Skip to main content Skip to main navigation menu Skip to site footer
Type: Monograph
Published: 2019-07-03
Page range: 1–265
Abstract views: 379
PDF downloaded: 16

A Catalogue and Historical Study of the Odonata of Ecuador

Managing Director, International Odonata Research Institute, 4525 NW 53rd Lane, Gainesville, FL 32653, USA.
Research Associate, Florida State Collection of Arthropods, PO Box 585, Wautoma, WI 54982, USA.
Odonata Anisoptera Zygoptera dragonfly damselfly new records Ecuador

Abstract

We present a list of 425 valid Odonata species from Ecuador, providing a brief overview of each genus, a photograph of a representative species of each genus, and all records known to us for each species. Thirty-eight of the 425 species were previously unrecorded from Ecuador. Data were derived from published records, public and private collections, and field records accumulated since the 1960’s by the authors and others who have contributed specimen information to this study. A historical study and a comprehensive list of synonyms are presented, and questionable species records are discussed. The physiography of the country, including the general climate of each subregion and current pressures on the environment, is briefly discussed. At present, 44 species of Odonata are considered endemic to Ecuador, and eight species are considered endangered, vulnerable, or near threatened on the IUCN Red List.

 

References

  1. Alexander, C.P. (1953) Notes on the Tipulidae of Ecuador (Order Diptera). Part I. Revista Ecuatoriana de Entomologia y Parasitologia, 1 (1), 1–9.

    Altamiranda-Saavedra, M. (2009) Diversidad de Libélulas (Insecta-Odonata) Para dos uso de Suelo, en un Bosque Seco Tropical. Revista Facultad Nacional de Agronomía, 62 (2), 5071–5079.

    Altamiranda-Saavedra, M., Palacino-Rodríguez, F. & Lobo-Hernández, M. (2014) Daily abundance at the breeding site and reproductive behavior of Polythore gigantea (Odonata: Polythoridae). Odonatologica, 43 (3–4), 169–182.

    Álvarez-Covelli, C., Álvarez-Covelli, M.A. & Palacino-Rodríguez, F. (2015) Abdomen or wings? Comparing two body places for marking in Mesamphiagrion laterale (Odonata: Coenagrionidae). Odonatologica, 44 (3), 343–348.

    Amaya-Perilla, C. & Palacino-Rodríguez, F. (2012) An updated list of the Dragonflies (Odonata) of Meta Department, Colombia, with forty-six new department records. Bulletin of American Odonatology, 11 (2), 29–38.

    Amaya-Vallejo, V. & Ledezma, J. (2010) Libélulas (Odonata: Anisoptera) de la Collección Entomológica del Museo de Historia Natural Noel Kempff Mercado, Santa Cruz de la Sierre, Bolivia. Kempffiana, 6 (2), 40–47.

    Amaya-Vallejo, V. & Novelo-Gutiérrez, R. (2011) Larva of Palaemnema mutans Calvert, 1931 (Odonata: Platystictidae). Zootaxa, 3049, 59–63.

    Amaya-Vallejo, V. & Novelo-Gutiérrez, R. (2012) Desmogomphus anchicayensis spec. nov. from Colombia (Anisoptera: Gomphidae). Odonatologica, 41 (1), 25–29.

    Amaya-Vallejo, V., Novelo-Gutiérrez, R. & Realpe, E. (2017) Perigomphus basicornis sp. nov. (Odonata: Gomphidae) from Anchicayá, Colombia. Zootaxa, 4294 (3), 385–400.

    https://doi.org/10.11646/zootaxa.4294.3.10

    Amaya-Vallejo, V., Novelo-Gutiérrez, R. & Realpe, E. (2018) The larva and female of Perigomphus basicornis Amaya-Vallejo, Novelo-Gutiérrez & Realpe, 2017, and the first record of Perigomphus pallidistylus (Belle, 1972) for Colombia (Insecta: Odonata: Gomphidae). PeerJ, 6. e5279.

    https://doi.org/10.7717/peerj.5279

    Asahina, S. (1961) Dragonflies taken by Dr. Sekiguehi in the Galapagos Islands. Publications of the Entomological Laboratory, University of Osaka Prefecture, 6, 1–3.

    Barragan, A., Dangles, O., Cardenas, R. & Onore, G. (2009) The History of Entomology in Ecuador. Annales de la Société entomologique de France, New Series, 45 (4), 410–423.

    https://doi.org/10.1080/00379271.2009.10697626

    Belle, J. (1963) Dragon flies of the genus Zonophora with special reference to its Surinam representatives. Studies on the Fauna Suriname and other Guyanas, 5 (16), 60–69.

    Belle, J. (1966) Surinam dragon-flies of the Agriogomphus complex of genera. Studies on the Fauna Suriname and other Guyanas, 8 (29), 29–60.

    Belle, J. (1970) Studies on South American Gomphidae (Odonata) with special reference to the species from Surinam. Studies of the Fauna of Suriname and other Guyanas, 11 (43), 1–158.

    Belle, J. (1972) Further studies on South American Gomphidae (Odonata). Tijdschrift voor entomologie, 115 (5), 217–240.

    Belle, J. (1973) A revision of the New World genus Progomphus Selys, 1854 (Anisoptera: Gomphidae). Odonatologica, 2 (4), 191–308.

    Belle, J. (1975) On Agriogomphus tumens (Calvert, 1905) with a description of its male (Anisoptera: Gomphidae). Odonatologica, 4 (4), 237–242.

    Belle, J. (1978) The male sex of Aphylla alia Calvert and A. boliviana Belle (Anisoptera: Gomphidae). Odonatologica, 7 (2), 159–162.

    Belle, J. (1979) A new species of Peruviogomphus Klots, 1944 from Ecuador (Anisoptera: Gomphidae). Odonatologica, 8 (2), 111–114.

    Belle, J. (1981) A new species of Phyllogomphoides from Ecuador (Odonata: Gomphidae). Entomologische berichten, 41, 173–176.

    Belle, J. (1983a) On the species of the polygonus group of Progomphus with a description of a new species (Odonata: Gomphidae). Tijdschrift voor entomologie, 126, 137–144.

    Belle, J. (1983b) A review of the genus Zonophora Selys (Odonata: Gomphidae). Tijdschrift voor entomologie, 126, 145–173.

    Belle, J. (1984) A synopsis of the South American species of Phyllogomphoides. Tijdschrift voor entomologie, 127, 79–100.

    Belle, J. (1987) Three new species of Macrothemis from northern South America (Odonata: Libellulidae). Zoologische Mededelingen, 61 (19), 287–294.

    Belle, J. (1988a) A synopsis of the species of Phyllocycla Calvert, with descriptions of four new taxa and a key to the genera of Neotropical Gomphidae (Odonata: Gomphidae). Tijdschrift voor entomologie, 131, 73–102.

    Belle, J. (1988b) Epigomphus gibberosus, a new species from Peru, with lectotype designations for the eligible species of the genus Epigomphus (Odonata: Gomphidae). Tijdschrift voor entomologie, 131, 135–140.

    Belle, J. (1989) A revision of the New World genus Neuraeschna Hagen, 1867 (Odonata: Aeshnidae). Tijdschrift voor entomologie, 132, 259–284.

    Belle, J. (1991a) A visit to the Galapagos Islands. Selysia, 20 (1), 2.

    Belle, J. (1991b) The ultimate instar larvae of the Central American species of Progomphus Selys, with a description of P. belyshevi spec. nov. from Mexico (Anisoptera: Gomphidae). Odonatologica, 20 (1), 9–27.

    Belle, J. (1992a) Studies on ultimate instar larvae of Neotropical Gomphidae, with the description of Tibiagomphus gen. nov. (Anisoptera). Odonatologica, 21 (1), 1–24.

    Belle, J. (1992b) A revision of the South American species of Aphylla Selys, 1854 (Odonata: Gomphidae). Zoologische mededelingen, Leiden, 66 (12), 239–264.

    Belle, J. (1994) Four new species of Aphylla from Brazil (Odonata: Gomphidae). Entomologische Berichten, 54 (7), 138–144.

    Belle, J. (1996) Higher classification of the South American Gomphidae (Odonata). Zoologische mededelingen, Leiden, 70, 297–324

    Belle, J. (1998) Synopsis of the neotropical genus Rhodopygia. Zoologische mededelingen, Leiden, 72 (1), 1–13.

    Bick, G.H. & Bick, J.C. (1985) A revision of the picta group of Polythore, with a description of a new species, P. lamerceda spec. nov., from Peru (Zygoptera: Polythoridae). Odonatologica, 14 (1), 1–28.

