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Type: Monograph
Published: 2021-05-07
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Annotated world checklist of the Trombiculidae and Leeuwenhoekiidae (1758–2021) (Acari: Trombiculoidea), with notes on nomenclature, taxonomy, and distribution

Nebraska Department of Agriculture, 301 Centennial Mall South, Lincoln, Nebraska 68509 USA.
Walter Reed Biosystematics Unit, Department of Entomology, Smithsonian Institution, MSC, MRC 534, 4210 Silver Hill Road, Suitland, Maryland 20746 USA, and Department of Entomology, Walter Reed Army Institute of Research, 503 Robert Grant Avenue, Silver Spring, Maryland 20910, USA.
Walter Reed Biosystematics Unit, Department of Entomology, Smithsonian Institution, MSC, MRC 534, 4210 Silver Hill Road, Suitland, Maryland 20746 USA, and Department of Entomology, Walter Reed Army Institute of Research, 503 Robert Grant Avenue, Silver Spring, Maryland 20910, USA. Mailing address: 204 Draymore Way, Cary, North Carolina 27519, USA
Acari checklist chigger mites Trombiculoidea taxonomy biogeography

Abstract

The superfamily Trombiculoidea is a large and diverse group of acarines that comprises six families; of these, the families Trombiculidae and Leeuwenhoekiidae are characterized by larvae commonly known as chiggers that are parasites of terrestrial vertebrates, including humans, and some species are of medical importance as vectors of chigger-borne rickettsiosis (scrub typhus), caused by the rickettsia Orientia tsutsugamushi.  This paper presents an annotated checklist of 3,013 generally accepted chigger species, together with their distribution by zoogeographic region, and a non-comprehensive list of synonyms.  A total of 58 new combinations are proposed by transferring species to different genera, treating some subgenera as genera, or updating current generic names. The checklist updates earlier catalogs of the world chigger fauna, but because the literature on chigger taxonomy is voluminous and scattered, similar, regional monographs were utilized in constructing the classificatory foundation.  This list may not contain every species, synonym or taxonomic rearrangement that has been published, but it reflects the most recent arrangement of chigger taxa.  Distribution records are primarily drawn from collection sites listed in the original species descriptions, together with subsequently published geographical records.

 

References

  1. Abo-Taka, S.M. & Comroy, H.L. (1995) A new species of Euschoengastia (Acari: Trombiculidae) from a domestic dog in the U.S.A. Alexandria Journal of Agricultural Research, 40, 245–250.

    Abonnenc, E. (1955) Sur un nouveau Trombiculidae (Acarien): Euschoengastia rouchoni, parasite d’un phacochère. Annales de Parasitologie Humaine et Comparée, 30, 219–224.

    Abonnenc, E. & Taufflieb, R. (1957a) Deux trombiculidés nouveaux de la région du Tchad, Euschoengastia pilosa n. sp. et Trombicula youhensis n. sp., présence de Schoengastia radfordi Jadin et Vercammen-Grandjean, 1954. Bulletin de la Société de Pathologie Exotique, 50, 556–567.

    Abonnenc, E. & Taufflieb, R. (1957b) Sur un nouveau trombiculide larvaire de la région du Tchad: Euschoengastia (Helenicula) vercammengrandjeani n. sp. Bulletin de la Société de Pathologie Exotique, 50, 86–89.

    *Abou-Taka, C.M. (1985) Schoutedenichia infrequens sp.n. a new species of trombiculid mites from Tadzhikistan (Trombiculidae). Parazitologiya, 19, 245–247. (in Russian) (cited in Kudryashova 1998)

    Abussalam, M. (1939) A new trombiculid larva parasitic on the house rat (Rattus rattus). Indian Journal of Entomology, 1, 83–86.

    Ah, H.S. (1960) Shunsennia gracilis a new species of trombiculid mite from Korea (Acarina, Trombiculidae). Parasitology, 50, 577–580.

    Ah, H.S. (1964) New intranasal chiggers of the genus Microtrombicula from some bats in Korea (Acarina: Trombiculidae). Journal of Medical Entomology, 1, 186–191.

    https://doi.org/10.1093/jmedent/1.2.186

    Ahamad, M., Ibrahim, H., Bujang, M.K., Mohd Sah, S.A., Mohamed, N., Mohd Nor, S., Ahmad, A.H. & Ho, T.Z. (2013) A survey of acarine ectoparasites of bats (Chiroptera) in Malaysia. Journal of Medical Entomology, 50, 140–146.

    https://doi.org/10.1603/ME11240

    *Allred, D.M. (1957) Mites found on the mice genus Peromyscus in Utah. Great Basin Naturalist, 17, 95–102. (cited in Kudryashova 1998)

