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Type: Article
Published: 2021-01-28
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Higher classification of the Monostilifera (Nemertea: Hoplonemertea)

Faculty of Science, Hokkaido University, N10 W8, Sapporo 060-0810, Japan
Nemertea Enopla higher taxa marine invertebrate phylogeny systematics taxonomy

Abstract

The nemertean order Monostilifera consists of 594 species in 127 genera and is distributed worldwide. Within the Monostilifera, two suborders have been recognized, Cratenemertea and Eumonostilifera. Within the latter, two, unranked clade names, Oerstediina and Amphiporina, were recently proposed without formal taxonomic definition. In this article, I give morphological circumscriptions and clade definitions for Cratenemertea, Eumonostilifera, Oerstediina, Plectonemertidae, Oerstediidae, and Amphiporina. Oerstediina and Amphiporina are placed on the Linnaean rank of infraorder. Constituent genera and species for each higher taxon are tabulated. The genus Amphiporella Friedrich, 1939 is herein replaced with Germanemertes nom. nov. to avoid homonymy with the Carboniferous fossil bryozoan genus Amphiporella Girty, 1910. Loxorrhochmidae Diesing, 1862 is declared a nomen oblitum relative to Tetrastemmatidae Hubrecht, 1897, a nomen protectum under Article 23.9 of the International Code of Zoological Nomenclature. There remain 308 species of eumonostiliferans whose infraorder affiliation is uncertain due to the lack of information on vascular morphology and molecular sequence data. The suborder affiliation of the two species Cinclidonemertes mooreae Crandall, 2010 and Verrillianemertes schultzei Senz, 2001 is left uncertain.

 

References

  1. Abildgaard, P.C. (1806) Planaria dorsalis. In: Müller, O.F. (Author) & Abildgaard, P.C., Vahl, M., Holten, J.S. & Rathke, J. (who prepared descriptions and figures), Zoologia Danica sev animalium Daniae et Norvegiae rariorvm ac minus notorvm descriptiones et historia. Volvmen qvartvm. Vol. 4. Explicationi iconvm fascicvli qvarti ejvsdem operis inserviens. N. Christensen, Havniae (locative case of Havnia, Copenhagen), pp. 25.

    [https://www.biodiversitylibrary.org/item/104862#page/231/mode/1up]

    Akhmatova, A.F., Chernyshev, A.V. & Zaslavskaya, N.I. (2012) The species composition of the nemertean genus Oerstedia (Nemertea: Hoplonemertea) in the Far Eastern seas of Russia. Russian Journal of Marine Biology, 38, 423–430.

    https://doi.org/10.1134/S1063074012060028

    Andrade, S.C.S., Strand, M, Schwartz, M., Chen, H., Kajihara, H., Döhren, J.v., Sun, S., Junoy, J., Thiel, M., Norenburg, J.L., Turbeville, J. M., Giribet, G. & Sundberg, P. (2012) Disentangling ribbon worm relationships: multi-locus analysis supports traditional classification on the phylum Nemertea. Cladistics, 28, 141–159.

    https://doi.org/10.1111/j.1096-0031.2011.00376.x

    Beaumont, W.I. (1900) Fauna and flora of Valencia Harbour, Ireland. XI.—Report on the Nemertea. Proceedings of the Royal Irish Academy, Series 3, 5, 815–831.

    Benham, W.B. (1901) The Platyhelmia, Mesozoa, and Nemertini. In: Lankester, E.R. (Ed.), A Treatise on Zoology. Adam & Charles Black, London, pp. 1–204.

    [https://archive.org/details/treatiseonzoolog04lankrich/page/204/mode/2up]

    Berg, G. (1973) On morphology and distribution of some hoplonemertean species along Scandinavian coasts (Nemertini). Zoologica Scripta, 2, 63–70.

    https://doi.org/10.1111/j.1463-6409.1974.tb00740.x

    Berg, G. (1976) Redescription of Zygonemertes algensis (Bürger, 1895) (Nemertini, Hoplonemertini) from the Mediterranean Sea. Pubblicazioni della Stazione Zoologica di Napoli, 40, 11–18.

    Berg, G. (1985) Annulonemertes gen. nov., a new segmented hoplonemertean. In: Conway Morris, S., George, J.D., Gibson, R. & Platt, H.M. (Eds.), The Origins and Relationships of Lower Invertebrates. Clarendon, Oxford, pp. 200–209.

    Bergendal, D. (1900) Über ein Paar sehr eigenthümliche nordische Nemertinen. Zoologischer Anzeiger, 23, 313–328.

    [https://www.biodiversitylibrary.org/item/97248#page/341/mode/1up]

    Bergendal, D. (1903) Till kännedomen om de nordiska Nemertinerna. 4. Förteckning öfver vid Sveriges vestkust iakttagna Nemertiner. Arkiv för Zoologi, 1, 85–156.

    [https://www.biodiversitylibrary.org/item/30516#page/103/mode/1up]

    Blainville, H.M. de (1827) Espèces molles, sans articulations distinctes. (Genre Malacobdella, de Bl.). In: Levrault, F.G. (Ed.), Dictionnaire des sciences naturelles. Tome quarante-septième. Vol. 47. Normant, Paris, pp. 270–271.

    [https://www.biodiversitylibrary.org/item/81586#page/276/mode/1up]

    Blanchard, E. (1847) Recherches sur l’organisation des vers. Annales des Sciences Naturelles, Zoologie, Série 3, 8, 119–149.

    [https://www.biodiversitylibrary.org/item/23613#page/125/mode/1up]

    https://doi.org/10.5962/bhl.title.51506

    Böhmig, L. (1893) Tetrastemma graecensis. In: Mojsisovics, A. von (Reporter), Bericht der II. Section, für Zoologie. Mittheilungen des naturwissenschaftlichen Vereines für Steiermark. Vol. 29. Naturwissenschaftlichen Vereine für Steiermark, Graz, pp. LXXXII–LXXXIX.

    [https://www.biodiversitylibrary.org/item/45856#page/64/mode/1up]

    Brinkmann, A. (1914–1915) Uniporus, ein neues Genus der Familie Drepanophoridae Verrill. Bergens Museums Aarbok, 6, 1–29.

    [https://www.biodiversitylibrary.org/item/130365#page/233/mode/1up]

    Brinkmann, A. (1917) Pelagic nemerteans from the “Michael Sars” North Atlantic Deep-Sea Expedition 1910. Report on the Scientific Results of the “Michael Sars” North Atlantic Deep-Sea Expedition 1910, 3 (2), 1–18.

    [https://www.biodiversitylibrary.org/item/120117#page/19/mode/1up]

    Brinkmann, A. Jr. (1943) A new land nemertean. Nature, 151, 253.

    https://doi.org/10.1038/151253a0

    Bürger, O. (1890) Untersuchungen über die Anatomie und Histologie der Nemertinen nebst Beiträgen zur Systematik. Zeitschrift für wissenschaftliche Zoolgie, 50, 1–277.

    [https://www.biodiversitylibrary.org/item/135257#page/11/mode/1up]

    Bürger, O. (1893) Südgeorgische und andere exotische Nemertinen. Zoologische Jahrbücher, Abtheilung für Systematik, Geographie und Biologie der Thiere, 7, 207–240.