    Bick, G.H. & Bick, J.C. (1986) The genus Polythore exclusive of the picta group (Zygoptera: Polythoridae). Odonatologica, 15 (3), 245–273.

    Bick, G.H. & Bick, J.C. (1988) A review of males of the genus Philogenia, with descriptions of five new species from South America (Zygoptera: Megapodagrionidae). Odonatologica, 17 (1), 9–32.

    Bick, G.H. & Bick, J.C. (1990a) A revision of the neotropical genus Cora Selys, 1853 (Zygoptera: Polythoridae). Odonatologica, 19 (2), 117–143.

    Bick, G.H. & Bick, J.C. (1990b) Polythore neopicta spec. nov. from Peru (Odonata: Polythoridae). Opuscula Zoologica Fluminensia, 49, 1–7.

    Bick, G.H. & Bick, J.C. (1991) Two new damselflies: Cora dorada spec. nov., from Ecuador and C. parda spec. nov. from Peru (Zygoptera: Polythoridae). Odonatologica, 20 (4), 453–458.

    Bick, G.H. & Bick, J.C. (1992) A study of the family Polythoridae, with details on the genus Euthore Selys, 1869 (Zygoptera). Odonatologica, 21 (3), 275–288.

    Bick, G.H. & Bick, J.C. (1995) A review of the genus Telebasis with descriptions of eight new species (Zygoptera: Coenagrionidae). Odonatologica, 24 (1), 11–44.

    Biodiversity Heritage Library (2019) Available from http://www.biodiversitylibrary.org/ (accessed 14 May 2019)

    Borror, D.J. (1931) The genus Oligoclada (Odonata). Miscellaneous Publications of the Museum of Zoology, University of Michigan, 22, 1–42.

    Borror, D.J. (1942) A revision of the libelluline genus Erythrodiplax (Odonata). Ohio State University, Columbus, Contributions in Zoology and Entomology, No. 4 (Biological Series), 1–286.

    Borror, D.J. (1945) A key to the new world genus of Libellulidae (Odonata). Annals of the Entomological Society of America, 38 (2), 168–194.

    https://doi.org/10.1093/aesa/38.2.168

    Borror, D.J. (1957) New Erythrodiplax from Venezuela (Odonata: Libellulidae). Acta Biologica Venezuelica, 2 (5), 31–42.

    Bota-Sierra, C.A. (2012) Rediscovery of Proneura prolongata (Zygoptera: Protoneuridae) and other new Odonata records from Colombian Amazon. Agrion, 16 (1), 34–41.

    Bota-Sierra, C.A. (2014a) A brief look at the Odonata from the Páramo ecosystems in Colombia, with the descriptions of Oxyallagma colombianum sp. nov. and Rhionaeschna caligo sp. nov. (Odonata: Coenagrionidae, Aeshnidae, Libellulidae). Zootaxa, 3856 (2), 192–210.

    https://doi.org/10.11646/zootaxa.3856.2.2

    Bota-Sierra, C.A. (2014b) Nine new records of Odonata for Colombia from the Orinoco Basin (Lestidae, Calopterygidae, Heteragrionidae, Coenagrionidae, Libellulidae). Notulae odonatologicae, 8 (4), 77–116.

    Bota-Sierra, C.A. (2017a) Exploring Colombian Odonata. Agrion, 21 (2), 78–80.

    Bota-Sierra, C.A. (2017b) Two new species of the family Philogeniidae (Odonata: Zygoptera) from the Western Colombian Andes. International Journal of Odonatology, 20 (3–4), 137–150.

    https://doi.org/10.1080/13887890.2017.1344733

    Bota-Sierra, C.A., Baena-Bejarano, N. & Bermúdez, C. (2010) First records of Gomphomacromia fallax (Odonata, Corduliidae) from Colombia. Revista Colombiana de Entomología, 36 (2), 333–334.

    Bota-Sierra, C.A., Corso, A., Janni, O., Sandoval-H, J. & Viganò, M. (2018) Seventeen new dragonfly records from Colombia and the confirmation of the synonymy of Philogenia monotis and P. tinalandia (Insecta: Odonata). International Journal of Odonatology, 21 (2), 115–127.

    https://doi.org/10.1080/13887890.2018.1462262

    Bota-Sierra, C.A., Moreno-Arias, C. & Faasen, T. (2015) Preliminary list of Odonata from the Colombian Amazon, with descriptions of Inpabasis nigridorsum sp. nov. & Diaphlebia richteri sp. nov. (Coenagrionidae & Gomphidae). International Journal of Odonatology, 18 (3), 249–268.

    https://doi.org/10.1080/13887890.2015.1081637

    Bota-Sierra, C.A., Maufray, B., Palacino-Rodríguez, F., Hofmann, J., Tennessen, K.J., Rache, L., & Tognelli, M.F. (2016) Estado de conservación de las libélulas de los Andes Tropicales. In: Tognelli, M.F., Lasso, C.A., Bota-Sierra, C.A., Jiménez-Segura, L.F. & Cox, N.A. (Eds.), Estado de Conservación y Distribución de la Biodiversidad de Agua Dulce en los Andes Tropicales. UICN, Gland, Suiza, Cambridge and Arlington, pp. 67–86.

    Bota-Sierra, C.A. & Novelo-Gutierrez, R. (2017) The genus Heteragrion (Odonata: Zygoptera) in Northwestern Colombia, with the description of Heteragrion tatama sp. nov. Zootaxa, 4347 (3), 553–571.

    https://doi.org/10.11646/zootaxa.4347.3.8

    Bota-Sierra, C.A., Novelo-Gutierrez, R. & Amaya-Vallejo, V. (2017) The rediscovery and redescription of Epigomphus pechumani Belle, 1970 (Odonata: Gomphidae), with a description of its female from the Western Colombian Andes. Zootaxa, 4306 (3), 419–427.

    https://doi.org/10.11646/zootaxa.4306.3.8

    Bota-Sierra, C.A., Sanchez-Herrera, M. & Palacino-Rodriguez, F. (2018) Odonata from protected areas in Colombia with new records and description of Cora verapax sp. nov. (Zygoptera: Polythoridae). Zootaxa, 4462 (1), 115–131.

    https://doi.org/10.11646/zootaxa.4462.1.5

    Bota-Sierra, C.A. & Sandoval-H, J. (2017) The female of Oreiallagma oreas (Odonata: Coenagrionidae), with notes on the species natural history. International Journal of Odonatology, 20 (3-4), 165–172.

    https://doi.org/10.1080/13887890.2017.1362363

    Bota-Sierra, C.A. & Wolff Echeverri, M.I. (2013) Taxonomic revision of Mesamphiagrion Kennedy, 1920 from Colombia (Odonata: Coenagrionidae), with description of four new species. Zootaxa, 3718 (5), 401–440.

    https://doi.org/10.11646/zootaxa.3718.5.1

    Brauer, F. (1865a) Zweiter Bericht über die auf der Weltfahrt der Kais. Fregatte Novara gesammelten Neuropteren. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 15, 501–512.

    Brauer, F. (1865b) Veirter Bericht über die auf der Weltfahrt der Kais. Fregatte Novara gesammelten Neuropteren. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 15, 903–908.

    Brauer, F. (1868) Verzeichniss der bis jetzt bekannten Neuropteren im Sinne Linnés. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 18, 711–742.

    Bridges, C.A. (1993) Catalogue of the family-group, genus-group and species-group names of the Odonata of the world. 3rd Edition. C.A. Bridges, Urbana, Illinois, 905 pp.

    https://doi.org/10.5962/bhl.title.15283

    Brown, F.M. (1941) A gazeteer of entomological stations in Ecuador. Annals of the Entomological Society of America, 34 (4), 809–851.

    https://doi.org/10.1093/aesa/34.4.809

    Brown, F.M. & Mcintyre, W.C. (date unknown) Collecting stations in the northeastern part of Ecuador. Part I. the Río Pastaza watershed. Unpublished manuscript with maps in the International Odonata Research Library, Florida State Collection of Arthropods, Gainesville, Florida, 16 + 2 (maps) pp. [M.J. Westfall, Jr. archives]

    Burmeister, H. (1839) Neuroptera. In: Handbuch der Entomologie. 2 (2). T.C.F. Enslin, Berlin, pp. 757–1050.

    Bybee, S.M. & Tennessen, K.J. (2008) Description of the female and nymph of Philogenia mangosisa from southern Ecuador (Odonata: Megapodagrionidae). Zootaxa, 1787, 63–68.

    Byers, C.F. (1939) A study of the dragonflies of the genus Progomphus (Gomphoides) with a description of a new species. Proceedings of the Florida Academy of Science, 4, 19–85.