    *Allred, D.M. and Beck, D.E. (1953) A new species of Acomatacarus (Acarina, Trombiculidae) from Utah. Great Basin Naturalist, 13, 87–90. (cited in Walters et al. 2011)

    *Allred, D.M. and Beck, D.E. (1966) Mites of Utah mammals. Brigham Young University Science Bulletin, Biological Series, 8, 1–122. (cited in Walters et al. 2011)

    Amanguliev, A. (1973) New species of chiggers from Eltonella Audy, 1956 (Acariformes: Trombiculidae). Izvestiia Akademii Nauk Kirgizskoi SSR, Khimiko-Tekhnologicheskie i Biologicheskie Nauki (Bulletin of the Academy of Sciences of Kirgizskaya SSR, Series of Chemical-Tekhnological and Biological Sciences), 4, 89–92. (in Russian)

    *Amanguliev, A. (1977) Materials on trombiculid mites from some gerbil species. In: Ecological and medical significance of gerbils of the USSR fauna. Moscow, pp. 241–242. (in Russian) (cited in Kudryashova 1998)

    *Amanguliev, A. (1984) New species of Neotrombicula kugitangica Amanguliyev sp.nov. of Neotrombicula Hirst, 1925 (Acariformes, Trombiculidae) genus from Turkmenistan. Izvestiia Akademii Nauk Turkmenskoy SSR, Seriia Biologicheskikh Nauk (Bulletin of the Academy of Sciences of Turkmenskaya SSR, Series of Chemical-Tekhnological and Biological Sciences), 1984 (1), 62–64. (in Russian) (in Zoological Record; cited in Kudryashova, 1998)

    Amanguliev, A. (1996) New species and findings of Acariformes, Trombiculidae in Turkmenistan. Izvestiia Akademii Nauk Turkmenskoy SSR, Seriia Biologicheskikh Nauk (Bulletin of the Academy of Sciences of Turkmenskaya SSR, Series of Chemical-Tekhnological and Biological Sciences), 2, 60–62. (in Russian with English summary)

    Amanguliev, A., Shammakov, S. & Atayev, C. (1972) New species of mites of Odontacarus (Acariformes, Leeuwenhoekiidae). Izvestiia Akademii Nauk Turkmenskoy SSR, Seriia Biologicheskikh Nauk (Bulletin of the Academy of Sciences of Turkmenskaya SSR, Series of Chemical-Tekhnological and Biological Sciences), 5, 83–85. (in Russian with English summary)

    *André, M (1926) Notes complémentaires et synonymiques concernant divers Thrombidiidae. Bulletin du Muséum National d’Histoire Naturelle, Paris, 2e Série, 32, 375–377. (cited in Wharton and Fuller 1952)

    *André, M. (1928a) Observations biologiques sur la larve du Thrombicula autumnalis Shaw. Bulletin de la Société Zoologique de France, 53, 368–370. (cited in Wharton and Fuller 1952)

    *André, M. (1928b) Note sur le genre Thrombicula Berlese, 1905. Bulletin du Muséum National d’Histoire Naturelle, Paris, 2e Série, 34, 208–211. (cited in Wharton and Fuller 1952)

    André, M. (1929) Nouvelle forme larvaire de Thrombicula parasite sur un saurien de Palestine. Bulletin du Muséum National d’Histoire Naturelle, Paris, 2e Série, 1, 401–405.

    *André, M. (1930a) Contribution à l’étude d’un acarien: le Thrombicula autumnalis Shaw. Mémoires de la Société Zoologique de France, 29, 39–138. (cited in Wharton and Fuller 1952)

    *André, M. (1930b) Sur une larve d’acarien parasite de l’homme et des animaux en Uruguay, appartenant au genre Thrombicula Berlese. Annales de Parasitologie Humaine et Comparée, 8, 355–361. (cited in Wharton and Fuller 1952)

    André, M. (1932a) Note sur un acarien Yougo-Slavie appartenant au genre Thrombicula Berlese, 1905. Compte Rendu, de l’Association Française pour l’Avancement des Sciences, Bruxelles 1932, pp. 273–276.

    André, M. (1932b) Un acarien nouveau du Sud Algerien appartenant au genre Thrombicula (T. algerica n. sp.). Bulletin de la Société Zoologique de France, 57, 284–288.

    *André, M. (1933) Thrombicula autumnalis Shaw ou T. russica Oud. (Acariens). Bulletin de la Société Entomologique de France, 38, 154–156. (cited in Wharton and Fuller 1952)

    André, M. (1937) Note sur différents Thrombicula adultes trouvés en France avec description d’une espèce nouvelle. Bulletin du Muséum National d’Histoire Naturelle, Paris, 2e Série, 9, 313–318.

