Abstract
Describing ontogenetic morphological change is an important part of integrative taxonomy; still, most taxonomic studies are based only on adult characters. Here, we provide illustrations and a morphological description of the sea stars from Bahia, including ontogenetic and intraspecific variation, and identify taxonomic issues. A total of 293 specimens from different localities along the Bahia State coastline and comparative material from other localities were examined. Eighteen species (11 genera, eight families) of Asteroidea were identified; Astropectinidae was the most representative family. All species identified also occur in subtropical Brazilian waters and most species are from shallow water habitats with soft bottoms. Most observed ontogenetic variation was quantitative in nature, such as the increase in the number of spines in the furrow and of spinelets in the paxillae with specimen growth. Genera that require further taxonomic studies are Astropecten and Othilia, whose specimens are commonly misidentified in local studies. One third of the species from Bahia are currently classified as “Vulnerable” in the Brazilian Red List, but baseline data on the population biology of these species are scarce. An illustrated identification key to the 65 Brazilian sea star species is also provided. This taxonomic study will facilitate the identification of specimens occurring along the Brazilian coast and help scientists and policy makers to establish the conservation status of the Brazilian species.
References
Abreu-Pérez, M., Solís-Marín, F.A. & Laguarda-Figueras, A. (2005) Catálogo de los equinodermos (Echinodermata: Asteroidea y Ophiuroidea) nerítico-bentónicos del Arquipiélago Cubano. Revista de Biologia Tropical, 53, 29–52.
Agassiz, L. (1835) Prodrome d’une monographic des radiares ou echinodermes. Memoires de la Société des Sciences Naturelles de Neuchatel, 1, 168–199.
Agostini, V.O. & Ozorio, C.P. (2018) Caracterização da macrofauna de substratos consolidados do Litoral Norte do Rio Grande do Sul, Brasil. Arquivos de Ciências do Mar, 51 (1), 26–48.
https://doi.org/10.32360/acmar.v51i1.31293
Alcantara, A.A. & Costa, F.M. (2019) Resposta comportamental de Echinaster (Othilia) brasiliensis Müller & Troschel, 1842, (Echinodermata, Asteroidea) com base em preferência alimentar e observação em cativeiro. Acta Scientiae et Technicae, 7 (1), 21–25.
https://doi.org/10.17648/uezo-ast-v7i1.263
Alitto, R.A.S., Bueno, M.L., Domenico, M. & Borges, M. (2016) Annotated checklist of Echinoderms from Araçá Bay, Southeastern Brazil. Check List, 12 (1), 1–15.
https://doi.org/ 10.15560/12.1.1836
Alvarado, J.J., Solís-Marín, F.A. & Ahearn, C. (2008) Equinodermos (Echinodermata) del Caribe Centroamericano. Revista de Biologia Tropical, 56, 37–55.
Alvarado, J.J. & Solís-Marín, F.A. (2013) Echinoderm research and diversity in Latin America. Vol. 1. Springer, Berlin, 658 pp.
https://doi.org/10.1007/978-3-642-20051-9
Alvarado, J.J., Chacón-Monge, J.L., Solís-Marín, F.A., Pineda-Enríquez, T., Caballero-Ochoa, A.A., Rivera, S.S. & Chaves, R.R. (2017) Equinodermos del Museo de Zoología de la Universidad de Costa Rica. Revista de Biología Tropical, Universidad de Costa Rica, 65 (1), S272–S287.
https://doi.org/10.15517/rbt.v65i1-1.31695
Alves, O.F.S. & Cerqueira, W.R.P. (2000) Echinodermata das praias de Salvador (Bahia, Brasil). Revista Brasileira de Zoologia, 17 (2), 543–553.
https://doi.org/10.1590/S0101-81752000000200024
Alves, R.N., Neto, N.L., Brooks, S.E. & Albuquerque, U.P. (2009) Commercialization of animal-derived remedies as complementary medicine in the semi-arid region of Northeastern Brazil. Journal of Ethnopharmacology, 124, 600–608.
https://doi.org/10.1016/j.jep.2009.04.049
Alves, R.N. & Dias, T.L.P. (2010) Uso de invertebrados na medicina popular no Brasil e suas implicações para conservação. Tropical Conservation Science, 3 (2), 159–174.
https://doi.org/10.1177/194008291000300204
Alves R.R. & Alves H.N. (2011) The faunal drugstore: animal-based remedies used in traditional medicines in Latin America. Journal of Ethnobiology and Ethnomedicine, 10, 1746–4269.
https://doi.org/10.1186/1746-4269-7-9
Alves, R.R.N. & Rosa, I.L. (2007) Biodiversity, traditional medicine and public health: where do they meet? Journal of Ethnobiology and Ethnomedicine, 3, 1–9.
https://doi.org/10.1186/1746-4269-3-14
Alves, R.R.N., Rosa, I.L., Neto, N.L. & Voeks, R. (2012) Animals for the Gods: Magical and Religious Faunal Use and Trade in Brazil. Human Ecology, 40, 751–780.
https://doi.org/10.1007/s10745-012-9516-1
Alves S.L.S., Pereira, A.D. & Ventura, C.R.R. (2002) Sexual and asexual reproduction of Coscinasterias tenuispina (Echinodermata: Asteroidea) from Rio de Janeiro, Brazil. Marine Biology, 140, 95–101.
https://doi.org/10.1007/s002270100663
Amaral, A.C.Z. & Rossi-Wongtschowski, C.L.D.B. (2004) Biodiversidade Bentônica da Região Sudeste-Sul do Brasil –Plataforma Externa e Talude Superior. Série Documentos REVIZEE—Score Sul. Instituto Oceanográfico—USP, São Paulo, 216 pp.
Asbjornsen, P.C. (1856) Description d’un nouveau genre des Astéries. In: Sars, M., Koren, J. & Danielssen, D.C. (Eds.), Fauna littoralis Norvegiae. Vol. 2. Beyer (1846–1877), Bergen, pp. 95–101.
Ávila-Pires, T.C.S.A. (1983) Contribuição ao estudo do gênero Echinaster Müller & Troschel, 1840 (Echinodermata, Asteroidea) no litoral brasileiro. Anais da Academia Brasileira de Ciências, 55 (4), 431–448.
Bayer, F., Voss, G. & Robin, R. (1970) Bioenvironmental and radiological safety feasibility studies Atlantic-Pacific interoceanic canal. In: Report on the marine fauna and benthic shelf slope communities of the isthmian region. Vol. 1. University of Miami, Rosenstiel School of Marine and Atmospheric Science, Florida, pp. 99.
Bell, F.J. (1917) Echinoderma, Part I. Actinogonidiata. Natural History Reports of the British Antarctic “Terra Nova” Expedition, Zoology, 4 (1), 1–10.
Bell, F.J. (1881) Account of the Echinodermata collected during the Survey of H.M.S. “Alert” in the Straits of Magellan and on the Coast of Patagonia. Proceedings of the Zoological Society of London, 1881, 87–101.
Benavides-Serrato, M., Borrero-Pérez, G.H., Navas, G.R. & Solano, O.D. (2005) Listado taxonómico de los asteroideos (Echinodermata: Asteroidea) de la plataforma y el talud superior del caribe colombiano. Revista de Biología Tropical, 53 (13), 171–194.