    Calvert, P.P. (1898) The Odonate genus Macrothemis and its allies. Proceedings of the Boston Society of Natural History, 28 (12), 301–332.

    Calvert, P.P. (1901) Odonata. In: Godman, F.D. & Salvin, O. (Eds.), Biologia Centrali-Americana—Insecta Neuroptera. R.H. Porter & Dulau Co., London, pp. 17–72.

    Calvert, P.P. (1902a) Odonata. In: Godman, F.D. & Salvin, O. (Eds.), Biologia Centrali-Americana—Insecta Neuroptera. R.H. Porter & Dulau Co., London, pp. 73–128.

    Calvert, P.P. (1902b) Illustrations of Odonata—Argia by Hermann A. Hagen. With a list and bibliography of the species by Philip P. Calvert. Bulletin of the Museum of Comparative Zoology at Harvard College, 39 (4), 103–120.

    Calvert, P.P. (1903) Odonata. In: Godman, F.D. & Salvin, O. (Eds.), Biologia Centrali-Americana—Insecta Neuroptera. R.H. Porter & Dulau Co., London, pp. 129–144.

    Calvert, P.P. (1905) Odonata. In: Godman, F.D. & Salvin, O. (Eds.), Biologia Centrali-Americana—Insecta. Neuroptera. R.H. Porter & Dulau Co., London, pp. 145–212.

    Calvert, P.P. (1906) Odonata. In: Godman, F.D. & Salvin, O. (Eds.), Biologia Centrali-Americana—Insecta Neuroptera. R.H. Porter & Dulau Co., London, pp. 213–308.

    Calvert, P.P. (1907) Odonata. In: Godman, F.D. & Salvin, O. (Eds.), Biologia Centrali-Americana—Insecta Neuroptera. R.H. Porter & Dulau Co., London, pp. 309–404.

    Calvert, P.P. (1909) Contributions to a knowledge of the Odonata of the Neotropical region, exclusive of Mexico and Central America. Annals of the Carnegie Museum, 6 (3), 73–280.

    Calvert, P.P. (1920) The Costa Rican species of Epigomphus and their mutual adaptations (Odonata). Transactions of the American Entomological Society, 46, 323–354.

    Calvert, P.P. (1924) The generic characters and the species of Philogenia Selys (Odonata: Agrionidae). Transactions of the American Entomological Society, 50, 1–56.

    Calvert, P.P. (1931) The generic characters and the species of Palaemnema (Odonata: Agrionidae). Transactions of the American Entomological Society, 62, 1–110.

    Calvert, Philip P. (1934) The Rates of Growth, Larval Development and Seasonal Distribution of Dragonflies of the Genus Anax (Odonata: Aeshnidæ). Proceedings of the American Philosophical Society, 73 (1), 1–70.

    Calvert, P.P. (1947a) Aeshna psilus, a new species of the group of Ae. cornigera Brauer (Odonata: Aeshnidae). Notulae naturae of the Academy of Natural Sciences of Philadelphia, 194, 1–11.

    Calvert, P.P. (1947b) Odonata of Voyages under the auspices of the New York Zoological Society. Entomological News, 58 (9), 227–230.

    Calvert, P. P. (1948) Odonata (Dragonflies) of Kartabo, Bartica District, British Guiana. Zoologica, 33 (2), 47–87.

    Calvert, P.P. (1952) New taxonomic entities in Neotropical Aeshnas (Odonata: Aeshnidae). Entomological News, 63 (10), 253–264.

    Calvert, P.P. (1956) The Neotropical species of the “subgenus Aeschna” sensu selysii 1883 (Odonata). Memoirs of the American Entomological Society, 15, 1–251.

    Campion, H. (1920) Some new or little-known gomphine dragonflies from South America. Annals and Magazine of Natural History, Series 9, 6 (31), 130–141.

    https://doi.org/10.1080/00222932008632424

    Campos, F. (1900) Observaciones Entomologicas. Revista chilena de historia natural, 4 (4), 47–49.

    Campos, F. (1922) Catálogo Sistemático y Sinonímico de los Odonatos del Ecuador. Revista del Colegio Nacional Vicente Rocafuerte, 8–9, 1–75.

    Campos, F. (1925) Suplemento No. 1. al Catálogo Sistemático y Sinonímico de los Odonatos del Ecuador. Revista del Colegio Nacional Vicente Rocafuerte, 19–20, 35–42.

    Campos, F. (1927) Las Libelulas de la Provincia de El Oro. Revista del Colegio Nacional Vicente Rocafuerte, 30–31, 79–81.

    Campos, F. (1930) Una Colonia de Gamasidos afectando a nuestro grillo comun (Gryllus assimilis Fabr.). Revista del Colegio Nacional Vicente Rocafuerte, 40–41, 19–20.

    Campos, F. (1931) La luz electrica y su influencia en los odonatos. Revista del Colegio Nacional Vicente Rocafuerte, 13, 3–5.

    Campos, F. (1932) La Invasion de Libelulas en La Tarde de ayer. El Telegrafo, Ano XLIV, 16.853 Lunes 14 de Marzo 1931, 1.

    Carvalho A.L. (1992) Revalidation of the genus Remartinia Navás, 1911, with description of a new species and a key to the genera of Neotropical Aeshnidae (Anisoptera). Odonatologica, 21 (3), 289–298.

    Carvalho, A.L. & Werneck-de-Carvalho, P.C. (2005) Descrição da larva de Orthemis cultriformis Calvert, 1899 (Insecta, Odonata, Libellulidae). Archivos do Museo Nacional, Río de Janeiro, 63, 267–273.

    Central Intelligence Agency (2017) The World Factbook. Available from: https://www.cia.gov/library/publications/the-world-factbook/geos/ec.html (accessed 8 February 2017)

    Clements, J.F., Schulenberg, T.S, Iliff, M.J., Roberson, D., Fredericks, T.A., Sullivan, B.L. & Wood, C.L. (2016) The eBird/Clements checklist of birds of the world. Version 2016. Available from: from: http://www.birds.cornell.edu/clementschecklist/download/ (accessed 11 August 2016)

    Cook, C. (1989) Philogenia redunca, a new dragonfly from Ecuador (Odonata: Megapodagrionidae). Florida Entomologist, 72 (3), 419–424.

    https://doi.org/10.2307/3495177

    Corbet, P.S. (1999) Dragonflies: Behavior and ecology of Odonata. Cornell University Press, Ithaca, New York and Harley Books, Colchester, 864 pp.

    Cordero-Rivera, A., Encalada, A.C., Sánchez–Guillén, R.A., Santolamazza–Carbone, S. & von Ellenrieder, N. (2016) The status of Rhionaeschna galapagoensis (Currie, 1901) with notes on its biology and a description of its ultimate instar larva (Odonata, Aeshnidae). Animal Biodiversity and Conservation, 39 (1), 45–63.

    https://doi.org/10.32800/abc.2016.39.0045

    Costa, J.M. (1990) Macrothemis newtoni spec. nov., a new dragonfly from Brazil (Anisoptera: Libellulidae). Odonatologica, 19 (1), 75–79.

    Costa, J.M. & Santos, T.C. (1997) Intra- and interspecific variation in the genus Uracis Rambur, 1842, with a key to the known species (Anisoptera: Libellulidae). Odonatologica, 26 (1), 1–7.

    Costa, J.M. & Santos, T.C. (2009) Description of the larva of Orthemis schmidti (Odonata, Libellulidae). Iheringia, Série Zoologia, 99 (2), 129–131.

    https://doi.org/10.1590/S0073-47212009000200002

    Cowley, J. (1934) Notes on Odonata (Corduliidae) of the Neotropical region. Stylops, 3, 91–95.

    https://doi.org/10.1111/j.1365-3113.1934.tb01553.x

    Cowley, J. (1941) A new species of Protoneura from Peru, and a review of the group of Protoneura tenuis Selys (Odonata, Protoneuridae). Transactions of the Entomological Society of London, 91 (6), 145–173.

    https://doi.org/10.1111/j.1365-2311.1941.tb00258.x

    Currie, R.P. (1901) Papers from the Hopkins Stanford Galapagos Expedition, 1898–1899. Proceedings of the Washington Academy of Sciences, 3, 381–389.

    https://doi.org/10.5962/bhl.part.26342

    Daigle, J.J. (1996) The rat patrol's excellent Ecuador adventure! Argia, 8 (3), 10–13.

    Daigle, J.J. (2000) Metaleptobasis mauffrayi spec. nov. from Ecuador and Peru (Zygoptera: Coenagrionidae). Odonatologica, 29 (4), 325–328.