    [https://www.biodiversitylibrary.org/item/121145#page/221/mode/1up]

    https://doi.org/10.5962/bhl.part.24065

    Bürger, O. (1895) Die Nemertinen des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. Fauna und Flora des Golfes von Neapel und der angrenzenden Meeres-Abschnitte herusgegeben von der Zoologischen Station zu Neapel. 22. Monographie. R. Friedländer & Sohn, Berlin, xvi + 743 pp., 31 pls.

    [https://www.biodiversitylibrary.org/item/49221#page/8/mode/1up]

    https://doi.org/10.5962/bhl.title.14030

    Bürger, O. (1897–1907) Nemertini (Schnurwürmer). Dr. H.G. Bronn’s Klassen und Ordnungen des Tier-Reichs, wissenschaftlich dargestellt in Wort und Bild. Vierter Band [Volume 4]. Supplement. C.F. Winder’sche Verlagshandlung, Leipzig, viii + 542 pp. [pp. 1–176 published in 1897; pp. 177–240 in 1898; pp. 241–288 in 1899; pp. 289–384 in 1903; pp. 385–480 in 1905; pp. 481–542 in 1907, https://www.biodiversitylibrary.org/item/40724#page/7/mode/1up

    Bürger, O. (1904a) Nemertini. Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. 20. Lieferung. Platyhelminthes. R. Friedländer und Sohn, Berlin, xviii + 151 pp.

    [https://www.biodiversitylibrary.org/item/15763#page/7/mode/1up]

    Bürger, O. (1904b) Nemertinen. Résultats du voyage du S. Y. Belgica en 1897–1898–1899 sous le commandement de A. de Gerlache de Gomery. Rapports scientifiques publié aux frais du Gouvernement Belge, sous la direction de la Commission de la Belgica. Zoologie. J.-E. Buschmann, Antwerp, 10 pp.

    Bürger, O. (1909) Die Nemertinen. Deutsche Tiefsee Expedition 1898–1899, 16, 170–221.

    Carus, J.V. (1885) Prodromus faunae Mediterraneae sive descriptio animalium Maris Mediterranei incolarum quam comparata silva rerum quatenus innotuit adiectis locis et nominibus vulgaribus eorumque auctoribus. Vol. I. Coelenterata, Echinodermata, Vermses, Arthropoda. E. Schweizerbart’sche Verlagshandlung (E. Koch), Stuttgart, xi + 524 pp.

    [https://www.biodiversitylibrary.org/item/45304#page/181/mode/1up]

    https://doi.org/10.5962/bhl.title.11523

    Chernyshev, A.V. (1991a) Sacconemertopsis belogurovi sp. nov. (Hoplonemertini, Tetrastemmatidae)—novaya solonovatovodnaya nemertina iz Yuzhnogo Primorya. Biologicheskie Nauki, 5, 45–49.

    Chernyshev, A.V. (1991b) Tortus tokmakovae sp. n. (Hoplonemertini; Emplectonematidae) from the Sea of Japan. Zoologischeskii Zhurnal, 70, 129–132.

    Chernyshev, A.V. (1992) On the names of some nemertines. Zoologischeskii Zhurnal, 71, 134–136.

    Chernyshev, A.V. (1993) A review of the genera of nemerteans allied to Oerstedia (Monostilifera, Tetrastemmatidae) with description of four new species. Zoologischeskii Zhurnal, 72, 11–20.

    Chernyshev, A.V. (1998a) New monostiliferous nemerteans (Enopla, Monostilifera) from the Sea of Japan. Zoologicheskii Zhurnal, 77, 397–403.

    Chernyshev, A.V. (1998b) Nemerteans of the genus Tetrastemma (Enopla, Monostilifera) from the Far East seas of Russia. Zoologicheskii Zhurnal, 77, 995–1002.

    Chernyshev, A.V. (1998c) On the genus Gurjanovella Uschakov, 1926 (Nemertea: Enopla). Zoosystematica Rossica, 7, 9–11.

    Chernyshev, A.V. (1999) Nemerteans of the genus Antarctonemertes (Enopla, Monostilifera). Zoologicheskii Zhurnal, 78, 939–948.

    Chernyshev, A.V. (2002) Description of a new species of the genus Poseidonemertes (Nemertea, Monostilifera) with establishment of the family Poseidonemertidae. Zoologicheskii Zhurnal, 81, 909–916.

    Chernyshev, A.V. (2003a) Classification system of the higher taxa of enoplan nemerteans (Nemertea, Enopla). Russian Journal of Marine Biology, 29 (Supplement 1), S57–S65.

    https://doi.org/10.1023/B:RUMB.0000011717.06390.30

    Chernyshev, A.V. (2003b) A new species of Tetrastemma from the Sea of Japan and redescription of Tetrastemma laminariae Uschakov, 1928 (Nemertea: Tetrastemmatidae). Zoosystematica Rossica, 12, 19–22.

    Chernyshev, A.V. (2004) Two new genera of nemertean worms of the family Tetrastemmatidae (Nemertea: Monostilifera). Zoosystematica Rossica, 12, 151–156.

    Chernyshev, A.V. (2005a) System of families of enoplan nemerteans of the order Eumonostilifera (Nemertea: Enopla). Russian Journal of Marine Biology, 31 (Supplement 1), S27–S33.

    https://doi.org/10.1007/s11179-006-0012-6

    Chernyshev, A.V. (2005b) Redescription of Korotkevitschia pelagica (Korotkevitsch, 1961) (Enopla: Hoplonemertea: Cratenemertea), a pelagic nemertean from Antarctica. Zootaxa, 862 (1), 1–14.

    https://doi.org/10.11646/zootaxa.862.1.1

    Chernyshev, A.V. (2011a) Comparative Morphology, Systematics and Phylogeny of the Nemerteans. Dalnauka, Vladivostok, 309 pp. [in Russian]

    Chernyshev, A.V. (2011b) Nemertean worms (Nemertea) of the Vietnamese coastal waters. In: Lutaenko, K.A. (Ed.), Proceedings of the Workshop Coastal Marine Biodiversity and Bioresources of Vietnam and Adjacent Areas to the South China Sea Nha Trang, Vietnam. 24–25 November 2011. Dalnauka, Vladivostok, pp. 21–25.

    Chernyshev, A.V. (2016) Nemerteans of the coastal waters of Vietnam. In: Adrianov, A.V. & Lutaenko, K.A. (Eds.), Biodiversity of the Western Part of the South China Sea. Dalnauka, Vladivostok, pp. 279–314.

    Chernyshev, A.V. & Polyakova, N.E. (2018a) Nemerteans of the Vema-TRANSIT expedition: First data on diversity with description of two new genera and species. Deep-Sea Research, Part II, 148, 64–73.

    https://doi.org/10.1016/j.dsr2.2017.06.004

    Chernyshev, A.V. & Polyakova, N.E. (2018b) Nemerteans from deep-sea expedition SokhoBio with description of Uniporus alisae sp. nov. (Hoplonemertea: Reptantia s.l.) from the Sea of Okhotsk. Deep-Sea Research, Part II, 154, 121–139.

    https://doi.org/10.1016/j.dsr2.2017.09.022

    Chernyshev, A.V. & Polyakova, N.E. (2019) Nemerteans from the deep-sea expedition KuramBio II with descriptions of three new hoplonemerteans from the Kuril–Kamchatka Trench. Progress in Oceanography, 178, 102148.

    https://doi.org/10.1016/j.pocean.2019.102148

    Chernyshev, A.V., Timoshkin, O. & Kawakatsu, M. (1998) Prostoma ohmiense sp. n., a new species of freshwater nemertean from lake Biwa-ko, central Japan, with special reference to the taxonomy and distribution of the genus Prostoma Dugès, 1828. Bulletin of Fuji Women’s College, Series II, 36, 51–66.