Benavides-Serrato, M., Borrero-Pérez, G. & Diaz-Sanchez, C. (2011) Equinodermos del caribe colombiano: Crinodea, Asteroidea y Ophiuroidea. Vol. 1. Marquillas, Santa Marta, 384 pp.
Bernasconi, I. (1941) Los equinodermos de la expedicion del buque oceanografico comodoro rivadavia A.R.A. Physis, 19, 37–49.
Bernasconi, I. (1943) Los Asteroideos sulamericanos de la família Luidiidae. Anales del Museo Argentino de Ciencias Naturales, 7, 1–20.
Bernasconi, I. (1955) Equinoideos y asteroideos de la colección del instituto oceanografico de la Universidad de San Pablo. Primera contribución. Boletim do Instituto Oceanográfico, 6, 51–57.
https://doi.org/10.1590/S0373-55241955000100002
Bernasconi, I. (1956) Dos nuevos equinodermos de la costa del Brasil. Neotropica, 2, 33–36, 2 figs.
Bernasconi, I. (1957) Equinoideos y asteroideos de la colección del instituto oceanografico de la Universidad De San Pablo. Segunda Contribución. Boletim do Instituto Oceanográfico, 7, 119–148.
https://doi.org/10.1590/S0373-55241956000100008
Bernasconi, I. (1958) Asteroideos de la colección del Instituto Oceanográfico de la Universidad de San Pablo. Boletim do Instituto Oceanográfico, 9, 13–19.
https://doi.org/10.1590/S0373-55241958000100002
Bernasconi, I. (1973) Los equinodermos colectados por el “Walter Herwig” en el Atlántico Sudoeste. Revista del Museo Argentino de Ciencias Naturales, 3, 287–334.
Bitter, R. (1984) Composición de la dieta de Astropecten marginatus (Echinodermata: Asteroidea). Golfo Triste, región Centro-Occidental. Boletin del Instituto Oceanografico, Universidad de Oriente, 23, 169–176.
Blainville, E. (1834) Manuel d’Actinologie ou de Zoophytologie. F.G. Levrault, Paris, 694 pp., 99 pls.
https://doi.org/10.5962/bhl.title.8768
Blake, D.B. (1987) Classification and phylogeny of post-Paleozoic sea stars (Asteroidea: Echinodermata). Journal of Natural History, 21, 481–528.
https://doi.org/10.1080/00222938700771141
Blake, D.B. (1990) Adaptive zones of the class Asteroidea (Echinodermata). Adaptive zone, 46 (3), 701.
Borges, M., Alitto, R.A.S. & Amaral, A.C.Z. (2015) From baby to adult: ontogenetic series of nine species of Ophiuroidea from Atlantic Southwestern. Revista de Biologia Tropical, 63, (Suppl. 2), 361–381.
Borrero-Pérez, G.H., Benavides-Serrato, M., Néstor, H.C., Galeano-Galeano, E., Gavio, B., Medina, J. & Abril-Howard, A. (2019) Echinoderms of the Seaflower Biosphere Reserve: state of knowledge and new findings. Frontiers in Marine Science, 6, 188.
https://doi.org/10.3389/fmars.2019.00188
Böttger, S.A. & Mcclintock, J.B. (2002) Effects of inorganic and organic phosphate exposure on aspects of reproduction in the common sea urchin Lytechinus variegatus (Echinodermata: Echinoidea). Journal of Experimental Zoology, 7, 660–671.
https://doi.org/10.1002/jez.10104
Brites, A.D., Hadel, V.F. & Tiago, C.G. (2008a) Luidia senegalensis (Lamarck, 1816). In: Machado, A.B.M., Drummond, G.M. & Paglia, A.P. (Eds.), Livro Vermelho da Fauna Brasileira Ameaçada de Extinção. Vol. 1. Ministério do Meio Ambiente e Fundação Biodiversitas, Brasília, pp. 181–182.
Brites, A.D., Hadel, V.F. & Tiago, C.G. (2008b). Echinaster (Othilia) brasiliensis and Echinaster (Othilia) echinophorus. In: Machado, A.B.M.; Drummond, G.M. & Paglia, A.P. (Eds.), Livro Vermelho da Fauna Brasileira Ameaçada de Extinção. Vol. 1. Ministério do Meio Ambiente e Fundação Biodiversitas, Brasília, pp. 182–183.
Brites, A.D., Hadel, V.F. & Tiago, C.G. (2008c) Narcissia trigonaria Sladen, 1889. In: Machado, A.B.M.; Drummond, G.M. & Paglia, A.P. (Eds.), Livro Vermelho da Fauna Brasileira Ameaçada de Extinção. Vol. 1. Ministério do Meio Ambiente e Fundação Biodiversitas, Brasília, pp. 189–190.
Brito, I.M. (1960) Asteroides dos estados do Rio de Janeiro e de São Paulo. Universidade do Brasil, Faculdade Nacional de Filosofia, Centro de Estudos Zoológicos, 5, 1–13.
Brito, I.M. (1962) Ensaio de cataìlogo dos equinodermas do Brasil. Universidade do Brasil, Faculdade Nacional de Filosofia, Centro de Estudos Zooloìgicos, 13, 1–10.
Brito, I.M. (1968) Asteroìides e equinoìides do Estado da Guanabara e adjacências. Boletim do Museu Nacional, 260, 1–51.
Brito, I.M. (1971) Contribuição ao conhecimento dos Equinodermas da Ilha da Trindade, Brasil. Arquivos do Museu Nacional, 54, 261–265.
Bueno, M.L., Alitto, R.A.S., Guilherme P.D.B., Domenico, M.D. & Borges, M. (2018) Guia ilustrado dos Echinodermata da porção sul do Embaiamento Sul Brasileiro. Pesquisa e Ensino em Ciências Exatas e da Natureza, 2 (2), 169–237. https://doi.org/10.29215/pecen.v2i2.1071
Bumbeer, J., Cattani, A.P., Chierigatti, N.B. & Rocha, R.M. (2016) Biodiversity of benthic macroinvertebrates on hard substrates in the Currais Marine Protected Area, in southern Brazil. Biota Neotropica, 16 (4), e20160246.
https://doi.org/10.1590/1676-0611-BN-2016-0246
Campbell, A.C. (1983) Form and function of pedicellariae: A review. In: Jangoux, M. & Lawrence, J.M. (Eds.), Echinoderm studies. Vol. 1. Balkema, Rotterdam, pp. 139–167.
https://doi.org/10.1201/9781003079071-6
Campbell, D.B. & Turner, R.L. (1984) Echinaster graminicola, a new species of spinulosid sea star (Echinodermata: Asteroidea) from the west coast of Florida, U.S.A. Proceedings of the Biological Society of Washington, 97, 167–178.
https://doi.org/10.1126/science.226.4671.167
Campos, L.S., Barboza, C.A.M., Alcântara, P.F., Moura, R.B., Frensel, R. & Wanderley, P. (2010a) Echinodermata. In: Lavrado, H.P. & Brasil, A.C.S. (Eds.), Biodiversidade da região oceânica profunda da Bacia de Campos: megafauna e ictiofauna demersal. Vol. 1. SAG Serv, Rio de Janeiro, pp. 235–292.