    Daigle, J.J. (2002) Telebasis bickorum spec. nov. from Bolivia (Zygoptera: Coenagrionidae). Odonatologica, 32 (2), 177–180.

    Daigle, J.J. (2003) Metaleptobasis minteri spec. nov. from Ecuador (Zygoptera: Coenagrionidae). Odonatologica, 32 (4), 371–374.

    Daigle, J.J. (2005) Heteragrion bickorum spec. nov. from Ecuador (Zygoptera: Megapodagrionidae). Odonatologica, 34 (2), 165–168.

    Daigle, J.J. (2014) Two new Heteropodagrion species from Ecuador (Odonata: Megapodagrionidae). Odonatologica, 43 (1–2), 35–42.

    Daigle, J.J., Dunkle, S., Mauffray, B & Tennessen, K.J. (1996) More on the rat pack in Ecuador. Argia, 8 (4), 18.

    Daigle, J.J., Mauffray, B & Tennessen, K.J. (1997) Go south, young men, go south. Argia, 9 (1), 11–13.

    Daigle, J.J. & Tennessen, K.J. (2000) Heteragrion cooki spec. nov. from Ecuador (Zygoptera: Megapodagrionidae). Odonatologica, 29 (3), 255–259.

    Dalzochio, M.S. (2009) Descrição da larva de último estádio de Micrathyria pseudeximia Westfall (Odonata, Libellulidae). EntomoBrasilis, 2 (2), 54–57.

    https://doi.org/10.12741/ebrasilis.v2i2.56

    Dalzochio, M.S., Périco, E., Renner, S. & Sahlén, G. (2018) Description of the final stadium larva of Erythrodiplax media (Odonata: Libellulidae) with preliminary key to known South American larvae in the genus. International Journal of Odonatology, 21 (2), 93–104.

    https://doi.org/10.1080/13887890.2018.1462260

    Davies, D.A.L. & Tobin, P. (1984) The dragonflies of the world: a systematic list of the extant species of Odonata. Vol 1 Zygoptera, Anisozygoptera. Societas Internationalis Odonatologica Rapid Communications, 3 (Supplement), 1–127.

    Davies, D.A.L. & Tobin, P. (1985) The dragonflies of the world: a systematic list of the extant species of Odonata. Vol. 2. Anisoptera. Societas Internationalis Odonatologica Rapid Communications, 5 (Supplement), 1–151.

    De Marmels., J.C. (1982) Archilestes guayaraca, nuevo Zygoptera de Venezuela (Odonata: Lestidae). Boletin de Entomologia Venezolana, New Series, 2 (14), 111–113.

    De Marmels, J.C. (1987) On the type specimens of some neotropical Megapodagrionidae, with a description of Heteragrion pemon spec. nov. and Oxystigma caerulans spec. nov. from Venezuela (Zygoptera). Odonatologica, 16 (3), 225–238.

    De Marmels, J.C. (1988) Odonata del estado Tachira. Revista Cientifica Unet, 2 (1), 91–111.

    De Marmels, J.C. (1989) Odonata or dragonflies from Cerro de la Neblina and the adjacent lowland between the Río Baria. The Casiquiare and the Río Negro (Venezuela). I. Adults. Academia de las Ciencias Fisicas, Matematicas y Naturales, Caracas, Venzeuela, 25, 1–87.

    De Marmels, J.C. (1990) An updated checklist of the Odonata of Venezuela. Odonatologica, 19 (4), 333–345.

    De Marmels, J.C. (1992) Caballitos del diablo (Odonata) de las Sierras de Tapirapeco y Unturan, en el extremo sur de Venezuela. [Dragonflies (Odonata) from the Sierras of Tapirapeco and Unturan, in the extreme south of Venezuela.] Acta Biológica Venezuelica, 14 (1), 57–78.

    De Marmels, J.C. (1999) Rare Venezuelan dragonflies (Odonata) evaluated for their possible inclusion in the national red data book. International Journal of Odonatology, 2 (1), 55–67.

    https://doi.org/10.1080/13887890.1999.9748112

    De Marmels, J.C. (2001a) Revision of Megapodagrion Selys, 1886. (Insecta, Odonata, Megapodagrionidae). Dissertation zur Erlangung der Naturwissenschaftlichen Dokenorwürde (Dr. sc. nat) vorgelegt der Mathematisch-naturwissenschaftlichen Fakultät der Universität Zürich, Zürich, 220 pp.

    De Marmels, J.C. (2001b) Aeshna (Hesperaeschna) condor sp. Nov. from the Venezuelan Andes, with a redescription of A. (H.) joannisi, comments on other species, and descriptions of larvae (Odonata: Aeshnidae). International Journal of Odonatology, 4 (2), 119–134.

    https://doi.org/10.1080/13887890.2001.9748166

    De Marmels, J.C. (2001c) Sympetrum paramo sp. n. (Odonata: Libellulidae) from the Venezuelan high Andes, with a key to the species of Sympetrum Newman, 1833 found in Venezuela. Entomotropica, 16, 15–19.

    De Marmels, J. (2002) Phylogenetic relationships of Priscagrion Zhou & Wilson, 2001, with a description of Teinopodagrion croizati spec. nov. from Ecuador (Zygoptera: Megapodagrionidae). Odonatologica, 31 (4), 389–394.

    De Marmels, J. (2006) Una pequeña colección de libélulas (Odonata) de Colombia. Entomotropica, 21 (1), 69–71.

    De Marmels, J. (2012) Review of the “metallic group” of species of Argia Rambur known from Venezuela, with description of the larva of Argia Jocosa Hagen in Selys, 1865 (Odonata: Coenagrionidae). International Journal of Odonatology, 15 (3), 249–262.

    https://doi.org/10.1080/13887890.2012.717906

    De Marmels, J. (2016) Lista de los Odonata de Venezuela. Available from: http://odonata.miza-ucv.org.ve/sites/odonata.miza-ucv.org.ve/files/2016-04-13-Checklist_Odonata_Venezuela.pdf (accessed 19 June 2017)

    De Marmels, J. & Garrison, R.W. (2005) Review of the genus Leptagrion in Venezuela with new synonymies and descriptions of a new genus, Bromeliagrion, and a new species, B. rehni (Zygoptera: Coenagrionidae). Canadian Entomologist, 137 (3), 257–273.

    https://doi.org/10.4039/n04-096

    De Marmels, J. & Neiss, U.G. (2013) Description of the larva of Neuraeschna claviforcipata Martin, 1909 (Insecta: Odonata: Aeshnidae). Zootaxa, 3721 (1), 97–100.

    https://doi.org/10.11646/zootaxa.3721.1.7

    De Marmels, J. & Rácenis, J. (1982) An analysis of the cophysa-group of Tramea Hagen, with descriptions of two new species (Anisoptera: Libellulidae). Odonatologica, 11 (2), 109–128.

    de Souza, Luiz Onofre I., Costa, J.M. & Santos, T.C. (2010) Revalidation of Acanthagrion cuyabae (Odonata, Coenagrionidae) and description of the ♀, with a key to the Brazilian species of the viridescens group. Iheringia, Série. Zoologia, 100 (1), 79–83.

    https://doi.org/10.1590/S0073-47212010000100011

    de Souza, M.M., Pires, E.P, Brunismann, Â.G., Lucas Rocha Milani, L.R. & Pinto, Â.P. (2017) Dragonflies and damselflies (Odonata) from the wetland of the Rio Pandeiros, northern region of Minas Gerais State, Brazil, with a description of the male of Archaeogomphus vanbrinki Machado (Anisoptera: Gomphidae). International Journal of Odonatology, 20 (1), 13–26.

    https://doi.org/10.1080/13887890.2017.1281848

    Dijkstra, K.D.B., Kalkman, V.J., Dow, R.A., Stoks, F.R. & van Tol, J. (2014) Redefining the damselfly families: a comprehensive molecular phylogeny of Zygoptera (Odonata). Systematic Entomology, 39 (1), 68–96.

    https://doi.org/10.1111/syen.12035

    Dijkstra, K.D.B., Bechly, G., Bybee, S.M., Dow, R.A, Dumont, H.J., Fleck, G., Garrison, R.W., Hämäläinen, M., Kalkman, V.J, Karube, H., May, M.L., Orr, A.G., Paulson, D.R., Rehn, A.C., Theischinger, G., Trueman, J.W.H., Tol, J.V., von Ellenrieder, N. & Ware, J. (2013) The classification and diversity of dragonflies and damselflies (Odonata). In: Zhang, Z.-Q. (Ed.) Animal Biodiversity: An Outline of Higher-level Classification and Survey of Taxonomic Richness (Addenda 2013). Zootaxa, 3703 (1), 36–45.

    https://doi.org/10.11646/zootaxa.3703.1.9

    Dijkstra, K., Kipping, J. & Meziere, N. (2015) Sixty new dragonfly and damselfly species from Africa (Odonata). Odonatologica, 44 (4), 447–678.