    Coe, W.R. (1901) Papers from the Harriman Alaska Expedition. XX. The nemerteans. Proceedings of the Washington Academy of Sciences, 3, 1–110.

    [https://www.biodiversitylibrary.org/item/81277#page/17/mode/1up]

    Coe, W.R. (1902) The nemertean parasites of crabs. American Naturalist, 36, 431–450.

    [https://www.biodiversitylibrary.org/item/135794#page/458/mode/1up]

    https://doi.org/10.1086/278150

    Coe, W.R. (1904) Nemerteans of the Pacific coast of North America. Part II. Harriman Alaska Series, 11, 111–220.

    [https://www.biodiversitylibrary.org/item/16998#page/143/mode/1up]

    Coe, W.R. (1905) Nemerteans of the west and northwest coasts of America. Bulletin of the Museum of Comparative Zoölogy at Harvard College, 47, 1–318.

    [https://www.biodiversitylibrary.org/item/21167#page/5/mode/1up]

    Coe, W.R. (1940) Revision of the nemertean fauna of the Pacific coasts of North, Central, and northern South America. Allan Hancock Pacific Expeditions, 2, 247–322.

    Coe, W.R. (1943) Biology of the nemerteans of the Atlantic coast of North America. Transactions of the Connecticut Academy of Arts and Sciences, 35, 129–328.

    Coe, W.R. (1944) A new species of hoplonemertean (Paranemertes biocellatus) from the Gulf of Mexico. Journal of the Washington Academy of Sciences, 34, 407–409.

    Corrêa, D.D. (1948) Ototyphlonemertes from the Brazilian coast. Comunicaciones zoológicas del Museo de historía natural de Montevideo, 2, 1–12.

    Corrêa, D.D. (1954) Nemertinos do litoral Brasileiro. Boletim da Faculdade de Filosofia, Ciências e Letras, Universidade de São Paulo, 19, 1–90.

    https://doi.org/10.11606/issn.2526-3382.bffclzoologia.1954.120084

    Corrêa, D.D. (1955) Os gêneros Emplectonema Stimpson e Nemertopsis Bürger (Hoplonemertini Monostilifera). Boletim da Faculdade de Filosofia, Ciências e Letras, Universidade de São Paulo, 20, 67–78.

    https://doi.org/10.11606/issn.2526-3382.bffclzoologia.1955.120212

    Corrêa, D.D. (1961) Nemerteans from Florida and Virgin Islands. Bulletin of Marine Science of the Gulf and Caribbean, 11, 1–44.

    Crandall, F.B. (1993a) Major characters and enoplan systematics. Hydrobiologia, 266, 115–140.

    https://doi.org/10.1007/BF00013363

    Crandall, F.B. (1993b) The genus Valdivianemertes Stiasny-Wijnhoff 1923 (Nemertea, Enopla, Hoplonemertea): nomenclatural status and proper systematic position. Hydrobiologia, 266, 175–184.

    https://doi.org/10.1007/BF00013367

    Crandall, F.B. (1994) Case 2888. Valdivianemertes Stiasny-Wijnhoff, 1923 (Nemertea): proposed conservation. Bulletin of Zoological Nomenclature, 51, 298–301.

    [https://www.biodiversitylibrary.org/page/34357801#page/75/mode/1up]

    https://doi.org/10.5962/bhl.part.7228

    Crandall, F.B. (2001) A cladistic view of the Monostilifera (Hoplonemertea) with interwoven rhynchocoel musculature: a preliminary assessment. Hydrobiologia, 456, 87–110.

    https://doi.org/10.1023/A:1013031900458

    Crandall, F.B. (2010) A new family of the Hoplonemertea: Cinclidonemertes mooreae fam., gen. et sp. nov. (Nemertea: Hoplonemertea). Journal of Natural History, 44, 2411–2424.

    https://doi.org/10.1080/00222933.2010.508567

    Crandall, F.B. & Gibson, R. (1998) A second genus of pelagic Cratenemertidae (Nemertea, Hoplonemertea). Hydrobiologia, 365, 173–198.

    https://doi.org/10.1023/A:1003107332635

    Cuvier, [G.] (1817) Le règne animal distribué d’après son organisation, pour servir de base a l’histoire naturelle des animaux et d’introduction a l’anatomie comparée. Tome IV [Volume 4], contenant les Zoophytes, les tables, et les planches. A. Belin, Paris, vii + 255 pp.

    [https://www.biodiversitylibrary.org/item/91495#page/11/mode/1up]

    Dakin, W.J. (1915) Fauna of West Australia—III. A new nemertean, Geonemertes dendyi, sp. n., being the first recorded land nemertean from Western Australia. Proceedings of the General Meetings for Scientific Business of the Zoological Society of London, 1915, 567–570.

    [https://www.biodiversitylibrary.org/item/98466#page/341/mode/1up]

    https://doi.org/10.1111/j.1469-7998.1915.00567.x

    Davison, P.G., Robison, H.W. & McAllister, C.T. (2014) First record of ribbon worms (Nemertea: Tetrastemmatidae: Prostoma) from Arkansas. Journal of the Arkansas Academy of Science, 68, 146–148.

    http://scholarworks.uark.edu/jaas/vol68/iss1/25

    De Queiroz, K. (2013) Nodes, branches, and phylogenetic definitions. Systematic Biology, 62, 625–632.

    https://doi.org/10.1093/sysbio/syt027

    Delle Chiaje, S. (1841) Descrizione e notomia degli animali invertebrati della Sicilia citeriore osservati vivi negli anni 1822–1830. Tomo Terzo. Molluschi acefali, bracciopedi, cirropedi; Crostacei, anellosi. C. Batelli, Naples, 142 pp.

    [https://www.biodiversitylibrary.org/item/40311#page/273/mode/1up]

    https://doi.org/10.5962/bhl.title.10031

    Dendy, A. (1892) On an Australian land nemertine (Geonemertes australiensis, n. sp.). Proceedings of the Royal Society of Victoria, New Series, 4, 85–122.

    [https://www.biodiversitylibrary.org/item/34240#page/119/mode/1up]

    Dendy, A. (1895 [imprint 1894]) Notes on a New Zealand land nemertine. Transactions and Proceedings of the New Zealand Institute, 27, 191–194.

    [https://www.biodiversitylibrary.org/item/22767#page/219/mode/1up]

    Diesing, K.M. (1862) Revision der Turbellarien. Abtheilung: Rhabdocoelen. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe, 45, 191–320.

    [https://www.biodiversitylibrary.org/item/30219#page/217/mode/1up]

    Diesing, K.M. (1863) Nachträge zur Revision der Turbellarien. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe, 46, 173–188.