Campos, L.S., Moura, R.B., Alcântara, P.F., Vasconcelos, R.F., Curbelo-Fernandez, M.P., Veríssimo, I. & Cavalcanti, G.H. (2010b) On two new records of the Family Brisingidae (Echinodermata: Asteroidea) from the Brazilian continental margin. In: Harris, L.G., Boettger, S.A., Walker, C.W. & Lesser, M.P. (Eds.), Proceedings of the 12th International Echinoderm Conference, Durham, New Hampshire, 7–11 August 2006. CRC Press Boca Raton, Florida and Balkema Taylor & Francis Group, London, pp. 139–146.
https://doi.org/10.1201/9780203869543-c21
Campos, L.S., Moura, R.B., Veríssimo, I., Curbelo-Fernandez, M.P., Cavalcanti, G.H. & Brasil, A.C.S. (2010c) ROV imaging of deep-sea echinoderms from the Brazilian continental margin, Southwest Atlantic. In: Harris, L.G., Boettger, S.A., Walker, C.W. & Lesser, M.P. (Eds.), Proceedings of the 12th International Echinoderm Conference, Durham, New Hampshire, 7–11 August 2006. CRC Press Boca Raton, Florida and Balkema Taylor & Francis Group, London, pp. 147–152.
https://doi.org/10.1201/9780203869543-c22
Carmo, A.V., Okada, N.B. & Freret-Meurer, N.V. (2015) Caracterização do microhabitat da estrela-do-mar Linckia guildingii Gray, 1840 que ocorre na Praia de Araçatiba, Ilha Grande, Angra dos Reis, Brasil. Revista Pensar Biologia, 1 (1), 22.
Carrera-Rodriguez, C.J. & Tommasi, L.R. (1977) Asteroidea de la plataforma continental de Río Grande do Sul (Brasil), coleccionados durante los viajes del N/Oc. “Prof. W. Besnard” para el proyecto Río Grande do Sul. Boletim do Instituto Oceanográfico da Universidade de São Paulo, 26 (1), 51–130.
https://doi.org/10.1590/S0373-55241977000100004
Caso, M.E. (1944) Estudio sobre astéridos de México. Algunas especies interesantes de astéridos litorales. Anales del Instituto de Biologia Universidad Nacional Autonoma de Mexico, 15, 237–257.
Christensen, A.M. (1970) Feeding biology of the sea star Astropecten irregularis Pennant. Ophelia, 8, 1–134.
https://doi.org/10.1080/00785326.1970.10429554
Clark, A.H. (1916) Six new starfishes from the Gulf of California and adjacent waters. Proceedings of the Biological Society of Washington, 29, 51–62.
Clark, A.H. (1934) A new starfish from Puerto Rico. Smithsonian Miscellaneous Collections, 91 (14), 1–3.
Clark, A.H. (1938) A new genus of starfishes from Puerto Rico. Smithsonian Miscellaneous Collections, 91 (29), 1–7, 1 pl.
Clark, A.H. (1948) Two new starfishes and a new brittle star from Florida and Alabama. Proceedings of the Biological Society of Washington, 61, 55–66.
Clark, A.M. (1981) Notes on Atlantic and other Asteroidea. 1. Family Benthopectinidae. Bulletin of the British Museum of Natural History, Zoology, 41, 91–135.
Clark, A.M. (1983) Notes on Atlantic and other Asteroidea. 3. The families Ganeriidae and Asterinidae, with description of a new asterinid genus. Bulletin of the British Museum of Natural History, 45 (7), 359–380.
https://doi.org/10.5962/bhl.part.28006
Clark, A.M. (1987) Notes on Atlantic and other Asteroidea Echinasteridae. Bulletin of the British Museum of Natural History, 53, 65–78.
Clark, A.M. & Davies, P.S. (1966) Echinoderms of the Maldives. Annals and Magazine of Natural History, 13, 8 (94), 597–612.
https://doi.org/10.1080/00222936508651619
Clark, A.M. & Downey, M.E. (1992) Starfishes of the Atlantic. Chapman and Hall, London, 794 pp.
https://doi.org/10.5860/choice.30-2659
Clark, A.M. & Rowe, F.W.E. (1971) Monograph of Shallow-water Indo-West Pacific Echinoderms. Bulletin of the British Museum of Natural History, 690 IX, 1–238, 31 pls.
Clark, A. & Tortonese, E. (1986) Echinaster sepositus madseni n. subsp., from West Africa (Echinodermata: Asteroidea). Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale, Milano, 127, 65–71.
Clark, H.E.S. & McKnight, D.G. (2000) The marine fauna of New Zealand: Echinodermata: Asteroidea (sea-stars), Order Paxillosida, Order Notomyotida. NIWA Biodiversity Memoir, Wellington, New Zealand, 196 pp.
Clark, H.E.S. & McKnight, D.G. (2001) The Marine Fauna of New Zealand: Echinodermata: Asteroidea (sea-stars), Order Valvatida. NIWA Biodiversity Memoir, Wellington, 270 pp.
Clark, H.L. (1901) The echinoderms of Porto Rico. Bulletin of the United States Fish Commission, 20, 233–263, 4 pls.
Clark, H.L. (1910) The echinoderms of Peru. Bulletin of the Museum of Comparative Zoology at Harvard, 52, 321–358, 14 pls.
Clark, H.L. (1921) The echinoderm fauna of Torres Strait: Its composition and its origin. Carnegie Institution of Washington Publication, 10, 1–223.
https://doi.org/10.5962/bhl.title.14613
Clark, H.L. (1923) The Echinoderm fauna of South Africa. Annals of the South African Museum, 13 (7), 221–435, 23 pls.
Clark, H.L. (1933) A handbook of the littoral echinoderms of Porto Rico and the other West Indian Islands. Scientific Survey of Puerto Rico, 16, 1–147.
Clark, H.L. (1941) Reports on the scientific results of the Atlantis Expedition to the West Indies, under the joint auspices of the University of Havana and Harvard University. The echinoderms (other than holothurians). Memorias de la Sociedad Cubana de Historia Natural Felipe Poey, 15 (1), 1–154.
Cobb, J.C., Lawrence, J.M., Herrera, J.C. & Janies, D. (2019) A new species of Astropecten (Echinodermata: Asteroidea: Paxillosida: Astropectinidae) and a comparison of the Astropecten species from the Gulf of Mexico and the East Florida Shelf. Zootaxa, 4612 (3), 301–325.
https://doi.org/10.11646/zootaxa.4612.3.1
Collin, R., Venera-Pontón, D.E., Paulay, G. & Boyle, M.J. (2020) World Travelers: DNA Barcoding Unmasks the Origin of Cloning Asteroid Larvae from the Caribbean. Biology Bulletin, 239 (2), 73–79.
https://doi.org/10.1086/710796
Costa, P.A.S., Mincarone, M.M., Braga, A.C., Martins, A.S., Lavrado, H.P., Haimovici, M. & Falcão, A.P.C. (2015) Megafaunal communities along a depth gradient on the tropical Brazilian continental margin. Marine Biology Research, 11 (10), 1053–1064. https://doi.org/10.1080/17451000.2015.1062521
Costa-Neto, E.M. (1999) Healing with animals in Feira de Santana City, Bahia, Brazil. Journal of Ethnopharmacology, 65, 225–230.
https://doi.org/10.1016/S0378-8741(98)00158-5
Costa-Neto, E.M. & Marques, J.G.W. (2000) Faunistic resources used as medicines by artisanal fishermen from Siribinha Beach, State of Bahia, Brazil. Journal of Ethnobiology, 20 (1), 93–109.