    Donoso, D.A, Salazar, F., Maza, F., Cardenas, RE. & Odangles, O. (2009) Diversity and distribution of type specimens deposited in the Invertebrate section of the Museum of Zoology QCAZ, Quito, Ecuador. Annales de la Société entomologique de France, New Series, 45 (4), 437–454.

    https://doi.org/10.1080/00379271.2009.10697628

    Donnelly, T.W. (1989) Three new species of Epigomphus from Belize and Mexico (Odonata: Gomphidae). Florida Entomologist, 72 (3), 428–435.

    https://doi.org/10.2307/3495179

    Donnelly, T.W. (1992) The Odonata of Central Panama and their position in the neotropical odonate fauna, with a checklist, and descriptions of new species. In: Quintero, D. & Aiello, A. (Eds.), Insects of Panama and Mesoamerica: Selected studies. Oxford University Press, Oxford, pp. 52–90.

    Donnelly, T.W. (1995) Orthemis ferruginea – an adventure in Caribbean biogeography. Argia, 7 (4), 9–12.

    Donnelly, T.W. (1996) The status of Lestes apollinaris Navás and L. henshawi Calvert. Bulletin of American Odonatology, 4 (3), 69–74.

    Donnelly, T.W. (1998) Ecuador epilogue: a trip to La Selva. Argia, 9 (4), 15–16.

    Donnelly, T.W. (2001) Rediscovery of Orthemis sulphurata. Argia, 13 (2), 11–12.

    Dunkle, S.W. (1981) Dunkle and Knopf Blitz Ecuador and Colombia. Selysia, 10 (1), 5–6.

    Dunkle, S.W. (1982) Perithemis rubita spec. nov., a new dragonfly from Ecuador (Anisoptera: Libellulidae). Odonatologica, 11 (1), 33–39.

    Dunkle, S.W. (1986) Four new species of Philogenia damselflies from Ecuador and Columbia (Zygoptera: Megapodagrionidae). Odonatologica, 15 (1), 43–50.

    Dunkle, S.W. (1989) Dragonflies of the Florida Peninsula, Bermuda, and the Bahamas. Scientific Publishers, Gainesville, Florida, 155 pp.

    Dunkle, S.W. (1991a) Aeolagrion axine spec. nov., a new damselfly from Ecuador (Zygoptera: Coenagrionidae). Odonatologica, 20 (2), 239–244.

    Dunkle, S.W. (1991b) Review of the neotropical damselfly family Dicteriadidae (new spelling), with an annotated bibliography (Zygoptera). Odonatologica, 20 (4), 401–416.

    Dunkle, S.W. (1993) The female of Aphylla silvatica Belle from Peru (Anisoptera: Gomphidae). Odonatologica, 22 (4), 465–468.

    Dunkle, S.W. (1995) Geographical variation in Micrathyria mengeri Ris, with a description of M. mengeri watsoni ssp. nov. (Anisoptera: Libellulidae). Odonatologica, 24 (1), 45–51.

    Enciclopedia del Ecuador (2017) Personajes Historicos -Francisco Campos Rivadeneira. Available from: http://www.enciclopediadelecuador.com/personajes-historicos/francisco-campos-rivadeneira/ (accessed 16 February 2017)

    Encyclopedia.com (2007) “Ecuador.” Worldmark Encyclopedia of Nations. Available from: http://www.encyclopedia.com/doc/1G2-2586700155.html (accessed 10 February 2016)

    Encyclopedia.com (2017) Ecuador. Worldmark Encyclopedia of Nations. Available from: http://www.encyclopedia.com/places/latin-america-and-caribbean/south-american-political-geography/ecuador (accessed 10 February 2017)

    Evenhuis, N.L. (2011) The insect and spider collections of the world. The insect and spider collections of the world website. Available from: http://hbs.bishopmuseum.org/codens/ (accessed 20 March 2016)

    Fleck, G. (2004) Contribution a la connaissance des Odonates de Guyane française: les larves des Macrothemis pumila Karsch, 1889 et de Brechmorhoga praedatrix Calvert, 1909. Notes biologiques et conséquences taxonomiques (Anisoptera: Libellulidae). Annales de la Société entomologique de France, 40, 177–184.

    https://doi.org/10.1080/00379271.2004.10697414

    Fleck, G., De Marmels, J. & Grand, D. (2004) La larve de Tholymis citrina Hagen, 1867 (Odonata, Anisoptera, Libellulidae). Bulletin de la Société entomologique de France, 109, 455–457.

    Fleck, G. & Neiss, U.G. (2012) The larva of the genus Paracordulia Martin, 1907 (Odonata: Corduliidae s. s.) and a generic key to the larvae of Corduliidae s. l. occurring in South America. Zootaxa, 3412 (1), 62–68.

    https://doi.org/10.11646/zootaxa.3412.1.4

    Förster, F. (1914) Beiträge zu den Gattungen und Arten der Libellen (N. III). Archiv für Naturgeschichte, 80, 59–83.

    https://doi.org/10.5962/bhl.part.26477

    Förster, S. (2001) The Dragonflies of Central America Exclusive of Mexico and the West Indies, A Guide to Their Identification. 2nd Edition. Gunnar Rehfeld, Wolfenbüttel, 141 pp.

    Fraser, F.C. (1946a) Notes on Amazonian Odonata in the Leeds Museum. Part I. Transactions of the Entomological Society of London, 96 (2), 11–46.

    https://doi.org/10.1111/j.1365-2311.1946.tb00441.x

    Fraser, F.C. (1946b) Notes on Amazonian Odonata in the Leeds Museum. Part II. Transactions of the Entomological Society of London, 97 (18), 443–472.

    https://doi.org/10.1111/j.1365-2311.1946.tb00272.x

    Garrison, R.W. (1982) Archilestes neblina, a new damselfly from Costa Rica, with comments on the variability of A. latialatus Donnelly (Odonata: Lestidae). Occasional Papers of the Museum of Zoology, University of Michigan, 702, 1–12.

    Garrison, R.W. (1990) A synopsis of the genus Hetaerina with descriptions of four new species (Odonata: Calopterygidae). Transactions of the American Entomological Society, 116 (1), 175–259.

    Garrison, R.W. (1999) The genus Neoneura, with keys and description of a new species, Neoneura jurzitzai spec. nov. (Zygoptera: Protoneuridae). Odonatologica, 28 (4), 343–375.

    Garrison, R.W. (2004) An analysis of the Psaironeura tenuissima complex, including synonymy of P. machadoi De Marmels with P. bifurcata Sjöstedt (Zygoptera: Protoneuridae). Odonatologica, 33 (1), 83–89.

    Garrison, R.W. (2006) A synopsis of the genera Mnesarete Cowley, Bryoplathanon gen. nov., and Ormenophlebia gen. nov. (Odonata Calopterygidae). Contributions to Science Series, 506, 1–84.

    Garrison, R.W. (2009) A synopsis of the genus Telebasis (Odonata: Coenagrionidae). International Journal of Odonatology, 12 (1), 1–121.

    https://doi.org/10.1080/13887890.2009.9748331

    Garrison, R.W. (2014) Review of Oxystigma Selys with the synonymy of Oxystigma williamsoni Geijskes (Odonata Heteragrionidae). Zootaxa, 3780 (2), 347–364.

    https://doi.org/10.11646/zootaxa.3780.2.7

    Garrison, R.W. & Ramón Cabrera, G.M. (2019) Argia mauffrayi n. sp. from Ecuador (Odonata: Coenagrionidae). Zootaxa, 4545 (2), 286–292.

    https://doi.org/10.11646/zootaxa.4545.2.8

    Garrison, R.W. & von Ellenrieder, N. (2005) Neuragrion mysticum (Odonata: Megapodagrionidae) demystified. The Canadian Entomologist, 137 (2), 169–173.

    https://doi.org/10.4039/n04-097

    Garrison, R.W. & von Ellenrieder, N. (2007) The true Argia difficilis Selys, 1865, with the description of Argia yungensis sp. nov. (Odonata: Coenagrionidae). Transactions of the American Entomological Society, 133 (1–2), 189–204.

    https://doi.org/10.3157/0002-8320(2007)133[189:TTADSW]2.0.CO;2

    Garrison, R.W. & von Ellenrieder, N. (2009) Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa, 2145, 47–68.