    [https://www.biodiversitylibrary.org/item/30211#page/215/mode/1up]

    Dugès, A. (1828) Recherches sur l’organisation et les mœurs des Planariées. Annales des Sciences Naturelles, 15, 139–183, pls. IV–V.

    [https://www.biodiversitylibrary.org/item/29174#page/145/mode/1up]

    https://doi.org/10.5962/bhl.part.18554

    Ehrenberg, C.G. (1828–1831) Phytozoa Turbellaria Africana et Asiatica in Phytozoorum Tabula IV. et V. delineata. In: Hemprich, F.G. & Ehrenberg, C.G. (Eds.), Symbolae physicae, seu icones et descriptiones corporum naturalium novorum aut minus cognitorum quae ex itineribus per Libyam, Aegyptium, Nubiam, Dongalam, Syriam, Arabiam et Habessiniam, pars Zoologica II, animalia evertebrata exclusis insectis. Officina Academica: Berlin, pp. 53–67 (without pagination), pls. IV–V (plates published in 1828; text in 1831).

    [https://www.biodiversitylibrary.org/item/189273#page/7/mode/1up]

    https://doi.org/10.5962/bhl.title.107403

    Envall, M. (1996) Ototyphlonemertes correae sp. nov. and a redescription of O. duplex (Nemertea: Monostilifera: Ototyphlonemertidae), with a phylogenetic consideration of the genus. Journal of Zoology, 238, 253–277.

    https://doi.org/10.1111/j.1469-7998.1996.tb05393.x

    Felsenstein, J. (1978) Cases in which parsimony or compatibility methods will be positively misleading. Systematic Zoology, 27, 401–410.

    https://doi.org/10.2307/2412923

    Friedrich, H. (1933) Morphologische Studien an Nemertinen der Kieler Bucht, I und II. Zeitschrift für wissenschaftliche Zoologie, 144, 496–509.

    Friedrich, H. (1935a) Studien zur Morphologie, Systematik und Ökologie der Nemertini der Kieler Bucht. Archiv für Naturgeschichte, 4, 293–375.

    Friedrich, H. (1935b) Neue Hoplonemertinen der Kieler Bucht. Schriften des Naturwissenschaftlichen Vereins für Schleswig-Holstein, 21, 10–19.

    Friedrich, H. (1936) Nemertini. Die Tierwelt der Nord- und Ostsee, IV.d, 1–69.

    Friedrich, H. (1938) Atyponemertes korscheltii n. gen. n. spec., ein abweichender Hoplonemertinentypus aus der Nordsee. Zoologischer Anzeiger, 124, 22–30.

    Friedrich, H. (1939) Einige neue Hoplonemertinen aus der Ostsee. Kieler Meeresforschungen, 3, 233–251.

    Friedrich, H. (1955) Beiträge zu einer Synopsis der Gattungen der Nemertini Monostilifera nebst Bestimmungsschlüssel. Zeitschrift für wissenschaftliehe Zoologie, 158, 133–192.

    Friedrich, H. (1957) Beiträge zur Kenntnis der Arktischen Hoplonemertinen. Videnskabelige Meddeleser fra Dansk naturhistorisk Forening i Kjobenhavn, 119, 129–154.

    Friedrich, H. (1968) Sagaminemertes, eine bemerkenswerte neue Gattung der Hoplonemertinen und ihre systematische Stellung. Zoologischer Anzeiger, 180, 33–36.

    Friedrich, H. (1970) Nemertinen aus Chile. Sarsia, 40, 1–80.

    https://doi.org/10.1080/00364827.1970.10411156

    Frutos, I., Montalvo, S. & Junoy, J. (1998) A new species of Prosorhochmus (Hoplonemertea, Monostilifera) from the Chafarinas Islands (western Mediterranean). Journal of Zoology, 245, 293–298.

    https://doi.org/10.1111/j.1469-7998.1998.tb00104.x

    Gering, G. (1912) Neue Nemertinen der schwedischen Westküste. Zoologischer Anzeiger, 39, 520–523.

    [https://www.biodiversitylibrary.org/item/95291#page/534/mode/1up]

    Gibson, R. (1973) A new littoral hoplonemertean (Divanella evelinae gen. et sp. nov.) from the coast of Brazil. Bulletin of Marine Science, 23, 793–810.

    Gibson, R. (1974) A new species of commensal hoplonemertean from Australia. Zoological Journal of the Linnean Society, 55, 247–266.

    https://doi.org/10.1111/j.1096-3642.1974.tb01647.x

    Gibson, R. (1982a) Nemertea. In: Parker, S.P. (Ed.), Synopsis and Classification of Living Organisms. Vol. 1. McGraw-Hill, New York, pp. 823–846.

    Gibson, R. (1982b) Nemerteans of the Great Barrier Reef. 5. Enopla Hoplonemertea (Monostilifera). Zoological Journal of the Linnean Society, 75, 269–296.

    https://doi.org/10.1111/j.1096-3642.1982.tb01949.x

    Gibson, R. (1985) The need for a standard approach to taxonomic descriptions of Nemertea. American Zoologist, 25, 5–14.

    https://doi.org/10.1093/icb/25.1.5

    Gibson, R. (1986a) Redescription and taxonomic reappraisal of Nemertopsis actinophila Bürger, 1904 (Nemertea: Hoplonemertea: Monostilifera). Bulletin of Marine Science, 39, 42–60.

    Gibson, R. (1986b) A new monostiliferous hoplonemertean, Meionemertes polygonimos gen. et sp. nov., from the South Pacific (Nemertea). Zoologischer Anzeiger, 216, 279–288.

    Gibson, R. (1988) Evolutionary relationships between mono- and polystiliferous hoplonemerteans: Nipponnemertes (Cratenemertidae), a “missing link” genus? Hydrobiologia, 156, 61–74.

    https://doi.org/10.1007/BF00027979

    Gibson, R. (1989) Redescription and taxonomic reappraisal of Amphiporus elongatus Stephenson (Nemertea, Enopla) from the Firth of Clyde. Zoologica Scripta, 18, 357–364.

    https://doi.org/10.1111/j.1463-6409.1989.tb00130.x

    Gibson, R. (1990a) The macrobenthic nemertean fauna of Hong Kong. In: Morton, B.S. (Ed.), Proceedings of the Second International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China. Vol. 1. Hong Kong University Press, Hong Kong, pp. 33–212.

    Gibson, R. (1990b) The macrobenthic nemertean fauna of the Albany region, Western Australia. In: Wells, F.E., Walker, D.I., Kirkman, H. & Lethbridge, R. (Eds.), Proceedings of the Third International Marine Biological Workshop: The Marine Flora and Fauna of Albany, Western Australia. Vol. 1. Western Australian Museum, Perth, pp. 89–194.

    Gibson, R. (1994) Nemerteans. Field Studies Council, Shrewsbury, vii + 224 pp.

    Gibson, R. (1995) Nemertean genera and species of the world: an annotated checklist of original names and description citations, synonyms, current taxonomic status, habitats and recorded zoogeographic distribution. Journal of Natural History, 29, 271–561.

    https://doi.org/10.1080/00222939500770161

    Gibson, R. (1999) Further studies on the nemertean fauna of Rottnest Island, Western Australia. In: Walker, D.I. & Wells, F.E. (Eds.), The Seagrass Flora and Fauna of Rottnest Island, Western Australia. Western Australian Museum, Perth, pp. 359–376.

