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Revalidation of Vareuptychia Forster, 1964, description of Vanima gen. nov., and notes on Euptychia cleophes Godman & Salvin, 1889 (Lepidoptera: Nymphalidae: Satyrinae)

Departamento de Biologia Animal and Museu da Biodiversidade, Instituto de Biologia, Laboratório de Ecologia e Sistemática de Borboletas, Universidade Estadual de Campinas, Campinas, São Paulo, Brazil.
Department of Zoology, Laboratório de Estudos de Lepidoptera Neotropical, Universidade Federal do Paraná, Curitiba, Paraná, Brazil.
Department of Zoology, Laboratório de Estudos de Lepidoptera Neotropical, Universidade Federal do Paraná, Curitiba, Paraná, Brazil.
Department of Life Sciences, Natural History Museum, London, UK.
Research Museum Alexander Koenig, Bonn, Germany.
Departamento de Biologia Animal and Museu da Biodiversidade, Instituto de Biologia, Laboratório de Ecologia e Sistemática de Borboletas, Universidade Estadual de Campinas, Campinas, São Paulo, Brazil.
McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida, USA.
McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida, USA.
Departament of Entomology, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima, Peru.
Lepidoptera Amazon rainforest Central America Cissia dry forests Euptychiina integrative taxonomy Megisto phylogeny


Vareuptychia Forster, 1964 stat. rest. is revalidated and comprises two species, V. similis (Butler, 1867) comb. rest. and V. themis (Butler, 1867) comb. nov. Vanima Zacca, Casagrande & Mielke gen. nov. is described to contain Euptychia labe Butler, 1870 (the type species), E. palladia Butler, 1867 and E. lesbia Staudinger, [1886]. The taxonomy of these two genera was initially revised based on morphological and distributional data, and subsequently tested and supported with a Maximum Likelihood analysis using four genes (COI, GAPDH, RpS5 and EF1-a). Lectotypes are designated for Euptychia similis Butler, 1867, E. themis Butler, 1867, E. undina Butler, 1870 and E. lesbia Staudinger, [1886]. No DNA sequences were obtained for Euptychia cleophes Godman & Salvin, 1889 but its transfer to Megisto Hübner, [1819] is supported by morphological evidence. For all taxa treated in this study, a taxonomic catalog, diagnosis, (re)description and illustrations of adults, venation and genitalia are provided, as well as comments on intraspecific variation, sexual dichromatism, ecology and distribution maps.



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