Abstract
Four new crane fly species of subgenus Tipula (Vestiplex) Bezzi, 1924, T. (V.) borthi sp. nov. (Sichuan, Qinghai and Gansu provinces, erectiloba species group), T. (V.) maoxianensis sp. nov. (Sichuan province, virgatula species group), T. (V.) xinduqiaoensis sp. nov. and T. (V.) auriculatalobata sp. nov. (Sichuan and Yunnan provinces, divisotergata species group) are described and illustrated. The newly described species can be separated using the identification keys provided for local species of the erectiloba, virgatula and divisotergata groups.
References
Alexander, C.P. (1932) New or little-known Tipulidae from eastern Asia (Diptera). X. Philippine Journal of Science, 49, 105–136.
Alexander, C.P. (1933) New or little-known Tipulidae from eastern Asia (Diptera). XIV. Philippine Journal of Science, 51, 507–544.
Alexander, C.P. (1934) New or little-known Tipulidae from eastern Asia (Diptera). XXI. Philippine Journal of Science, 55, 19–60.
Alexander, C.P. (1935) New or little-known Tipulidae from eastern Asia (Diptera). XXV. Philippine Journal of Science, 57, 81–148.
Alexander, C.P. (1936) New or little-known Tipulidae from eastern Asia (Diptera). XXXI. Philippine Journal of Science, 60, 323–360.
Alexander, C.P. (1959) Undescribed species of crane-flies from the Himalaya mountains (Tipulidae, Diptera). IV. Journal of the New York Entomological Society, 67, 223–235.
Alexander, C.P. (1963) Some Tipulidae from Tibet and upper Burma in the British museum (Natural History) (Diptera). Bulletin of the British Museum (Natural History), Entomology, 14, 319–340.
https://doi.org/10.5962/bhl.part.8785
Alexander, C.P. (1965) New subgenera and species of crane-flies from California (Diptera: Tipulidae). Pacific Insects, 7, 333–386.
Alexander, C.P. & Byers, G.W. (1981) Tipulidae. In: McAlpine, J.F. (Coord.), Manual of Nearctic Diptera. Monograph 27 (1). Research Branch Agriculture Canada, Ottawa, pp. 153–190.
Bezzi, M. (1924) Una nuova Tipula delle Alpi con ali ridotte anche nel maschio (Dipt.). Annali del Museo Civico di Storia naturale di Genova, 51, 228–233.
Dia, A. & Theowald, Br. (1982) Tipula (Vestiplex) relicta nov. spec., ein frühpleistozänes Relikt aus Libanon (Diptera, Tipulidae). Bulletin Zoologisch Museum, Universiteit van Amsterdam, 9, 21–24.
Edwards, F.W. (1928) Some nematocerous Diptera from Yunnan and Tibet. Annals and Magazine of Natural History, Series 10, 1 (6), 681–703.
https://doi.org/10.1080/00222932808672840
Frommer, S.I. (1963) Gross morphological studies of the reproductive system in representative North American crane flies (Diptera: Tipulidae). The University of Kansas Science Bulletin, 44, 535–625.
Gelhaus, J.K. (2005) Systematics and biogeography of the desert crane fly subgenus Tipula (Eremotipula) Alexander (Diptera: Tipulidae). Memoirs of the American Entomological Society, 46, 1–235.
Hemmingsen, A.M. (1956) Deep-boring ovipository instincts of some crane-fly species (Tipulidae) of the subgenera Vestiplex Bezzi and Oreomyza Pok. and some associated phenomena. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, 118, 243–315.
Mannheims B (1953) 15. Tipulidae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region, 3 (5) 1, Lief. 173, pp. 113–136.
Mannheims, B. (1963) 15. Tipulidae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region, 3 (5) 1, Lief. 238, pp. 137–176.
McAlpine, J.F. (1981) Morphology and terminology adults. In: McAlpine, J.F. (Coordinator), Manual of Nearctic Diptera. Monograph 27. Research Branch, Agriculture Canada, Ottawa, pp. 9–63.
Oosterbroek, P. (2019) Catalogue of the Craneflies of the World (CCW). Available from: https://ccw.naturalis.nl/index.php
(accessed 5 August 2019)Savchenko, E.N. (1960) A contribution to the taxonomy of crane-flies (Diptera, Tipulidae) of the subgenus Vestiplex Bezzi of the genus Tipula L. Horae Societatis Entomologicae Rossicae, 47, 143–216 [in Russian]
Savchenko, E.N. (1964) Crane-flies (Diptera, Tipulidae), Subfam. Tipulinae, Genus Tipula L., 2. Fauna USSR, New Series, 89 (Insecta Diptera), 4, 1–503. [in Russian]
Starkevich, P. (2012) Taxonomic and phylogentic review of the subgenus Tipula (Vestiplex) Bezzi, 1924 (Diptera, Tipulidae). Doctoral dissertation, Vilnius University, Vilnius, 219 pp. [in Lithuanian]
Theowald, Br. & Mannheims, B. (1963) Die Arten der Tipula (Vestiplex) excisa-Gruppe in der Paläarktis (Diptera, Tipulidae). Bonner Zoologische Beitrage, 13, 360–402.