    Gibson, R. (2002) The Invertebrate Fauna of New Zealand: Nemertea (Ribbon Worms). National Institute of Water and Atmospheric Research, Wellington, 87 pp. [https://docs.niwa.co.nz/library/public/Memoir%20118_Marine%20Fauna%20of%20NZ_Nemertea%20(Ribbon%20Worms)%20-%202002.pdf]

    Gibson, R. & Crandall, F.B. (1989) The genus Amphiporus Ehrenberg (Nemertea, Enopla, Monostiliferoidea). Zoologica Scripta, 18, 453–470.

    https://doi.org/10.1111/j.1463-6409.1989.tb00140.x

    Gibson, R. & Crandall, F.B. (1991) Case 2707. Amphiporus Ehrenberg, 1831 (Nemertea): proposed designation of Planaria lactiflorea Johnston, 1828 as the type species. Bulletin of Zoological Nomenclature, 48, 22–24.

    [https://www.biodiversitylibrary.org/page/12230693#page/38/mode/1up]

    Gibson, R. & Jennings, J.B. (1967) “Leucine aminopeptidase” activity in the blood system of rhynchocoelan worms. Comparative Biochemistry and Physiology, 23, 645–651.

    https://doi.org/10.1016/0010-406X(67)90415-X

    Gibson, R. & Jennings, J.B. (1969) Observations on the diet, feeding mechanisms, digestion and food reserves of the entocommensal rhynchocoelan Malacobdella grossa. Journal of the Marine Biological Association of the United Kingdom, 47, 17–32.

    https://doi.org/10.1017/S0025315400046397

    Gibson, R. & Junoy, J. (1991) A new species of Tetrastemma (Nemertea: Enopla: Monostiliferoidea) from Ría de Foz, north-western Spain, found living in the mantle cavity of the bivalve mollusc Scrobicularia plana. Zoological Journal of the Linnean Society, 103, 225–240.

    https://doi.org/10.1111/j.1096-3642.1991.tb00904.x

    Gibson, R. & Moore, J. (1971) Occurrence of the freshwater hoplonemertean Prostoma graecense (Böhmig) in Tasmania. Freshwater Biology, 1, 193–195.

    https://doi.org/10.1111/j.1365-2427.1971.tb01556.x

    Gibson, R. & Moore, J. (1976) Freshwater nemerteans. Zoological Journal of the Linnean Society, 58, 177–218.

    https://doi.org/10.1111/j.1096-3642.1976.tb00828.x

    Gibson, R. & Moore, J. (1978) Freshwater nemerteans: new records of Prostoma and a description of Prostoma canadiensis sp. nov. Zoologischer Anzeiger, 201, 77–85.

    Gibson, R. & Moore, J. (1985) The genus Prosorhochmus Keferstein, 1862 (Hoplonemertea). Journal of Zoology, 206, 145–162. https://doi.org/10.1111/j.1469-7998.1985.tb05641.x

    Gibson, R. & Moore, J. (1998) Further observations on the genus Geonemertes with a description of a new species from the Philippine Islands. Hydrobiologia, 365, 157–171.

    https://doi.org/10.1023/A:1003199031726

    Gibson, R., Moore, J. & Crandall, F.B. (1982) A new semi-terrestrial nemertean from California. Journal of Zoology, 196, 463–474.

    https://doi.org/10.1111/j.1469-7998.1982.tb03518.x

    Gibson, R., Moore, J., Ruppert, E.E. & Turbeville, J.M. (1986) A new species of Prosorhochmus (Hoplonemertea, Monostilifera) from South Carolina. Journal of Zoology, 209, 327–335.

    https://doi.org/10.1111/j.1469-7998.1986.tb03595.x

    Gibson, R. & Strand, M. (2002) A new genus and species of monostiliferous hoplonemertean (Enopla: Hoplonemertea: Monostilifera) from New Zealand. Zootaxa, 50, 1–20.

    Gibson, R. & Sundberg, P. (1992) Three new nemerteans from Hong Kong. In: Morton, B. (Ed.), The Marine Flora and Fauna of Hong Kong and Southern China III. Vol. 1. Hong Kong University Press, Hong Kong, pp. 97–129.

    Gibson, R. & Sundberg, P. (2001) Some nemerteans (Nemertea) from Queensland and the Great Barrier Reef, Australia. Zoological Science, 18, 1259–1274.

    https://doi.org/10.2108/zsj.18.1259

    Gibson, R. & Tait, N.N. (1984) A redescription of Tetrastemma unilineatum Joubin 1910, and Antarctic Hoplonemertean (Nemertini). Zoologischer Anzeiger, 212, 139–150.

    Gibson, R. & Wang, H. (2002) A new genus and species of freshwater monostiliferous hoplonemertean (Nemertea, Enopla) from the People’s Republic of China. Hydrobiologia, 489, 185–196.

    https://doi.org/10.1023/A:1023236905555

    Gibson, R., Wickham, D.E. & Kuris, A.M. (1990) A new genus and species of monostiliferoidean nemertean (Nemertea: Enopla) found on an egg mass of the anomuran decapod Paralithodes camtschatica. Zoological Journal of the Linnean Society, 98, 185–198.

    https://doi.org/10.1111/j.1096-3642.1990.tb01215.x

    Girard, C. (1852) Hecate elegans. [In an untitled article on ‘descriptions of two new genera and two species of Nemertes’, presented by the Secretary of the Boston Society of Natural History]. Proceedings of the Boston Society of Natural History, 4, 186.

    [https://www.biodiversitylibrary.org/item/35742#page/194/mode/1up]

    Girard, C. (1893) Recherches sur les Planariés et les Némertiens de l’Amérique du Nord. Annales des Sciences Naturelles, Zoologie, Série 7, 15, 145–310.

    [https://www.biodiversitylibrary.org/item/111186#page/155/mode/1up]

    https://doi.org/10.5962/bhl.title.15104

    Giribet, G., Carranza, S., Baguñà, J., Riutort, M. & Ribera, C. (1996) First molecular evidence for the existence of a Tardigrada + Arthropoda clade. Molecular Biology and Evolution, 13, 76–84.

    https://doi.org/10.1093/oxfordjournals.molbev.a025573

    Girty, G.H. (1910) New genera and species of Carboniferous fossils from the Fayetteville Shale of Arkansas. Annals of the New York Academy of Sciences, 20, 189–238.

    [https://www.biodiversitylibrary.org/item/26492#page/253/mode/1up]

    https://doi.org/10.1111/j.1749-6632.1910.tb55149.x

    Gonzalez-Cueto, J., Castro, L.R. & Quiroga, S. (2017) Nipponnemertes incainca sp. n. Adoption of the new taxonomic proposal for nemerteans (Nemertea, Cratenemertidae). ZooKeys, 693, 1–15.

    https://doi.org/10.3897/zookeys.693.12015

    Graff, L. von (1879) Geonemertes chalicophora, eine neue Landnemertine. Morphologisches Jahrbuch, 5, 430–449.