Cunha, R., Tavares, M. & Mendonça, J.B. (2020) Asteroidea (Echinodermata) from shallow-waters of the remote oceanic archipelago Trindade and Martin Vaz, southeastern Atlantic, with taxonomic and zoogeographical notes. Zootaxa, 4742 (1), 31–56.
https://doi.org/10.11646/zootaxa.4742.1.2
Cutress, B.M. (1996) Changes in dermal ossicles during somatic growth in Caribbean littoral sea cucumbers (Echinoidermata: Holothuroidea: Aspidochirotida). Bulletin of Marine Science, 58 (1), 44–116.
Dall, W.H. (1902) Illustrations and descriptions of new, unfigured, or imperfectly known shells, chiefly American in the U. S. National Museum. United States National Museum, Proceedings, 24 (1264), 499–566.
https://doi.org/10.5479/si.00963801.24-1264.499
Danielssen, D.C. & Koren, J. (1884) Asteroidea. In: Mohn, H. & Sars, G.O. (Eds.), Den Norske Nordhavs Expedition, 1876-1878. Vol. 4. Grøndahl & Søns bogtrykkeri, Christiania, pp. 1–119, 15 pls.
Dartnall, A.J., Byrne, M., Collins, J. & Hart, M.W. (2003) A new viviparous species of asterinid (Echinodermata, Asteroidea, Asterinidae) and a new genus to accommodate the species of pantropical exiguoid sea stars. Zootaxa, 359, 1–14.
de Blainville, H.M. (1830) Zoophytes. In: Levrault, F.G. (Ed.), Dictionnaire des sciences naturelles, dans lequel on traitre méthodiquement des differéns êtres de la nature, considérés soit en eux-mêmes, d’après l’état actuel de nos connoissances, soit relativement a l’utlité qu’en peuvent retirer la médicine, l’agriculture, le commerce et les arts. Vol. 60. Le Normat, Paris, 548 pp., 68 pls.
De Léo, F.C. & Pires-Vanin, A.M.S. (2006) Benthic megafauna communities under the influence of the South Atlantic Central Water intrusion onto the Brazilian SE shelf: a comparison between an upwelling and a non-upwelling ecosystem. Journal of Marine Systems, 60, 268–284.
https://doi.org/10.1016/j.jmarsys.2006.02.002
Dehn, P.F. (1980a) Growth and reproduction in Luidia clathrata (Say) (Echinodermata: Asteroidea). Unpublished PhD Dissertation, University of South Florida, Tampa, Florida, 183 pp.
Dehn, P.F. (1980b) The annual reproductive cycle of two populations of Luidia clathrata (Asteroidea). 1. Organ indices and occurrence of larvae. In: Jangoux, M. (Ed.), Echinoderms: present and past. A.A. Balkema Press, Rotterdam, pp. 361–367.
https://doi.org/10.1201/9781003078913-66
Dilman, A.B. (2013) Asteroid fauna of the northern Mid-Atlantic Ridge: additional records. Marine Biology Research, 9 (5–6), 563–586.
https://doi.org/10.1080/17451000.2012.749993
Döderlein, L. & Hartmeyer, R. (1910) Westindische Seeigel und Seesterne. Zoologische Jahrbucher Supplement, 11 (2), 145–156.
Döderlein, L. (1916) Über die Gattung Oreaster und Venvandte. Zoologische Jahrbücher, Abteilung für Systematik, 40, 409–440.
Döderlein, L. (1917) Die Asteriden der Siboga-Expedition. I. Die Gattung Astropecten und ihre Stammesgeschichte. In: Brill, E.J. (Ed.), Siboga-Expeditie. Uitkomsten op zoölogisch, botanisch, ozeanographisch en geologisch gebied verzameld in Nederlandsch Oost-Indie 1899−1900 aan boord H.M. Buchhandlung und druckerei vormals E.J. Brill, Leiden, pp. 1–191 pp., 17 pls.
https://doi.org/10.5962/bhl.title.11319
d’Orbigny, A.D. (1839) Echinodermes et Polypiers. In: Webb, P.B. & Berthelot, S. (Eds.), Histoire Naturelle des Iles Canaries, Zoologie. Vol. 2. Béthune, Paris, pp. 148–149.
d’Orbigny, A.C.D. (1853) Mollusques. In: Sagra, R. de La (Ed.), Histoire physique, politique et naturelle de l’Île de Cuba. Bertrand, Paris, pp. 1–380, 28 pls.
Downey, M.E. (1970) Drachmaster bullisi new genus and species of Ophidiasteridae (Echinodermata: Asteroidea) with a key to the Caribbean species of the family. Proceedings of the Biological Society of Washington, 83 (6), 77–82.
Downey, M.E. (1973) Starfishes from the Caribbean and the Gulf of México. Smithsonian Contributions Zoology, 126, 1–158.
https://doi.org/10.5479/si.00810282.126
Düben, M.W.V. & Koren, J. (1845) Nya skandinaviska fiskar. Öfversigt af Kongl. Veterskaps-Akademiens Förhandlingar, 2 (1), 9–12.
Entrambasaguas, L. (2003) Estudio de la fauna de equinodermos del archipiélago de Cabo Verde: escalas de variabilidad espacial y factores que explican su distribución. Tesis de licenciatura no publicada. Universidad de Murcia, Murcia. [unknown pagination]
Entrambasaguas, L. (2008) Estudio fauniìstico y ecoloìgico de los equinodermos del archipieìlago de Cabo Verde. Tesis doctoral no publicada, Universidad de Murcia, Murcia, 315 pp.
Farmanfarmaian, A. (1966) The respiratory physiology of echinoderms. In: Boolootian, R.A. (Ed.), Physiology of Echinodermata. John Wiley & Sons, New York, pp. 245–265.
Fau, M. & Villier, L. (2018) Post-metamorphic ontogeny of Zoroaster fulgens Thomson, 1873 (Asteroidea, Forcipulatacea). Journal of Anatomy, 233 (5), 644–665.
https://doi.org/10.1111/joa.12881
Fau, M. & Villier, L. (2020) Comparative anatomy and phylogeny of the Forcipulatacea (Echinodermata: Asteroidea): insights from ossicle morphology. Zoological Journal of the Linnean Society, 189 (3), 921–952.
https://doi.org/10.1093/zoolinnean/zlz127
Fernandes, M.L.B., Tommasi, L.R. & Lima, E.J.B. (2002) Filo Echinodermata de Pernambuco. In: Tabarelli, M. & Silva, J.M.C. (Eds.), Diagnóstico da Biodiversidade de Pernambuco. Vol. 2. Massangana, Recife, pp. 405–427.
Fernandez, W.S., Dias, G.M., Majer, A., Delboni, C., Denadai, M.R. & Turra, A. (2017) Resource partitioning between sympatric starfish from tropical unconsolidated substrate: implications for coexistence and top-down control on benthic prey. Estuarine, Coastal and Shelf Science, 196, 141–149.
https://doi.org/10.1016/j.ecss.2017.06.039
Fisher, W.K. (1906) The starfishes of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23, 987–1130.
Fisher, W.K. (1908) Necessary changes in the nomenclature of starfishes. Smithsonian Miscellaneous collections (Quaterly), 52, 87–93.