    *André, M. (1938) Contribution à l’étude des Trombidiidae de l’Afrique occidentale. Bulletin de la Société Zoologique de France, 63, 224–236. (cited in Wharton and Fuller 1952)

    André, M. (1943) Une espèce nouvelle de Leeuwenhoekia (Acarien) parasite de scorpions. Bulletin du Muséum National d’Histoire Naturelle, Paris, 2e Série, 15, 294–298.

    André, M. (1945) Un Thrombicula nouveau recueille en Afrique-Orientale (T. scapulosa n. sp.). Bulletin du Muséum National d’Histoire Naturelle, Paris, 2e Série, 17, 472–475.

    André, M. (1946) Sur une nouvelle forme larvaire de Neoschoengastia (Acarien) parasite de Meriones shawi. Bulletin du Muséum National d’Histoire Naturelle, Paris, 2e Série, 18, 162–164.

    André, M. (1949) Sur une novelle forme larvaire de Schoengastia (Acarien) parasite d’un caméléon malgache. Bulletin du Muséum National d’Histoire Naturelle, Paris, 2e Série, 21, 537–539.

    André, M. (1950) Une nouvelle forme larvaire de Thrombicula d’Afrique Equatoriale. Bulletin du Muséum National d’Histoire Naturelle, Paris, 2e Série, 22, 577–581.

    André, M. (1951a) Deux nouvelles espèces de Thrombicula d’Afrique Equatoriale. Bulletin de la Société de Pathologie Exotique, 44, 215–221.

    *André, M. (1951b) Nouvelle espèce de Schoengastiella (Acarien) d’Afrique Equatoriale. Bulletin du Muséum National d’Histoire Naturelle, Paris, 2e Série, 23, 368–371. (cited in Stekolnikov 2018b)

    André, M. (1951c) Sur une forme larvaire de Schoengastia (Acarien) parasite de petits mammifères en Oubangui-Chari. Bulletin du Museum National d’Histoire Naturelle, Paris, 2e Série, 23, 372–374.

    André, M. (1954) Description d’une nouvelle larve d’Ascoschoengastia (Acarien) parasite de rats en Indochine. Bulletin du Muséum National d’Histoire Naturelle, Paris, 2e Série, 26, 200–203.

    *André, M. (1958) Acariens thrombidions (adultes) de l’Angola. Publicações Culturais da Companhia de Diamantes de Angola, 35, 1–124. (cited in Stekolnikov 2018b)

    *Anonymous (1944) Scrub typhus fever (tsutsugamushi disease). War Department Technical Bulletin, Medical. (cited in Wharton and Fuller 1952)

    Asanuma, K. (1957a) Description of a new chigger mite, Makiena smadeli n. sp., from a Japanese hedge-sparrow, Prunella rubida rubida. Japanese Journal of Sanitary Zoology, 8, 231–234. (in Japanese with English summary)

    Asanuma, K. (1957b) A new chigger mite, Neoschoengastia hullinghorsti sp. nov., from the long-billed ringed plover, Charadrius placidus Gray, in Saitama Prefecture, Japan. Miscellaneous Reports of the Research Institute for Natural Resources, (45), 69–73.

    Asanuma, K. (1959a) A new subgenus and two new species of trombiculid mites, Trombicula (Tsutsugamushia) blumbergi, n. subgen. et sp., and Neoschoengastia baylissi n. sp. from the Japanese birds. Miscellaneous Reports of the Research Institute for Natural Resources, (51), 34–43.

    Asanuma, K. (1959b) Two new species of chigger mites, Trombicula (Leptotrombidium) hazatoi n. sp. and Ascoschoengastia (Laurentella) mcninchi, n. sp. from Japan. Japanese Journal of Hygiene, 14, 534–539. (in English and Japanese)

    Asanuma, K. & Saito, Y. (1957) Notes on chigger mites found on the Japanese hares, with description of Gahrliepia (Walchia) masoni n. sp. Japanese Journal of Sanitary Zoology, 8, 223–230. (in Japanese)

    Asanuma, K., Suzuki, M. & Fujikura, T. (1959) Trombicula (Leptotrombidium) kitaokai n. sp. from a vole, Microtus montebelli, in Fukushima Prefecture. Japanese Journal of Sanitary Zoology, 10, 136–138. (in Japanese with English summary)

    *Audy, J.R. (1947) Scrub typhus as a study in ecology. Nature (London), 159, 295–296. (cited in Wharton and Fuller 1952)

    *Audy, J.R. (1949) A summary topographical account of scrub typhus 1908-1946. Bulletin from the Institute for Medical Research, Federation of Malaya, 1, 82 pp. (cited in Wharton and Fuller 1952)

    *Audy, J.R. (1950) Occurrence of trombiculid mites on arthropods. Nature, 165, 193. (cited in Wharton and Fuller 1952)

    Audy, J.R. (1952) Trombiculid mites infesting bats in Malaya, with descriptions of three new species. Bulletin of the Raffles Museum, (24), 132–159.