    [https://www.biodiversitylibrary.org/item/45508#page/496/mode/1up]

    Griffin, B.B. (1898) Description of some marine nemerteans of Puget Sound and Alaska. Annals of the New York Academy of Sciences, 11, 193–217.

    [https://www.biodiversitylibrary.org/item/109565#page/265/mode/1up]

    https://doi.org/10.1111/j.1749-6632.1898.tb54969.x

    Grube, A.E. (1840) Actinien, Echinodermen und Würmer des Adriatischen- und Mittelmeers: nach eigenen Sammlungen beschrieben. J. H. Bon, Königsberg, 92 pp.

    [https://www.biodiversitylibrary.org/item/40483#page/5/mode/1up]

    https://doi.org/10.5962/bhl.title.23025

    Gulliver, G. (1879) Turbellaria. Philosophical Transactions of the Royal Society of London, 168, 557–563.

    [https://www.biodiversitylibrary.org/item/77050#page/573/mode/1up]

    https://doi.org/10.1098/rstl.1879.0058

    Gunnerus, J.E. (1770) Nogle smaa rare or meestendeelen nye Norske Søedyr. Skrifter som udi det Kjobenhavnske Selskab, 10, 166–176.

    [https://www.biodiversitylibrary.org/item/183235#page/184/mode/1up]

    Härlin, M. & Sundberg, P. (1995) Cladistic analysis of the eureptantic nemerteans (Nemertea: Hoplonemertea). Invertebrate Systematics, 9, 1211–1229.

    https://doi.org/10.1071/IT9951211

    Herrera-Bachiller, A., Pérez, J. & Junoy, J. (2014) Occurrence of the ribbon worm Emplectonema neesii (Nemertea: Hoplonemertea) on the Atlantic coast of Spain. Marine Biodiversity Records, 7, E8.

    https://doi.org/10.1017/S175526721300119X

    Hett, M.L. (1924) On a new land nemertean from New South Wales (Geonemertes hillii, sp. n.). Proceedings of the General Meetings for Scientific Business of the Zoological Society of London 1924 [Journal of Zoology 94], (3), 775–787.

    https://doi.org/10.1111/j.1096-3642.1924.tb03314.x

    Hickman, J.L. & Moore, J. (1990) A new species from Tasmania of the freshwater hoplonemertean, Potamonemertes. Papers and Proceedings of the Royal Society of Tasmania, 124, 51–59.

    https://doi.org/10.26749/rstpp.124.1.51.

    Hookabe, N., Tsuchida, S., Fujiwara, Y. & Kajihara, H. (2020) A new species of bathyal nemertean, Proamphiporus kaimeiae sp. nov., off Tohoku, Japan, and molecular systematics of the genus (Nemertea: Monostilifera). Species Diversity, 25 (2), 183–188.

    https://doi.org/10.12782/specdiv.25.183

    Hubrecht, A.A.W. (1874) Aanteekeningen over de Anatomie, Histologie en Ontwikkelingsgeschiedenis van eenige Nemertinen. J. L. Beijers, Utrecht, 66 pp. [http://objects.library.uu.nl/reader/index.php?obj=1874-9848&lan=en#page//16/41/44/164144307394886498884867375865787646542.jpg/mode/1up]

    Hubrecht, A.A.W. (1879) The genera of European nemerteans critically revised, with description of several new species. Notes from the Leyden Museum, 1, 193–232.

    [https://www.biodiversitylibrary.org/item/37312#page/205/mode/1up]

    Hubrecht, A.A.W. (1887) Report on the Nemertea collected by H. M. S. Challenger during the years 1873–76. Reports of the Scientific Results of the Voyage of H. M. S. Challenger 1873–76, Zoology, 19, 1–150.

    [https://www.biodiversitylibrary.org/item/29739#page/17/mode/1up]

    https://doi.org/10.5962/bhl.title.3925

    Humes, A.G. (1942) The morphology, taxonomy, and bionomics of the nemertean genus Carcinonemertes. Illinois Biological Monographs, 18, 1–105.

    https://doi.org/10.5962/bhl.title.50200

    International Commission on Zoological Nomenclature (1992) Opinion 1675. Amphiporus Ehrenberg, 1831 (Nemertea): Planaria lactiflorea Johnston, 1828 designated as the type species. Bulletin of Zoological Nomenclature, 49, 157.

    [https://www.biodiversitylibrary.org/item/44490#page/177/mode/1up]

    International Commission on Zoological Nomenclatutre (1999) International Code of Zoological Nomenclature. 4th Edition. International Trust for Zoological Nomenclature, London, 306 pp.

    International Commission on Zoological Nomenclature (2003) Opinion 2016 (Case 2888). Valdivianemertes Stiasny-Wijnhoff, 1923 (Nemertea): not concerved. Bulletin of Zoological Nomenclature, 60, 59–60.

    [https://archive.org/details/biostor-80528]

    Iwata, F. (1952) Nemertini from the coasts of Kyusyu. Journal of the Faculty of Science, Hokkaido University, Series 6, Zoology, 11, 126–148.

    [https://eprints.lib.hokudai.ac.jp/dspace/bitstream/2115/27119/1/11%281%29_P126-148.pdf]

    Iwata, F. (1954) The fauna of Akkeshi Bay. XX. Nemertini in Hokkaido. Journal of the Faculty of Science, Hokkaido University, Series 6, Zoology, 12, 1–39.

    [https://eprints.lib.hokudai.ac.jp/dspace/bitstream/2115/27137/1/12%281_2%29_P1-39.pdf]

    Iwata, F. (1960) Studies of the comparative embryology of nemerteans with special reference to their interrelationships. Publications from the Akkeshi Marine Biological Station, 10, 1–51.

    [https://eprints.lib.hokudai.ac.jp/dspace/bitstream/2115/68363/1/Akkeshi_No10.pdf]

    Iwata, F. (1970) On the brackish water nemertens from Japan, provided with special circulatory and nephridial organs useful for osmoregulation. Zoologischer Anzeiger, 184, 133–154.

    Iwata, F. (2001) Nipponnemertes fernaldi, a new species of swimming monostiliferous hoplonemertean from the San Juan Archipelago, Washington, U.S.A. Proceedings of the Biological Society of Washington, 114, 833–857.

    [https://www.biodiversitylibrary.org/item/107511#page/283/mode/1up]

    Iwata, F. (2006) On four monostiliferous hoplonemerteans, including three new genera and species from Washington State and British Columbia. Journal of Natural History, 40, 873–913.

    https://doi.org/10.1080/00222930600833800

    Iwata, F. (2007) Cyanonemertes elegans, new genus, new species, a marine hoplonemertean from Washington State, U.S.A. Proceedings of the Biological Society of Washington, 120, 197–213.

    https://doi.org/10.2988/0006-324X(2007)120[197:CENGNS]2.0.CO;2

    Johnston, G. (1828) Contributions to the British fauna. Zoological Journal, 3, 486–491.

    [https://www.biodiversitylibrary.org/item/19597#page/548/mode/1up]

    Johnston, G. (1837) Miscellanea zoologica. II. A description of some planarian worms. Magazine of Zoology and Botany, 1, 529–538.

