 pp.

    Xia, K.-L. & Liu, X.-W. (1992) Descriptions of three new species of Conocephalini (Orthoptera, Tettigoniidae). Contributions from the Shanghai Institute of Entomology, 9, 162–166.

    Zhou, M., Bi, W.-X. & Liu, X.-W. (2010) The genus Conocephalus (Orthoptera, Tettigonioidea) in China. Zootaxa, 2527, 49–60.

    https://doi.org/10.11646/zootaxa.2527.1.3