    Audy, J.R. (1954) Malaysian Parasites IX. Notes on the taxonomy of trombiculid mites, with description of a new subgenus. Studies from the Institute for Medical Research, Federation of Malaya, (26), 123–170.

    Audy, J.R. (1956a) Laurentella, a new subgenus of trombiculid mites, with notes on biology and medical importance. Bulletin of the Raffles Museum, (28), 5–26.

    Audy, J.R. (1956b) Malayan trombiculid mites. 1. Schoutedenichia vercammeni n. sp. with a note on the genus. Bulletin of the Raffles Museum, (28), 81–85.

    *Audy, J.R. (1956c) Malayan trombiculid mites. 2. Naked-eye observations on attached chiggers, with a simple checklist of Malayan species. Bulletin of the Raffles Museum, (28), 86–101. (cited in Audy et al. 1965)

    Audy, J.R. (1956d) Trombiculid mites infesting birds, reptiles, and arthropods in Malaya, with a taxonomic revision, and description of a new genus, two new subgenera, and six new species. Bulletin of the Raffles Museum, (28), 27–80.

    Audy, J.R. (1957) A checklist of trombiculid mites of the Oriental and Australasian Regions. Parasitology, 47, 217–294.

    Audy, J.R. (1968) Red mites and typhus. The Athlone Press, University of London. x + 191 pp.

    Audy, J.R. and Domrow, R. (1957) Malaysian parasites XXIII. A revision of Oriental and Australasian species of Euschoengastia Ewing, including Walchiella Fuller, and description of a new subgenus, Derrickiella (Acarina, Trombiculidae). Studies from the Institute for Medical Research, Federation of Malaya, (28), 121–152.

    *Audy, J.R., Lawrence, R.F. & Vercammen-Grandjean, P.H. (1961) Trombiculidae. In: Zumpt, F. (Ed.) The Arthropod Parasites of Vertebrates in Africa South of the Sahara (Ethiopian Region). Publications of the South African Institute of Medical Research, Johannesburg, 9, 457 pp. (cited in Kolebinova and Vercammen-Grandjean 1978)

    Audy, J.R. & Nadchatram, M. (1957a) Malaysian parasites XXVI. New intranasal species of Traubacarus n. gen. (Acarina: Trombiculidae). Studies from the Institute for Medical Research, Federation of Malaya, (28), 187–230.

    Audy, J.R. & Nadchatram, M. (1957b) Malaysian parasites XIX. Vercammenia, a new genus of chiggers hypodermal in Amphibia (Acarina, Trombiculidae). Studies from the Institute for Medical Research, Federation of Malaya, (28), 95–102.

    Audy, J.R. & Nadchatram, M. (1960) Malaysian parasites XXXIX. Susa, a new genus related to Ascoschoengastia Ewing (Acarina, Trombiculidae), with descriptions of two new species. Studies from the Institute for Medical Research, Federation of Malaya, (29), 154–162.

    Audy, J.R., Nadchatram M., Loomis, R.B. & Traub R. (1965) Trombicula minor Berlese (Acarina, Trombiculidae): Designation of neotype with larval and post-larval stages from Malayan bats, and new name, Myotrombicula dilarami, for T. minor, Cooreman 1960 nec Berlese 1905. Acarologia, 7, 1–33.

    Audy, J.R. & Traub, R. (1950) Trombicula sylvestris sp. nov. (Acarina, Trombiculidae) from Malaya. Bulletin of the Raffles Museum, (23), 325–327.

    *Audy, J.R. & Vercammen-Grandjean, P.H. (1961a) African Trombiculidae (Acarina). 1. An interim note on the genus Trombicula Berlese and the subgenus Microtrombicula Ewing. Annals of the Natal Museum, 15, 125–133. (cited in Stekolnikov 2018b)

    *Audy, J.R. & Vercammen-Grandjean, P.H. (1961b) African Trombiculidae (Acarina). 2. The genera Eutrombicula Ewing and Sauriscus Lawrence, with description of a new subgenus, Squamicola. Annals of the Natal Museum, 15, 135–140. (cited in Stekolnikov 2018b)

    Audy, J.R. & Womersley, H. (1957) Malaysian parasites XXIX. New species of Oriental and Australian Trombiculidae (Acarina). Studies from the Institute for Medical Research, Federation of Malaya, (28), 359–382.