Fisher, W.K. (1910) New genera of starfishes. Annals and Magazine of Natural History, Series 8, 5 (26), 171–173.
https://doi.org/10.1080/00222931008692747
Fisher, W.K. (1911) Asteroidea of the North Pacific and adjacent waters. 1. Phanerozonia and Spinulosa. Bulletin of the United States National Museum, 76, i–xiii + 1–420 pp., 122 pls.
https://doi.org/10.5479/si.03629236.76.i
Fisher, W.K. (1913) Four new genera and fifty-eight new species of starfishes from the Philippine Islands, Celebes, and the Moluccas. Proceedings of the US National Museum, 43, 599–648.
https://doi.org/10.5479/si.00963801.43-1944.599
Fisher, W.K. (1917) Notes on Asteroidea. Annals and Magazine of Natural History, Series 8, 20 (116), 166–172.
https://doi.org/10.1080/00222931709486985
Fisher, W.K. (1930) Asteroidea of the North Pacific and Adjacent Waters, Pt. 3: Forcipulatida. Bulletin of the United States National Museum, 76, 1–356, 93 pls.
https://doi.org/10.5479/si.03629236.76.3
Fisher, W.K. (1940) Asteroidea. Discovery Reports, 20, 69–306.
Fontanella, F.M. & Hopkins, T.S. (2003) Preliminary phylogeny of Echinaster (Othilia) from the Gulf of Mexico based on morphological characters (Echinodermata: Asteroidea). In: Féral, J.P. & David, B. (Eds.), Proceedings of the Sixth European Conference on Echinoderm Research, Banyuls-sur-Mer, France, 3–7 September 2001. Balkema, Rotterdam, pp. 91–95.
Fonseca, B.F. (2015) Contribuição da reprodução sexuada e assexuada e conectividade entre populações da estrela-do-mar Coscinasterias tenuispina (Lamarck, 1816) na costa do Estado do Rio de Janeiro. Unpublished Master Thesis, Museu Nacional, Universidade Federal do Rio de Janeiro, Rio de Janeiro, 64 pp.
Forbes, E. (1839) On the Asteriadae of the Irish Sea. Memoirs of the Wernerian Natural History Socierty of Edinburgh, 8, 114–129.
Forbes, E. (1841) A history of British Starfishes and other animals of the class Echinodermata. John Van Voorst, London, 267 pp.
https://doi.org/10.5962/bhl.title.2129
Franco, O.P., Patiño, G.S. & Ortiz, A.A. (2015) Actividad antibacteriana y antifúngica de la estrella de mar Oreaster reticulatus (Valvatida: Oreasteridae) y de los erizos de mar Mellita quinquiesperforata (Clypeasteroida: Mellitidae) y Diadema antillarum (Diadematoida: Diadematidae) del Caribe Colombiano. Revista de Biología Tropical, 63, 329–337.
Franz, D., Worley, E. & Merrill, A. (1981) Distribution patterns of common sea stars of the middle Atlantic continental shelf of the northwest Atlantic (Gulf of Maine to Cape Hatteras). The Biological Bulletin, 160, 394–418.
https://doi.org/10.2307/1540848
Gaymer, C.F. & Himmelman, J.H. (2008) A keystone predatory sea star in the intertidal zone is controlled by a higher-order predatory sea star in the subtidal zone. Marine Ecology Progress Series, 370, 143–153.
https://doi.org/10.3354/meps07663
GBIF Secretariat (2020) Coscinasterias tenuispina (Lamarck, 1816) in GBIF Backbone Taxonomy. Available from: https://www.gbif.org/species/2273832 (accessed 1 October 2020).
Gondim, A., Lacouth, P., Alonso, C. & Manso, C. (2008) Echinodermata da praia do Cabo Branco, JoaÞo Pessoa, Paraiìba, Brasil. Biota Neotropica, 8, 151–159.
https://doi.org/10.1590/S1676-06032008000200016
Gondim, A.I., Dias, T.L.P., Campos, F.F., Alonso, C. & Christoffersen, M.L. (2011) Macrofauna bêntica do Parque Estadual Marinho de Areia Vermelha, Cabedelo, Paraíba, Brasil. Biota Neotropica, 11 (2), 1–12.
https://doi.org/10.1590/S1676-06032011000200009
Gondim, A., Christoffersen, M. & Dias, T. (2014) Taxonomic guide and historical review of starfishes in northeastern Brazil (Echinodermata, Asteroidea). ZooKeys, 449, 1–56.
https://doi.org/10.3897/zookeys.449.6813
Gould, A.A. (1841) Report on the invertebrata of Massachusetts: comprising the Mollusca, Crustacea, Annelida, and Radiata. Folson, Wells & Thurston, Cambridge, xiii + 373 pp.
https://doi.org/10.5962/bhl.title.17646
Grant, R.E. (1841) Porifera. In: Bailliere, H. (Ed.), Outlines of comparative anatomy. Vol. 1. Baillière, London, pp. 5–9 + 310–313, pls. II–IV.
https://doi.org/10.5962/bhl.title.28330
Gray, J.E. (1840) A synopsis of the genera and species of the class Hypostoma (Asterias Linnaeus). Annals of the Magazine of Natural History, Series 1, 6 (37), 175–184 + 275–290.
https://doi.org/10.1080/03745484009443296
Gray, J.E. (1867) Notes on the Arrangement of Sponges, with the Descriptions of some New Genera. Proceedings of the Zoological Society of London, 1867 (2), 492–558, pls. XXVII–XXVIII.
Gray, I.E., Downey, M.E. & Cerame-Vivas, M.J. (1968) Sea-Stars of North Carolina. Fisheries Bulletin, 67 (1), 127–163, 40 figs.
Guerrazzi, M.C., Hadju, E., Duarte, L.F.L. & Morgado, E.H. (1999) Spongivory by the Brazilian starfish Echinaster brasiliensis. Memoirs of the Queensland Museum, Brisbane, 44, 214–214.
Guilherme, P.D. & Rosa, L.C. (2014) Seasonal variation in body size and diet of the sea star Astropecten marginatus (Paxillosida, Astropectinidae) off coast of Parana, Southern Brazil. Revista de Biologia Tropical, 62, 59–68.
https://doi.org/10.15517/rbt.v62i1.7888
Gurjão, L.M. & Lotufo, T.M.C. (2018) Native species exploited by marine aquarium trade in Brazil. Biota Neotropica, 18 (3), e20170387.
https://doi.org/10.1590/1676-0611-bn-2017-0387
Guzman, H.M. & Guevara, C.A. (2002) Annual reproductive cycle, spatial distribution, abundance, and size structure of Oreaster reticulatus (Echinodermata: Asteroidea) in Bocas del Toro, Panama. Marine Biology, 141, 1077–1084.
https://doi.org/10.1007/s00227-002-0898-2
Haeckel, E. (1878) Die Kometenform der Seesterne und der Generationswechsel der Echinodermum. Zeitschrift für Wissenschaftliche Zoologie, 30 (Supplement), 424–445.
Halpern, J.A. (1970a) A monographic revision of the goniasterid sea stars of the North Atlantic. Unpublished PhD Dissertation, University of Miami, Florida, 253 pp.
Halpern, J.A. (1970b) Goniasteridae (Echinodermata: Asteroidea) of the straits of Florida. Bulletin of Marine Science, 20 (1), 193–286.