    Garrison, R.W. & von Ellenrieder, N. (2010) Redefinition of Leptobasis Selys with the synonymy of Chrysobasis Racenis and description of L. mauffrayi sp. nov. from Peru (Odonata: Coenagrionidae). Zootaxa, 2438 (1), 1–36.

    https://doi.org/10.11646/zootaxa.2438.1.1

    Garrison, R.W. & von Ellenrieder, N. (2015) Damselflies of the genus Argia of the Guiana Shield (Odonata: Coenagrionidae). Zootaxa, 4042 (1), 1–85.

    https://doi.org/10.11646/zootaxa.4042.1.1

    Garrison, R.W. & von Ellenrieder, N. (2016) New World Odonata List (NWOL). A Synonymic List of the New World Odonata. Revised version. 18 February 2016. Available from: http://www.odonatacentral.org/docs/NWOL.pdf (accessed 15 February 2017)

    Garrison, R.W. & von Ellenrieder, N. (2017) New species of the damselfly genus Argia from Mexico, Central America and Ecuador with an emphasis on Costa Rica (Insecta: Odonata: Coenagrionidae). Zootaxa, 4235 (1), 1–93.

    https://doi.org/10.11646/zootaxa.4235.1.1

    Garrison, R.W. & von Ellenrieder, N. (2018) Damselflies of the genus Argia (Odonata: Coenagrionidae) from Ecuador with descriptions of five new species. Zootaxa, 4470 (1), 1–69.

    https://doi.org/10.11646/zootaxa.4470.1.1

    Garrison, R.W., von Ellenrieder, N. & Louton, J.A. (2006) The Dragonfly Genera (Odonata: Anisoptera of the New World. An Illustrated and Annotated Key. Johns Hopkins University Press, Baltimore, 368 pp.

    Garrison, R.W., von Ellenrieder, N. & Louton, J.A. (2010) Damselfly Genera of the New World. An Illustrated and Annotated Key to the Zygoptera. Johns Hopkins University Press, Baltimore, 490 pp.

    Garrison, R.W., von Ellenrieder, N. & O’Brien, M.F. (2003) An annotated list of the name-bearing types of species-group names in Odonata preserved in the University of Michigan Museum of Zoology. Occasional Papers of the Museum of Zoology, University of Michigan, 736, 1–73.

    Geijskes, D.C. (1943) Notes on Odonata of Surinam. III. The genus Coryphaeschna, with descriptions of a new species and the nymph of C. virens. Entomological News, 54 (3), 61–72.

    Geijskes, D.C. (1948) What is Oligoclada abbreviata (Rambur, 1942)? (Odonata: Libellulidae). Zoologische Mededelingen, Leiden, 58, 175–185.

    Geijskes, D.C. (1951) Notes on Odonata of Surinam. V. A new species of Misagria with a redescription of the genus (Odonata: Libellulidae). Entomological News, 62 (2), 70–76.

    Geijskes, D.C. (1959) The Aeschnine genus Staurophlebia. Notes on Odonata of Suriname VII. Studies on the Fauna of Suriname and other Guyanas, 3, 147–172.

    Geijskes, D.C. (1968) Anax longipes vs Anax concolor. Notes on Odonata of Suriname X. Studies on the Fauna of Suriname and other Guyanas, 10, 67–100.

    Geijskes, D.C. (1970) Generic characters of the south american Corduliidae, with descriptions of the species found in the Guyanas. Studies on the Fauna of Suriname and other Guyanas, 12 (1), 1–42.

    Geijskes, D.C. (1971) List of Odonata known from French Guyana, mainly based on a collection brought together by the mission of the “Museum National d’Histoire Naturelle”. Annales de la Société entomologique de France, New Series, 7 (3), 655–677.

    Geijskes, D.C. (1976) The Genus Oxystigma Selys, 1862 (Zygoptera: Megapodagrionidae). Odonatologica, 5 (3), 213–230.

    Gloger, H. (1962) Clave para la determinación de los géneros de Libellulidae (Odonata) de América del Sud. Neotropica, 8, 5–13.

    Gloger, H. (1964) Bemerkungen über die Odonaten-fauna der Galapagos-Inseln nach der Ausbeute von Juan Förster, 1959. Opuscula zoologica, 74, 1–6.

    Gloger, H. (1967) Sobre Acanthagrion (Odonata, Coenagrionidae) y géneros proximos. Acta Zoológica Lilloana, 21, 45–58.

    Gloyd, L.K. (1939) Two new species of Protoneura from South America. Occasional Papers of the Museum of Zoology, University of Michigan, 408, 1–21.

    Gloyd, L.K. (1980) The taxonomic status of the genera Superlestes and Cyptolestes Williamson, 1921 (Odonata: Lestidae). Occasional Papers of the Museum of Zoology, University of Michigan, 694, 1–3.

    González-Soriano, E., Delgado-Hernández, O. & Harp, G.L. (2004) Biological notes on Neoerythromma gladiolatum Williamson & Williamson, 1930, with description of its female. (Zygoptera: Coenagrionidae). Odonatologica, 33 (3), 327–331.

    Guerrero, G. (2015) La Manga del Cura decide pertenecer a Manabí, informó el CNE. In: El Universo. Available from: http://www.eluniverso.com/noticias/2015/09/27/nota/5153639/manga-cura-le-pertenece-manabi-dice-cne (accessed 16 March 2016)

    Guillermo-Ferreira, R & Bispo, P.C. (2013) Description of the larva of Telebasis griffinii (Martin, 1896) (Zygoptera: Coenagrionidae). Odonatologica, 42 (4), 403–407.

    Haber, B. & Wagner, D. (2014) Dragonflies and Damselflies of Ecuador (Odonata). Dragonflies of Ecuador. Available from: http://bdei2.cs.umb.edu/~whaber/Odonata_of_Ecuador/index.html (accessed 4 August 2016)

    Hagen, H.A. (1861) Synopsis of the Neuroptera of North America, with a list of the South American species. Smithsonian Miscellaneous Collections, Washington. Smithsonian Institution, Washington, 347 pp.

    https://doi.org/10.5962/bhl.title.60275

    Hagen, H.A. (1868) Odonaten Cubas (Fortsetzung). Entomologische Zeitung Stettin, 29, 274–287.

    Hagen, H.A. (1877) Synopsis of the Odonata of America. Proceedings of the Boston Society of Natural History, 18, 20–96.

    Hagen, H.A. & Calvert, P.P. (1902) Illustrations of Odonata, with a list and bibliography of the species. Bulletin of the Museum of Comparative Zoology at Harvard College, 34 (4), 105–120.

    Hanratty, D.M. (Ed.) (1989) A Country Study: Ecuador. Library of Congress, Federal Research Division, Washington D.C., 306 pp.

    Haseman, J.F. (1912) Some factors of geographical distribution in South America. Annals of the New York Academy of Sciences, 22, 9–112.

    https://doi.org/10.1111/j.1749-6632.1912.tb55158.x

    Hawking, J.H. & Ingram, B.A. (1994) Rate of larval development of Pantala flavescens (Fabricius) at its southern limit of range in Australia (Anisoptera: Libellulidae). Odonatologica, 23 (1), 63–68.

    Heckman, C.W. (2006) Encyclopedia of South American Aquatic Insects: Odonata—Anisoptera. Illustrated Keys to Known Families, Genera and Species in South America. Springer, A. A. Dordrecht, 723 pp.

    Heckman, C.W. (2008) Encyclopedia of South American Aquatic Insects: Odonata – Zygoptera. Illustrated Keys to Known Families, Genera and Species in South America. Springer, A. A. Dordrecht, 687 pp.

    https://doi.org/10.1007/978-1-4020-8176-7

    Hedström, I. & Sahlén, G. (2001) A key to the adult Costa Rican "helicopter" damselflies (Odonata: Pseudostigmatidae) with notes on their phenology and life zone preferences. Revista de biologia tropical, 49 (3–4), 1037–56.

    Hincks, W.D. (1934) Odonata (Paraneuroptera) from Peru and Colombia. I. Aeshnidae. Entomologist’s Record and Journal of Variation, 46 (7–8), 77–81.

    Hoffmann, J. (2009) Summary catalogue of the Odonata of Peru. IDF-Report, 16 (2009), 1–115.

    Hoffmann, J. (2010) Do climate changes influence dispersal and population dynamics of dragonflies in the western Peruvian Andes? BioRick, 5, 47–71.

    https://doi.org/10.3897/biorisk.5.842

    Howard, L.O. (1889) Scientific Results of explorations by the U. S. Fish Commission steamer Albatros. V. Annotated catalogue of the insects collected in 1878–88. Proceedings of the United States National Museum, 12 (71), 194.