    [https://www.biodiversitylibrary.org/item/110264#page/543/mode/1up]

    Joubin, L. (1890) Recherches sur les Turbellariés des côtes de France (Némertes). Archives de Zoologie Expérimentale et Générale, Série 2, 8, 461–602.

    [https://www.biodiversitylibrary.org/item/28409#page/519/mode/1up]

    Joubin, L. (1894) Les Némertiens. Société d’Éditions Scientifiques, Paris, 235 pp.

    [https://books.google.com.tw/books?id=svxGAQAAMAAJ&printsec=frontcover&source=gbs_ge_summary_r&cad=0#v=onepage&q&f=false]

    Joubin, L. (1902) Némertiens. In: Milne-Edwards, A. & E. Perrier, E. (Eds.), Expéditions scientifiques du “Travailleur” et du “Talisman” pendant les années 1880, 1881, 1882, 1883. Vol. 6. Masson, Paris, pp. 181–220.

    Joubin, L. (1910) Nemertinea. National Antarctic Expedition, 1901–1904, 5, 1–15.

    Junoy, J., Andrade, S.C.S. & Giribet, G. (2011) Phylogenetic placement of a new hoplonemertean species commensal on ascidians. Invertebrate Systematics, 24, 616–629.

    https://doi.org/10.1071/IS10036

    Junoy, J. & Gibson, R. (1992) Primeras citas de los nemertinos Oerstedia dorsalis (Abildgaard, 1806) y Tetrastemma vermiculus (Quatrefages, 1846) (Nemertea, Enopla, Monostiliferoidea) para las costas ibéricas. Boletín de la Real Sociedad Española de Historia Natural, 88, 165–177.

    Kajihara, H. (2002) Two new species of Zygonemertes (Nemertea: Enopla: Monostilifera) from Hokkaido, Japan. Species Diversity, 7, 121–143.

    https://doi.org/10.12782/specdiv.7.121

    Kajihara, H. (2007a) A taxonomic catalogue of Japanese nemerteans (phylum Nemertea). Zoological Science, 24, 287–326.

    https://doi.org/10.2108/zsj.24.287

    Kajihara, H. (2007b) Ototyphlonemertes dolichobasis sp. nov. (Nemertea: Hoplonemertea: Monostilifera: Ototyphlonemertidae), a new species of interstitial nemertean from Japan. Species Diversity, 12, 57–66.

    https://doi.org/10.12782/specdiv.12.57

    Kajihara, H. (2007c) Two species of Nemertopsis (Nemertea: Hoplonemertea: Monostilifera) living in association with Capitulum mitella (Crustacea: Cirripedia: Thoracica: Lepadomorpha). Zootaxa, 1446 (1), 43–58.

    https://doi.org/10.11646/zootaxa.1446.1.4

    Kajihara, H. (2017) Species diversity of Japanese ribbon worms (Nemertea). In: Motokawa, M. & Kajihara, H. (Eds.), Species Diversity of Animals in Japan. Springer Japan, Tokyo, pp. 419–444.

    https://doi.org/10.1007/978-4-431-56432-4_16

    Kajihara, H., Chernyshev, A.V., Sun, S.-C., Sundberg, P. & Crandall, F.B. (2008) Checklist of nemertean genera and species published between 1995 and 2007. Species Diversity, 13, 245–274.

    https://doi.org/10.12782/specdiv.13.245

    Kajihara, H., Gibson, R. & Mawatari, S.F. (2000) Redescription and taxonomic reassessment of Nemertellina minuta Friedrich, 1935, sensu Yamaoka, 1940 (Nemertea, Hoplonemertea, Monostilifera). Zoological Science, 17, 265–276.

    https://doi.org/10.2108/zsj.17.265

    Kajihara, H., Gibson, R. & Mawatari, S.F. (2001) A new genus and species of monostiliferous hoplonemertean (Nemertea: Enopla: Monostilifera) from Japan. Hydrobiologia, 456, 187–198.

    https://doi.org/10.1023/A:1013047207243

    Kajihara, H., Gibson, R. & Mawatari, S.F. (2003) Potamostoma shizunaiense gen. et sp. nov. (Nemertea: Hoplonemertea: Monostilifera): a new brackish-water nemertean from Japan. Zoological Science, 20, 491–500.

    https://doi.org/10.2108/zsj.20.491

    Kajihara, H. & Kuris, A. (2013) Ovicides paralithodis (Nemertea, Carcinonemertidae), a new species of symbiotic egg predator of the red king crab Paralithodes camtchaticus (Tilesius, 1815) (Decapoda, Anomura). ZooKeys, 258, 1–15.

    http://doi.org/10.3897/zookeys.258.4260

    Kajihara, H., Nishi, E., Kawabata, M., Kohtsuka, H. & Uyeno, D. (2015) Records of the poorly known ribbon worm Nipponnemertes ogumai (Nemertea: Monostilifera) and its phylogenetic position. Marine Biodiversity, 45, 175–182.

    https://doi.org/10.1007/s12526-014-0252-1

    Kajihara, H., Olympia, M., Kobayashi, N., Katoh, T., Chen, H.-X., Strand, M. & Sundberg, P. (2011) Systematics and phylogeny of the hoplonemertean genus Diplomma (Nemertea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society, 161, 695–722.

    https://doi.org/10.1111/j.1096-3642.2010.00650.x

    Karling, T.G. (1933) Ein Beitrag zur Kenntnis der Nemertinen des Finnischen Meerbusens. Memoranda Societatis pro Fauna et Flora Fennica, 10, 76–90.

    Keferstein, W. (1862) Untersuchungen über niedere Seethiere. Zeitschrift für wissenschaftliche Zoologie, 12, 1–147.

    [https://www.biodiversitylibrary.org/item/104150#page/9/mode/1up]

    Kennel, J.v. (1877–1878) Beiträge zur Kenntniss der Nemertinen. Arbeiten aus dem Zoologisch-Zootomischen Institut in Würzburg, 4, 305–360, pls. XVII–XIX.

    [https://www.biodiversitylibrary.org/item/106735#page/311/mode/1up]

    Kirsteuer, E. (1963) Beitrag zur Kenntnis der Systematik und Anatomie der adriatischen Nemertinen (Genera Tetrastemma, Oerstedia, Oerstediella). Zoologische Jahrbücher, Abteilung für Anatomie und Ontogenie der Tiere, 80, 555–616.

    Kirsteuer, E. (1965) Über das Vorkommen von Nemertinen in einem tropischen Korallenriff. 4. Hoplonemertini monostilifera. Zoologische Jahrbiicher, Abteilung für Systematik, Okologie und Geographie der Tiere, 92, 289–326.

    Kirsteuer, E. (1967) New marine nemerteans from Nossi Be, Madagascar. Zoologischer Anzeiger, 178, 110–122.

    Kirsteuer, E. (1974) Description of Poseidonemertes caribensis sp. n., and discussion of other taxa of Hoplonemertini Monostilifera with divided longitudinal musculature in the body wall. Zoologica Scripta, 3, 153–166.

    https://doi.org/10.1111/j.1463-6409.1974.tb00812.x

    Kölliker, A. (1845) Drei neue Gattungen von Würmern. Verhandlungen der Schweizerischen Naturforschenden Gessellschaft bei ihrer versammlung zu Chur, 29, 86–98.