Hendler, G., Muller, J.E., Pawson, D.L. & Kier, P.M. (1995) Sea Stars, Sea Urchins and Allies: Echinoderms of Florida and the Caribbean. Smithsonian Institution Press, Washington, 390 pp.
Hopkins, T.S., Fontanella, F.M. & Ventura, C.R.R. (2003) Morphological diagnosis of three Brazilian starfishes of the genus Echinaster (Othilia). In: Féral, J.P. & David, B. (Eds.), Proceedings of the Sixth European Conference on Echinoderm Research, Banyuls-Sur-Mer, France, 3–7 September 2001. Balkema, Rotterdam, pp. 97–103.
Hopkins, T.S. & Knott, K.E. (2010) The establishment of a neotype for Luidia clathrata (Say, 1825) and a new species within the genus Luidia (Asteroidea: Paxillosida: Luidiidae). In: Harris, L.G., Böttger, S.A., Walker, C.W. & Lesser, M.P. (Eds.), Proceedings of the 12th International Echinoderm Conference, 7–11 August 2006, Durham, New Hampshire. Taylor & Francis Group, London, pp. 207–212.
https://doi.org/10.1201/9780203869543-c32
Hyman, L.H. (1955) The Invertebrates: Echinodermata, The Coelomata Bilateria. McGraw Book Company INC, London, 763 pp.
Jangoux, M. (1982) Food and feeding mechanisms: Asteroidea. In: Jangoux, M. & Lawrence, J.M. (Eds.), Echinoderm Nutrition. A.A. Balkema, Rotterdam, pp. 117–159.
https://doi.org/10.1201/9781003078920-6
Jangoux, M. & Lambert, A. (1987) Étude comparative des pédicellaires des astérides (échinodermes). Bulletin de la Société Scientifique et Naturaliste de l’Ouest de France, 1987, 47– 56.
Jangoux, M. & Aziz, A. (1988) Les asterides (Echinodermata) récoltes autour de l’île de la Réunion par le N.O. “Marion-Dufresne” en 1982. Bulletin du Museum National D’Histoire Naturelle, Zoologie, 4 (10), 631–650.
Janies, D., Hernández-Díaz, Y.Q., Solís-Marín, F.A., Lopez, K., Alexandrov, B., Galac, M., Herrera, J., Cobb, J., Ebert, T.A. & Bosch. I. (2019) Discovery of adults linked to cloning oceanic starfish larvae (Oreaster: Asteroidea: Echinodermata). The Biological Bulletin, 236, 174–185.
https://doi.org/10.1086/703233
John, D.D. (1948) Notes on the asteroids in the British Museum (Natural History). I. The species of Astropecten. Novitates Zoologicae, 42, 485–508.
Kordas, R.L., Harley, C.D.G. & O’Connor, M.I. (2011) Community ecology in warming world: the influence of temperature on interspecific interactions in marine systems. Journal of Experimental Biology and Ecology, 400, 218–226.
https://doi.org/10.1016/j.jembe.2011.02.029
Koukouras, A., Sinis, A.I., Bobori, D., Kazantzidis, S. & Kitsos, M.S. (2007) The echinoderm (Deuterostomia) fauna of the Aegean Sea, and comparison with those of the neighboring seas. Journal of Biological Research, 7, 67–92.
Lamarck, J.B. (1816) Histoire naturelle des animaux sans vertebres. Meline, Bruxelles, 568 pp.
Lambert, P. (2000) Sea stars of British Columbia, Southeast Alaska and Puget Sound. University of British Columbia Press, Vancouver, British Columbia, 186 pp.
Lawrence, J.M. (1973) Level, content, and caloric equivalents of the lipid, carbohydrate, and protein in the body components of Luidia clathrata (Echinodermata: Asteroidea: Platyasterida) in Tampa Bay. Journal of Experimental Marine Biology and Ecology, 11, 263–274.
https://doi.org/10.1016/0022-0981(73)90026-9
Lawrence, J.M. & Cobb, J. (2017) Validation of Astropecten jarli Madsen, 1950 and implications for A. cingulatus Sladen, 1883 (Paxillosida: Astropectinidae). Zootaxa, 4269 (1), 101–114.
https://doi.org/10.11646/zootaxa.4269.1.4
Lawrence, J.M., Erwin, K. & Turner, R.L. (1974) Stomach contents of Luidia clathrata (Asteroidea). Florida Scientist, 37, 8.
Lawrence, J.M., Cobb, J.C. & Herrera, J.C. (2018a) Synonymy of Astropecten nitidus Verrill, 1915 with Astropecten cingulatus Sladen, 1883 and Astropecten comptus Verrill, 1915 with Astropecten articulatus (Say, 1825). Bulletin of the Peabody Museum of Natural History, 59 (2), 127–157.
https://doi.org/10.3374/014.059.0203
Lawrence, J.M., Cobb, J.C., Herrera, J.C., Durán-González, A. & Solís-Marín, F.A. (2018b) Morphological comparison of Astropecten cingulatus and a new species of Astropecten (Paxillosida, Astropectinidae) from the Gulf of Mexico. Zootaxa, 4407 (1), 86–100.
https://doi.org/10.11646/zootaxa.4407.1.5
Lima, E.J.B. & Fernandes, M.L.B. (2009) Diversidade de equinodermos (Echinodermata) no estado de Pernambuco (Brasil). Revista Brasileira de Zoociências, 11, 55–63.
Lima, R.J.P. (2018) Uso de Zooterápicos na Zona Rural do Município de Jaçanã, Semiárido Potiguar. Unpublished Honors Thesis, Instituto Federal de Educação, Ciência e Tecnologia da Paraíba, João Pessoa, Paraíba, 34 pp.
Lima-Verde, J.S. (1969) Primeira contribuição ao inventário dos equinodermas do Nordeste brasileiro. Arquivos de Ciências do Mar, 9 (1), 9–13.
Linnaeus, C. (1758) Systema Naturae. Impensis Direct, Laurentii Salvii, Holmiae, 824 pp.
Lopes, E.M. (2010) Variações morfológicas, ontogenia e compatibilidade gamética entre os morfotipos de Echinaster (Othilia) brasiliensis Muller & Troschel, 1840. Unpublished Master Thesis, Universidade Federal do Rio de Janeiro, Rio de Janeiro, 78 pp.
Lopes, E.M. & Ventura, C.R.R. (2016) Development of the sea star Echinaster (Othilia) brasiliensis, with inference on the evolution of development and skeletal plates in Asteroidea. Biology Bulletin, 230, 25–35.
https://doi.org/10.1086/BBLv230n1p25
Lopes, E.M., Perez-Portela, R., Paiva, P.C. & Ventura, R.C.C. (2016) The molecular phylogeny of the sea star Echinaster (Asteroidea: Echinasteridae) provides insights for genus taxonomy. Invertebrate Biology, 135 (3), 235–244.
https://doi.org/10.1111/ivb.12135
Lütken, C.F. (1860) Bidrag til kundskab om de ved kysterne af Mellenog Sydde ved kysterne af Mellenog Sydundskab. Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjobenhavn, 1859, 25–97.
Lütken, C. (1871) Fortasatte kritiske og beskrivende Bidrag til Kundskab om Sostjernerne (Asteriderne). Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, 1871, 227–304.