    Hunger, H. & Schiel, F. (2012) Description of Protallagma hoffmanni sp. nov. from the Peruvian Andes (Odonata: Coenagrionidae), including description of the larva. Zootaxa, 3202 (1), 28–50.

    https://doi.org/10.11646/zootaxa.3202.1.2

    Ingley, S.J., Bybee, S.M., Tennessen, K.J., Whitting, M.F. & Branham, M.A. (2012). Life on the fly: phylogenetics and evolution of the helicopter damselflies (Odonata, Pseudostigmatidae). Zoologica Scripta, 41, 637–650.

    https://doi.org/10.1111/j.1463-6409.2012.00555.x

    INEC (2017) El Instituto Nacional de Estadística y Censos del Ecuador. Available from: http://www.ecuadorencifras.gob.ec (accessed 15 February 2017)

    IUCN (2018) The IUCN Red List of Threatened Species. Version 2018-1. Available from: http://www.iucnredlist.org (accessed 16 August 2018)

    Johnson, J.T. (2016) Leptobasis linda sp. nov. from Ecuador (Odonata: Coenagrionidae). Zootaxa, 4171 (2), 373–381.

    https://doi.org/10.11646/zootaxa.4171.2.10

    Jørgensen, P.M. & León-Yánez, S. (1999) Catalogue of the vascular plants of Ecuador. Monographs in Systematic Botany, Missouri Botanical Garden, 75, 1–1182.

    Jurzitza, G. (1982) Die Unterscheidung der Mannchen von Zenithoptera fasciata (Linnaeus, 1758), Z. viola Ris,1910, und Z. lanei Santos, 1941 (Anisoptera: Libellulidae). Odonatologica, 11 (4), 331–338.

    Kalkman, V.J., Clausnitzer, V., Dijkstra, K.B., Orr, A.G., Paulson, D.R. & Tol, V.T. (2008) Global diversity of dragonflies (Odonata) in freshwater. Hydrobiologia, 595, 351–363.

    https://doi.org/10.1007/s10750-007-9029-x

    Karsch, F. (1890) Beiträge zur Kenntniss der Arten und Gattungen der Libellulinen. Berliner Entomologische Zeitschrift, 33 (2), 347–392.

    https://doi.org/10.1002/mmnd.18900330221

    Karsch, F. (1891a) Neue Odonaten von Ecuador. Societas Entomologica, 6 (14), 105.

    Karsch, F. (1891b) Neue Odonaten von Ecuador. Societas Entomologica, 6 (15), 113.

    Karsch, F. (1891c) Neue Odonaten von Ecuador. Societas Entomologica, 6 (16), 121–122.

    Kemp, C. (2017) The Lost Species: Great Expeditions in the Collections of Natural History Museums. University of Chicago Press, Chicago, Illinois, 256 pp.

    https://doi.org/10.7208/chicago/9780226386355.001.0001

    Kennedy, C.H. (1919) The Phylogeny of the Zygoptera, a Study of the Phylogeny of the Zygoptera from the Evidence Given by the Genitalia. Ph.D. thesis, Cornell University, 293 pp.

    Kennedy, C.H. (1920) Forty–two hitherto unrecognized genera and subgenera of Zygoptera, Ohio Journal of Science, 21, 83–88.

    https://doi.org/10.5962/bhl.part.14540

    Kennedy, C.H. (1922) The phylogeny and geographical distribution of the genus Libellula. Entomological News, 33 (3), 65–71, 105–111.

    Kennedy, C.H. (1923) The phylogeny and the distribution of the genus Erythemis (Odonata). Miscellaneous Publications, University of Michigan, Museum of Zoology, 11, 19–21.

    Kennedy, C.H. (1936) Telebasis flammeola, T. carota and T. livida, new dragonflies from Ecuador (Odonata: Coenagrionidae). Annals of the Entomological Society of America, 29 (4), 804–815.

    https://doi.org/10.1093/aesa/29.4.804

    Kennedy, C.H. (1937a) Staurophlebia auca, a new dragonfly from Ecuador (Odonata: Aeshnidae). Annals of the Entomological Society of America, 30 (2), 425–430.

    https://doi.org/10.1093/aesa/30.2.425

    Kennedy, C.H. (1937b) Perilestes castor and P. pollux, new dragonflies from Ecuador, with notes on P. kahli. Annals of the Entomological Society of America, 30 (2), 410–424.

    https://doi.org/10.1093/aesa/30.2.410

    Kennedy, C.H. (1938a) Palaemnema picicaudata, P. abbreviata and P. brucelli, new dragonflies from Ecuador, with notes on other Palaemnema (Odonata: Platystictidae). Annals of the Entomological Society of America, 31 (2), 249–266.

    https://doi.org/10.1093/aesa/31.2.249

    Kennedy, C.H. (1938b) Aeshna biliosa, A new dragonfly from Andean Equador and Peru (Odonata: Aeshnidae). Annals of the Entomological Society of America, 31 (4), 573–577.

    https://doi.org/10.1093/aesa/31.4.573

    Kennedy, C.H. (1939a) Archaepodagrion bicorne, a very primitive dragonfly from Eastern Ecuador (Odonata: Megapodagrionidae). Annals of the Entomological Society of America, 33 (1), 32–43.

    https://doi.org/10.1093/aesa/32.1.32

    Kennedy, C.H. (1939b) Protallagma runtuni, n. sp. of dragonfly from Ecuador with notes on the genus (Coenagriidae: Odonata). Annals of the Entomological Society of America, 32 (1), 177–187.

    https://doi.org/10.1093/aesa/32.1.177

    Kennedy, C.H. (1939c) Protoneura macintyrei, a new species of dragonfly from Ecuador (Odonata: Protoneurinae). Occasional Papers of the Museum of Zoology, University of Michigan, 410, 1–8, 2 pl.

    Kennedy, C.H. (1939d) The female of Aeshna biliosa (Odonata: Aeshnidae). Annals of the Entomological Society of America, 32 (2), 344–348.

    https://doi.org/10.1093/aesa/32.2.344

    Kennedy, C.H. (1940) The Miocora-like dragonflies from Ecuador with notes on Cora, Miocora, Kalocora, Josocora and Stenocora (Odonata: Polythorinae). Annals of the Entomological Society of America, 33 (2), 406–436.

    https://doi.org/10.1093/aesa/33.2.406

    Kennedy, C.H. (1941a) Perilestinae in Ecuador and Peru: Revisional notes and descriptions (Lestidae: Odonata). Annals of the Entomological Society of America, 34 (3), 658–688.

    ttps://doi.org/10.1093/aesa/34.3.658

    Kennedy, C.H. (1941b) Agnophilogenia monotis, new genus and species of dragonfly from humid Northwestern Ecuador (Odonata: Megapodagrionidae). Annals of the Entomological Society of America, 34 (2), 490–494.

    https://doi.org/10.1093/aesa/34.2.490

    Kennedy, C.H. (1942a) Palaemnema lorena and P. melanocauda new species of dragonflies from northwestern Ecuador (Platystictidae: Odonata). Annals of the Entomological Society of America, 35 (1), 97–104.

    https://doi.org/10.1093/aesa/35.1.97

    Kennedy, C.H. (1942b) Lestes henshawi, Ecuador, L. urubamba, Peru, and notes on other South American Lestes (Lestidae: Odonata). Revista de Entomologia, 13 (3), 275–279.

    Kennedy, C.H. (1946a) Archaeopodagrion bilobata, n. sp., from central Ecuador (Odonata: Megapodagrionidae). Annals of the Entomological Society of America, 39 (2), 171–176.

    https://doi.org/10.1093/aesa/39.2.171

    Kennedy, C.H. (1946b) Protallagma runtuni Kennedy, 1939, a synonym of Oxyallagma dissidens Selys: notes on Oxyagrion and related genera. Annals of the Entomological Society of America, 39 (3), 381–382.

    https://doi.org/10.1093/aesa/39.3.381

    Kimmins, D.E. (1945) A new species of Odonata from Ecuador. Annals Magazine Natural History, 11 (12), 187–189.

    https://doi.org/10.1080/00222934508527503

    Kimmins, D.E. (1969a) A list of the type specimens of Libellulidae and Corduliidae (Odonata) in the British Museum (Natural History). Bulletin of the British Museum (Natural History). Entomology, 22 (6), 277–306.

    https://doi.org/10.5962/bhl.part.9954

    Kimmins, D.E. (1969b) A list of the type-specimens of Odonata in the British Museum (Natural History), Part II. Bulletin of the British Museum (Natural History), Entomology, 23 (7), 289–314.

    https://doi.org/10.5962/bhl.part.15135

    Kimmins, D.E. (1970) A list of the type-specimens of Odonata in the British Museum (Natural History), Part III. Bulletin of the British Museum (Natural History), Entomology, 24 (6), 173–205.

    https://doi.org/10.5962/bhl.part.1521

    Kirby, W.F. (1889) A revision of the subfamily Libellulinae, with descriptions of new genera and species. Transactions of the Zoological Society of London, 12 (9), 249–348.

    https://doi.org/10.1111/j.1096-3642.1889.tb00016.x

    Kirby, W.F. (1890) A Synonymic Catalogue of Neuroptera Odonata, or Dragonflies, with an Appendix of Fossil Species. Guerney & Jackson, London, 202 pp.

    https://doi.org/10.5962/bhl.title.5534

    Kirby, W.F. (1897) List of the Neuroptera collected by Mr. E.E. Austen on the Amazons etc. during the recent expedition of Messrs. Siemsn Bros. Cable S. S. 'Faraday,' with descriptions of several new species of Odonata (dragonflies). Annals and Magazine of Natural History, Series 6, 19 (114), 598–617.

    https://doi.org/10.1080/00222939708680582

    Kompier, T. (2015) A Guide to the Dragonflies and Damselflies of the Serra dos Orgaos, South-eastern Brazil. REGUA Publications, Brussels, 379 pp.