    Koroleva, A.G., Chernyshev, A.V., Kiril’chik, S.V., Tasevska, O., Kostoski, G. & Timoshkin, O.A. (2014) The first finding of freshwater nemerteans in Ohrid Lake (Macedonia) with some comments on the taxonomy of the genus Prostoma (Nemertea, Monostilifera). Biology Bulletin, 41, 736–741.

    https://doi.org/10.1134/S1062359014090052

    Korotkevitsch, V.S. (1961) A new nemertean species and its position in the system. Zoologicheskii Zhurnal, 40, 1416–1420.

    Korotkevitsch, V.S. (1977) Nemertinui pribrezhnuikh rod kurilskikh ostrovov. In: Gulbin, V.V. (Ed.), Fauna Pribrezhnuikh zon Kurilskikh Ostrovov. Izdatelstvo Nauka, Moscow, pp. 49–124. [in Russian]

    Korotkevitsch, V.S. (1983) O stiletakh nemertin i novom vide Cratenemertes (Hoplonemertini, Amphiporidae) iz Antarktiki. Antarktika Doklady Komissii, 22, 137–143. [in Russian]

    Krämer, D. & Döhren, J. von (2015) Arenogigas armoricus, a new genus and species of a monostiliferous hoplonemertean (Nemertea) from the north-west coast of France. Zoological Science, 32, 605–614.

    https://doi.org/10.2108/zs140266

    Kulikova, V.I. (1987) New species of the genus Oerstediella (Nemertini, Hoplonemertini) from the Vostok Bay, Sea of Japan. Izvestiya Akademii Nauk SSSR, 6, 828–836.

    Kulikova, V.I. (1989) Novuii vid roda Tetrastemma (Nemertini, Hoplonemertini) iz yaponskogo morya. Izvestiya Akademii Nauk SSSR, Seriya Biologicheskaya, 4, 628–631.

    Kvist, S., Laumer, C. E., Junoy, J. & Giribet, G. (2014) New insights into the phylogeny, systematics and DNA barcoding of Nemertea. Invertebrate Systematics, 28, 287–308.

    https://doi.org/10.1071/IS13061

    Kvist, S., Chernyshev, A.V. & Giribet, G. (2015) Phylogeny of Nemertea with special interest in the placement of diversity from Far East Russia and northeast Asia. Hydrobiologia, 760, 105–119.

    https://doi.org/10.1007/s10750-015-2310-5

    Lagunas-Calvo, O., Hernández-Mena, D.I., Rivas, G., Contreras-Mirón, S., Mendoza-Garfias, B., Echeverría-Martínez, O.M. & Oceguera-Figueroa, A. (2018) Rediscovery of the nemertean Prostoma graecense from Xochimilco, 75 years after its first and only record. Revista Mexicana de Biodiversidad, 89, 39–45.

    https://doi.org/10.22201/ib.20078706e.2018.1.2094

    Leasi, F., Andrade, S.C.S. & Norenburg, J.L. (2016) At least some meiofaunal species are not everywhere. Indication of geographic, ecological and geological barriers affecting the dispersion of species of Ototyphlonemertes (Nemertea, Hoplonemertea). Molecular Ecology, 25, 1381–1397.

    https://doi.org/10.1111/mec.13568

    Leidy, J. (1850) Description of new genera of Vermes. Proceedings of the Academy of Natural Sciences of Philadelphia, 5, 124–126. https://www.biodiversitylibrary.org/item/84765#page/158/mode/1up

    Maclaren, N.H.W. (1901) On the blood vascular system of Malacobdella grossa. Zoologischer Anzeiger, 24, 126–129.

    [https://www.biodiversitylibrary.org/item/37592#page/142/mode/1up]

    Magarlamov, T.Y. & Chernyshev, A.V. (2009) Ultrastructure of larval epidermis of the nemertean Quasitetrastemma stimpsoni (Chernyshev, 1992) (Hoplonemertea). Russian Journal of Marine Biology, 35, 34–40.

    https://doi.org/10.1134/S1063074009010064

    Maslakova, S.A & Norenburg, J.L. (2001) Phylogenetic study of pelagic nemerteans (Pelagica, Polystilifera). Hydrobiologia, 456, 111–132.

    https://doi.org/10.1023/A:1013048419113

    Maslakova, S.A. & Norenburg, J.L. (2008a) Revision of the smiling worms, genus Prosorhochmus Keferstein, 1862, and description of a new species, Prosorhochmus belizeanus sp. nov. (Prosorhochmidae, Hoplonemertea, Nemertea) from Florida and Belize. Journal of Natural History, 42, 1219–1260.

    https://doi.org/10.1080/00222930801995747

    Maslakova, S.A. & Norenburg, J.L. (2008b) Revision of the smiling worms, genera Prosadenoporus Bürger, 1890 and Pantinonemertes Moore and Gibson, 1981 and description of a new species Prosadenoporus floridensis sp. nov. (Prosorhochmidae; Hoplonemertea; Nemertea) from Florida and Belize. Journal of Natural History, 42, 1689–1727.

    https://doi.org/10.1080/00222930802130286

    Maslakova, S.A., Thiel, M., Vásquez, N. & Norenburg, J.L. (2005) The smile of Amphiporus nelsoni Sanchez, 1973 (Nemertea: Hoplonemertea: Monostilifera: Amphiporidae) leads to a redescription and a change in family. Proceedings of the Biological Society of Washington, 118, 483–498.

    https://doi.org/10.2988/0006-324X(2005)118[483:TSOANS]2.0.CO;2

    Maslakova, S.A. & Döhren, J. von (2009) Larval development with transitory epidermis in Paranemertes peregrina and other hoplonemerteans. Biological Bulletin, 216, 273–292.

    https://doi.org/10.1086/BBLv216n3p273

    Mateos, E. & Giribet, G. (2008) Exploring the molecular diversity of terrestrial nemerteans (Hoplonemertea, Monostilifera, Acteonemertidae) in a continental landmass. Zoologica Scripta, 37, 235–243.

    https://doi.org/10.1111/j.1463-6409.2008.00324.x

    McDermott, J.J. (1976) Observations on the food and feeding behavior of estuarine nemertean worms belonging to the order Hoplonemertea. Biological Bulletin, 150, 57–68.

    https://doi.org/10.2307/1540589

    McDermott, J.J. & Gibson, R. (1993) Carcinonemertes pinnotheridophila sp. nov. (Nemertea, Enopla, Carcinonemertidae) from the branchial chambers of Pinnixa chaetopterana (Crustacea, Decapoda, Pinnotheridae): description, incidence and biological relationships with the host. Hydrobiologia, 266, 57–80.

    https://doi.org/10.1007/BF00013359

    McDermott, J.J. & Roe, P. (1985) Food, feeding behavior and feeding ecology of nemerteans. American Zoologist, 25, 113–125.

    https://doi.org/10.1093/ICB/25.1.113

    McIntosh, W.C. (1873–1874) s.n. In: A Monograph of the British Annelids. Part I. The Nemerteans. Ray Society, London, pp. 1–96, pls. I–X (published in 1873) + pp. 97–214, pls. XI–XXIII (published in 1874).

    [https://www.biodiversitylibrary.org/item/118767#page/7/mode/1up]

    Monastero, S. (1930) I ne