Madeira, P., Kroh, A., Cordeiro, R., Frias, D.E., Martins, A.M. & Ávila, S.P. (2019) The Echinoderm Fauna of the Azores (NE Atlantic Ocean). Zootaxa, 4639 (1), 1–231. https://doi.org/10.11646/zootaxa.4639.1
Madsen, F.J. (1950) The Echinoderms Collected by the Atlantide Expedition 1945-46. I. Asteroidea. With Remarks on Other Sea-Stars from Tropical and Northern West Africa. Atlantide Reports, 1, 167–222.
Magalhães, W.F., Martins, L.R. & Alves, O.F.S. (2005) Inventário dos Echinodermata do estado da Bahia. Brazilian Journal of Aquatic Science and Technology, 9 (1), 61–65.
https://doi.org/10.14210/bjast.v9n1.p61-65
Magris, R.A. & Destro, G.F.G. (2010) Predictive modeling of suitable habitats for threatened marine invertebrates and implications for conservation assessment in Brazil. Brazilian Journal of Oceanography, 58 (4), 57–68.
https://doi.org/10.1590/S1679-87592010000800008
Magris, R.A & Giarrizzo, T. (2020) Mysterious oil spill in the Atlantic Ocean threatens marine biodiversity and local people in Brazil. Marine Pollution Bulletin, 153, 1–3. https://doi.org/10.1016/j.marpolbul.2020.110961
Mah, C.L. (2018) New genera, species and occurrence records of Goniasteridae (Asteroidea; Echinodermata) from the Indian Ocean. Zootaxa, 4539 (1), 1–116.
https://doi.org/10.11646/zootaxa.4539.1.1
Mah, C.L. (2020a) World Asteroidea Database. Available from: http://www.marinespecies.org/asteroidea/ (accessed 30 April 2020)
Mah, C.L. (2020b) New species, occurrence records and observations of predation by deep-sea Asteroidea (Echinodermata) from the North Atlantic by NOAA ship Okeanos Explorer. Zootaxa, 4766 (2), 201–260.
https://doi.org/10.11646/zootaxa.4766.2.1
Mah, C.L. (2020c) Mystery of the Mud Star Ctenodiscus revealed. Available from: http://echinoblog.blogspot.com/2010/03/mystery-of-mud-star-ctenodiscus-revealed.html (accessed 1 June 2020).
Mah, C. & Foltz, D. (2011) Molecular phylogeny of the Valvatacea (Asteroidea: Echinodermata). Zoological Journal of the Linnean Society, 161 (4), 769–788.
https://doi.org/10.1111/j.1096-3642.2010.00659.x
Mah, C. & Blake, D. (2012) Global diversity and phylogeny of the Asteroidea (Echinodermata). Plos One, 7, 1–22.
https://doi.org/10.1371/journal.pone.0035644
Mah, C.L. & Foltz, D.W. (2014) New Taxa and Taxonomic Revisions to the Poraniidae (Valvatacea: Asteroidea) with Comments on Feeding Biology. Zootaxa, 3795 (3), 327–372.
https://doi.org/10.11646/zootaxa.3795.3.7
Mah, C.L. & Fujita, T. (2020) New species and occurrence records of Japanese Solasteridae and Ganeriidae including a new species of Paralophaster from the North Pacific with an overview of Hyalinothrix. Zootaxa, 4750 (1), 67–100.
https://doi.org/10.11646/zootaxa.4750.1.4
Manso, C.L.C. (1989) Os Echinodermata da plataforma continental interna entre Cabo Frio e Saquarema, Rio de Janeiro, Brasil. Revista Brasileira de Biologia, 49, 355–359.
Manso, C.L.C. (1993) Ofiuróides da Plataforma Brasileira. Parte 11: Norte do Estado do Rio de Janeiro, Estado do Espírito Santo, Sul do Estado da Bahia e Bancos Royal Charlott, Hotspur e Davis (Echinodermata: Ophiuroidea). Revista Brasileira de Biologia, 53 (2), 189–195.
https://doi.org/10.1590/S1676-06032008000300017
Manso, C.L.C., Alves, O.F.S. & Martins, L.R. (2008) Echinodermata da Baía de Todos os Santos e da Baía de Aratu (Bahia, Brasil). Biota Neotropica, 8 (3), 179–196.
https://doi.org/10.1590/S1676-06032008000300017
Martins, L. (2019) Ontogenetic variation of the ossicles in Pseudothyone belli (Ludwig, 1887) with a description of a new Sclerodactylidae (Echinodermata: Holothuroidea: Dendrochirotida) from Southwestern Atlantic. Marine Biology Research, 15 (7), 434–443. https://doi.org/10.1080/17451000.2019.1662051
Martins, I.X. & Queiroz, A.C.M. (2006) Equinodermos do litoral do estado do Ceará. In: Matthews-Cascon, H. & Lotufo, T.M.C. (Eds.), Biota marinha da costa oeste do Ceará. Ministério do Meio Ambiente, Série Biodiversidade, Brasília, pp. 200–219.
Martins, L., Souto, C., Magalhães, W.F., Alves, O.F.S., Rosa, I.L. & Sampaio, C.L.S. (2012) Echinoderm harvesting in Todos-os-Santos Bay, Bahia State, Brazil: the aquarium trade. Sitientibus série Ciências Biológicas, 12 (1), 53–59.
https://doi.org/10.13102/scb123
Martynov, A., Ishida, Y., Irimura, S., Tajiri, R., O’Hara, T. & Fujita, T. (2015) When ontogeny matters: a new Japanese species of brittle star illustrates the importance of considering both adult and juvenile characters in taxonomic practice. PLoS ONE, 10 (10), e0139463.
https://doi.org/10.1371/journal.pone.0139463
McClintock, J.B. (1994) Trophic biology of Antarctic shallow-water echinoderms. Marine Ecology Progress Series, 111, 191–202.
https://doi.org/10.3354/meps111191
McKnight, D.G. (2006) The marine fauna of New Zealand, Echinodermata: Asteroidea (Sea-stars). 3. Orders Velatida, Spinulosida, Forcipulatida, Brisingida with addenda to Paxillosida, Valvatida. NIWA Biodiversity Memoir, Wellington, 187 pp.
Mecho, A., Billett, D.S.M., Ramirez-Liodra, E., Aguzzi, J., Tyler, P.A. & Company, J.B. (2014) First records, rediscovery and compilation of deep-sea echinoderms in the middle and lower continental slope of the Mediterranean Sea. Scientia Marina, 78 (2), 281–302.
https://doi.org/10.3989/scimar.03983.30C
Micael, J. & Costa A.C. (2010) Echinodermata. In: Borges, P.A.V., Costa, A., Cunha, R., Gabriel, R., Gonçalves, V., Martins, A.F., Melo, I., Parente, M., Raposeiro, P., Rodrigues, P., Santos, R.S., Silva, L., Vieira, P. & Vieira, V. (Eds.), A list of the terrestrial and marine biota from the Azores. Princípia, Oeiras, pp. 321–323.