    Kormondy, E.J. (1958) The plesiallotype female of Oligoclada umbricola Borror, 1931. Entomological News, 69, 57–59.

    Law, G. (2015) Hierarchical administrative subdivision codes—Ecuador. Available from: http://www.statoids.com/uec.html (accessed 14 March 2016)

    Lencioni, F.A.A. (1999) The genus Phasmoneura, with description of Forcepsioneura gen. nov. and two new species (Zygoptera: Protoneuridae). Odonatologica, 28 (2), 127–137.

    Lencioni, F.A.A. (2005) The damselflies of Brazil—An illustrated identification guide. 1. The non-Coenagrionidae families. Print Editora, Sao Paulo, 324 pp.

    Lencioni, F.A.A. (2006) The damselflies of Brazil—An illustrated identification guide. 2-Coenagrionidae. Print Editora, Sao Paulo, 419 pp.

    Lencioni, F.A.A. (2014) Diagnoses and discussion of the group 1 and 2 Brazilian species of Heteragrion, with descriptions of four new species (Odonata: Megapodagrionidae). Zootaxa, 3685 (1), 1–80.

    https://doi.org/10.11646/zootaxa.3685.1.1

    Leonard, J.W. (1977) A revisionary study of the genus Acanthagrion (Odonata: Zygoptera). Miscellaneous Publications, University of Michigan, Museum of Zoology, 153, 1–173.

    Loaiza, M. T., & D. Anjos-Santos (2019) Una catástrofe para la ciencia y la historia de Latinoamérica. Hetaerina, Boletín de la Sociedad de Odonatología Latinoamericana, 1 (1), 9–10.

    Lozano, F. (2007) Filogenia del género Acanthagrion Selys (Odonata, Coenagrionidae) y revisión sistemática del grupo viridescens, Resolución: 186/2006. N de expediente: 04496/2006. Thesis, Facultad de Ciencias, Naturales y Museo, Universidad Nacional De La Plata. Available from: http://naturalis.fcnym.unlp.edu.ar/id/20120126001064 (accessed 14 May 2019)

    Lozano, F. (2013) Description of three females of the genus Acanthagrion (Odonata: Coenagrionidae) with a key to the females from Argentina. Zootaxa, 3646 (1), 23–38.

    https://doi.org/10.11646/zootaxa.3646.1.2

    Lozano, F., Garré, A. & Pessacq, P. (2007) Descripción del último estadio larval de Acanthagrion aepiolum (Odonata: Coenagrionidae). Revista de la Sociedad Entomológica Argentina, 66, 1–4.

    Lozano, F., Muzón, J. & Scattolini, C. (2012) Description of the final stadium larva of Telebasis obsoleta (Selys, 1876) (Odonata: Coenagrionidae). Zootaxa, 3186 (1), 54–58.

    https://doi.org/10.11646/zootaxa.3186.1.4

    Lozano, F., Rodriguez, J. & Molineri, C. (2017) Acanthagrion peruvianum Leonard, 1977 (Odonata Coenagrionidae) a junior subjective synonym of A. floridense Fraser, 1946 and description of its final stadium larva. Zoologischer Anzeiger, 270, 71–80.

    https://doi.org/10.1016/j.jcz.2017.10.002

    Machado, A.B.M. (1954) Elga santosi sp. n. e redescrição de Elga leptostyla Ris, 1911 (Odonata, Libellulidae). Revista Brasileira Biologia, 14 (3), 303–312.

    Machado, A.B.M. (1980) Helveciagrion n. g., com descrição de uma nova especie de Parque Estadual do Río Doce, Minas Gerais (Odonata-Coenagrionidae). Lundiana, 1, 59–87.

    Machado, A.B.M. (1992) A taxonomic note on Elga Ris, with E. newtonsantosi nom. nov. for E. leptostyla Machado, 1954 (Anisoptera: Libellulidae). Notulae Odonatologicae, 3 (9), 153–154.

    Machado, A.B.M. (2000) Studies on neotropical Protoneuridae. 10. Forsepsioneura lucia sp. n. from the Parque Estadual rola Moca, Minas Gerais, Brazil (Odonata, Zygoptera). Boletim do Museu de Biologia Mello Leitão, New Series, 11–12, 127–134.

    Machado, A.B.M. (2001) Studies on Neotropical Protoneuridae. 11. Two new species of Forcepsioneura Lencioni (Odonata-Zygoptera) with a key to the males of the genus. Revista Brasileira de Zoologia, 18 (3), 845–854.

    https://doi.org/10.1590/S0101-81752001000300021

    Machado, A.B.M. (2004) Studies on neotropical Protoneuridae. 15. Amazoneura gen. nov. with a description of A. juruaensis sp. nov. (Odonata, Zygoptera). Revista Brasileira de Zoologia, 21 (2), 333–336.

    https://doi.org/10.1590/S0101-81752004000200029

    Machado, A.B.M. (2005) Peruviogomphus bellei spec. nov. from the Amazonian region of Brazil (Anisoptera: Gomphidae). Odonatologica, 34 (1), 59–63.

    Machado, A.B.M. (2009a) Denticulobasis and Tuberculobasis: new genera close to Leptobasis, with description of ten new species (Zygoptera: Coenagrionidae). Zootaxa, 2108, 1–36.

    Machado, A.B.M. (2009b) Studies on neotropical Protoneuridae. 21. The status of Amazoneura Machado, 2004 (Odonata: Protoneuridae). Lundiana, 9 (1), 53–56.

    https://doi.org/10.11646/zootaxa.2384.1.3

    Machado, A.B.M. (2010) Seven new species of Telebasis from Brazil (Odonata: Coenagrionidae). Zootaxa, 2384, 53–64.

    Machado, A.B.M. (2012a) The apicale species group of Acanthagrion, with descriptions of four new species and a hook-moving apparatus (Zygoptera: Coenagrionidae). Odonatologica, 41 (3), 201–223.

    Machado, A.B.M. (2012b) A new species of Ischnura (Odonata: Coenagrionidae) from high altitude eastern Andes of Colombia. Zoologia, 29 (6), 598–600.

    https://doi.org/10.1590/S1984-46702012000600011

    Machado, A.B.M. & Costa, J.M. (1995) Navicordulia gen. nov., a new genus of neotropical Corduliinae, with descriptions of seven new species (Anisoptera: Corduliidae). Odonatologica, 24 (2), 187–218.

    Machado, A.B.M. & Lacerda, D.S. (2017) Revalidation of Platystigma Kennedy, 1920, with a synopsis of the quadratum species group and the description of three new species (Odonata: Pseudostigmatidae). Zootaxa, 4242 (3), 493–516.

    https://doi.org/10.11646/zootaxa.4242.3.4

    Maria, A. (1938) Catalogo de los Odonatos Colombianos. Revista chilena de historia natural, 62, 206–211.

    Martin, R. (1896) Sur les Odonates recueillis par le D.r festa au Darien et á Cuenca. Bollettino dei Musei di Zoologia ed Anatomia comparata della R. Universita di Torino, 11 (240), 1–3.

    Martin, R. (1897) Description d’Odonates nouveaux. Ext. Annales de la Société entomologique de France, 66 (4), 589–594.

    Martin, R. (1907) Cordulines. Collections Zoologiques du Baron Edm. de Selys Longchamps, Catalogue Systématique et descriptif, 17, 1–94.

    Martin, R. (1908) Aeschnines. Collections Zoologiqu