Micael, J., Alves, M.J., Jones, M.B. & Costa, A.C. (2012) Diversity of shallow-water asteroids (Echinodermata) in the Azorean Archipelago. Marine Biodiversity Records, 5 (3), 1–10.
https://doi.org/10.1017/S1755267211000534
Miranda, A.L.S., Lima, M.L.F., Sovierzoski, H.H. & Correia, M.D. (2012) Inventory of the Echinodermata collection from the Universidade Federal de Alagoas. Biota Neotropica, 12 (2), 135–146.
https://doi.org/10.1590/S1676-06032012000200014
Miranda, A.P.S. (2018) Diversidade e distribuição de equinodermos da Plataforma Continental Amazônica. Unpublished Honors Thesis, Universidade Federal do Pará, Belém, 62 pp.
Mironov, A.N., Dilman, A.B., Vladychenskaya, I.P. & Petrov, N.B. (2016) Adaptive strategy of the porcellanasterid sea stars. Biology Bulletin, 43 (6), 503–516.
https://doi.org/10.1134/S106235901606011X
MMA (Ministério do Meio Ambiente) (2018) Livro Vermelho da Fauna Brasileira Ameaçada de Extinção: Invertebrados. ICMBio/MMA, Brasília, 727 pp.
Möbius, K. (1859) Neue Seesterne des Hamburger und Kieler Museums. Abhandlungen und Verhandlungen Naturwissenschaftlicher Verein in Hamburg, 4 (2), 1–14.
https://doi.org/10.5962/bhl.title.11830
Monteiro, A.M.G. & Pardo, E.V. (1991) Dieta alimentar de Astropecten marginatus e Luidia senegalensis (Echinodermata: Asteroidea). Revista Brasileira de Biologia, 54, 49–54.
Mortensen, T. (1933) The Echinoderms of St. Helena (Asteroidea and Ophiuroidea). Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjobenhavn, 93, 401–472.
Müller, J. & Troschel, F.H. (1842) System der Asteriden. Friedrich Viewig und Sohn, Braunschweig, 134 pp.
Nardo, J.D. (1834) De Asteriis. Isis von Oken, Encyclopaedische Zeitun, 27, 716–717.
Netto, L.F. (2006) Echinodermata do Canal de São Sebastião, São Sebastião. Unpublished Master Thesis, Universidade de São Paulo, São Paulo, 201 pp.
Nisperuza, P.C., Padilla, C.J. & Quirós, R.J. (2016) Current status of the study of the caribbean cordobes echinoderms, Colombia. Revista Colombiana de Ciencia Animal, 8 (1), 112–119.
https://doi.org/10.24188/recia.v8.n1.2016.234
Nobre, C. & Campos-Creasey, L.S. (2000) Efeito da salinidade no desenvolvimento larval de Echinaster echinophorus (Lamarck 1816) (Echinodermata: Asteroidea). Brazilian Journal of Aquatic Science and Technology, 4, 61–68.
https://doi.org/10.14210/bjast.v4n1.p61-68
Nomura, H. & Fausto Filho, J. (1966) Shrimp survey in coastal and offshore waters of northeastern Brazil. Arquivos da Estação de Biologia Marinha da Universidade Federal do Ceará, 6, 15–29.
Nunes, J.M.C., Santos, A.C.C. & Santana, L.C. (2005) Novas ocorrências de algas marinhas bentônicas para o estado da Bahia, Brasil. Iheringia, 60, 99–106.
Nunes, T.B. (1975) Echinoderma da Baía de Aratu (Bahia, Brasil). Memories of the second Latin American Symposium of Oceanographic Biology, Universidad de Oriente, 1975, 179–189.
Oliveira, J.P., Oliveira, J. & Manso, C.L.C. (2010) Inventário da coleção de equinodermos do LABIMAR, Campus Prof°. Alberto Carvalho, Universidade Federal de Sergipe. Scientia Plena, 6, 1–14.
O’Loughlin, P.M. (2002) New genus and species of southern Australian and Pacific Asterinidae (Echinodermata, Asteroidea). Memoirs of Museum Victoria, 59 (2), 277–296.
https://doi.org/10.24199/j.mmv.2002.59.2
O’Loughlin, P.M. & Waters, J.M. (2004) A molecular and morphological revision of genera of Asterinidae (Echinodermata: Asteroidea). Memoirs of Museum Victoria, 61 (1), 1–40.
https://doi.org/10.24199/j.mmv.2004.61.1
Parra, M.G., Fidalgo, L.M., Martinez, J.M., Alvarez, A.M. & Iglesias, O.V. (2010) Leishmanicidal activity of Echinaster (Othilia) echinophorus crude extract. Revista do Instituto de Medicina Tropical de São Paulo, 52, 89–93.
https://doi.org/10.1590/S0036-46652010000200005
Patrizzi, N. & Dobrovolski, R. (2018) Integrating climate change and human impacts into marine spatial planning: a case study of threatened starfish species in Brazil. Ocean & Coastal Management, 161, 177–188.
https://doi.org/10.1016/j.ocecoaman.2018.05.003
Pawson, D.L. (1978) The echinoderm fauna of Ascension Island, South Atlantic Ocean, Smithsonian. Contributions Marine Sciences, 2, 1–31.
https://doi.org/10.5479/si.01960768.2.1
Pawson, D.L. (2007a) Phylum Echinodermata. Zootaxa, 1668 (1), 749–764.
https://doi.org/10.11646/zootaxa.1668.1.31
Pawson, D.L. (2007b) Narcissia ahearnae, a new species of sea star from the Western Atlantic (Echinodermata: Asteroidea: Valvatida). Zootaxa, 1386 (1), 53–58. https://doi.org/10.11646/zootaxa.1668.1.31
Pawson, D.L., Vance, D.J., Messing, C.G., Solis-Marin, F.A. & Mah, C.L. (2009) Echinodermata of the Gulf of Mexico. In: Felder, D.L. & Camp, D.K. (Eds.), Gulf of Mexico–Origins, Waters, and Biota. Texas A&M University Press, College Station, Texas, pp. 1177–1204.
Pazzoto, C.E.M., Ventura, C.R.R. & Silva, E.P. (2010) Genetic contribution of sexual and asexual reproduction to the recruitment of a sexually unbalanced population of Coscinasterias tenuispina (Echinodermata: Asteroidea) in Rio de Janeiro, Brazil. In: Harris, L.G., Boettger, S.A., Walker, C.W. & Lesser, M.P. (Eds.), Proceedings of the 12th International Echinoderm Conference, Durham, New Hampshire, 7–11 August 2006. CRC Press, Boca Raton, Florida, pp. 473–478.
https://doi.org/10.1201/9780203869543-c73
Pelaes, M.A.L. (2008) Composição, distribuição espacial e temporal das espécies de Echinodermata na plataforma rasa do sul da Bahia num trecho entre Itacaré e Canavieiras. Unpublished Master Thesis, Universidade Estadual de Santa Cruz, Ilhéus, Bahia, 97 pp.
Pechaszadeh, P.E. & Lera, M.E. (1983) Alimentación de tres especies tropicles de Luidia (Echinodermata: Asteroidea) en Golfo Triste, Venezuela. Caribbean Journal of Science, 19, 1–6.
Pérez-Ruzafa, A., Entrambasaguas, L. & Bacallado, J.J. (1999) Fauna de equinodermos (Echinodermata) de los fondos rocosos infralitorales del archipiélago de Cabo Verde. Revista de la Academia Canaria de Ciencias, 11 (3–4), 43–62.
Pérez-Ruzafa, A., Entrambasaguas, L., Espejo, C., Marcos, C. & Bacallado, J.J. (2002) Fau