    Banks, N. (1916) Acariens from Australian and Tasmanian ants and ant nests. Transactions of the Royal Society of South Australia, 40, 224–240.

    Bassini-Silva, R., Jacinavicius, F.C., Muñoz-Leal, S., Maturano, R., Welbourn, W.C., Ochoa, R., Bauchan, G.R. & Barros-Battesti, D.M. (2018) A new species of the genus Eutrombicula Ewing, 1938 (Trombidiformes: Trombiculidae) and new records for the species Eutrombicula batatas (Linnaeus, 1758) in Brazil. Acarologia, 58, 976–986.

    https://doi.org/10.24349/acarologia/20184304

    Bassini-Silva, R., Jacinavicius, F.C., Oliveira, M., Peinado, L.C., Faxina, C., Moreira-Lima, L.,Welbourn, C., Ochoa, R., Fischer, E., Hingst-Zaher, E., Famadas, K.M., Faccini, J.L.H. & Barros-Battesti, D.M. (2021) A revision of Parasecia (Trombidiformes: Trombiculidae) with a description of a new species, a new genus and a key to species. Journal of Medical Entomology, 58, 146–181. (published online, 2020)

    https://doi.org/10.1093/jme/tjaa149

    Bassini-Silva, R., Jacinavicius, F.C., Pinter, A., Fournier, G.F.S.R., Lugarini, C., Ferreira, A., Moreira-Lima, L., Hingst-Zaher, E., Welbourn, C., Ochoa, R. & Barros-Battesti, D.M. (2019) Eutrombicula tinami (Oudemans, 1910) (Trombidiformes: Trombiculidae) in Brazil: a neglected ectoparasite of several animals including humans. Acarologia, 59, 412–423.

    https://doi.org/10.24349/acarologia/20194343

    Bassini-Silva, R., Jacinavicius, F.C., Welbourn, C., Barros-Battesti, D.M. & Ochoa, R. (2021) Complete type catalog of Trombiculidae sensu lato (Acari: Trombidiformes) of the U.S. National Entomology Collection, Smithsonian Institution. Smithsonian Contributions to Zoology, 652, 1–152. https://smithsonian.figshare.com/articles/book/ Complete_Type_ Catalog_of_Trombiculidae_sensu_lato_Acari_Trombidiformes_of_the_U_S_National_Entomology_Collection_Smithsonian_Institution/13546616/1

    Bennett, S.G. & Loomis, R.B. (1981) A new species of Eutrombicula (Acari: Trombiculidae) from the Chiricahua Mountains, southeastern Arizona. International Journal of Acarology, 7, 105–108.

    https://doi.org/10.1080/01647958108683251

    *Berdyev, A., Meledzhaeva, M.A., Amanguliev, A. & Mamednyazov, O. (1980) Parasitic arthropods of Western Turkmenia and their importance in the foci of some transmissible diseases. In: Charyev, O. Ch. and Meledzhaeva, M.A. (Eds.) Animal Parasites of Western Turkmenia. Ylym, Ashkhabad, pp. 210–263. (in Russian) (cited in Schmidt 2020)

    Berlese, A. (1888) Acari Austro-Americani quos collegit Aloysius Balzan. Bollettino della Società Entomologica Italiana, 20, 171–182.

    *Berlese, A. (1893) Acari Myriapoda et Scorpiones hucusque in Italia Reperta. Ordo Prostigmata (Trombidiidae). Padua 1882-1893, 155 pp. (cited in Wharton and Fuller 1952)

    Berlese, A. (1905) Acari nuovi, manipulus IV Acari de Giava. Redia, 2, 154–156.

    Berlese, A. (1910) Brevi diagnosi di generi e specie novi di acari. Redia, 6, 346–388.

    Berlese, A. (1912) Trombidiidae. Redia, 8, 1–291.

    *Beron, P. & Guéorguiev, V. (1967) Essai sur la faune cavernicole de Bulgarie. II. Résultats des recherches biospéléologiques de 1961-1965. Bulletin de l’Institut de Zoologie et Musée, 24, 151–212. (cited in Kolebinova 1992)

    Bhat, H.R. (1971) Whartonia kumaonensis, a new species of chigger (Acarina: Leeuwenhoekiinae) from Kumaon, Uttar Pradesh, India. Oriental Insects, 5, 585–588.

    Bischoff de Alzuet, A.D. & Mauri, R.A. (1987) Ácaros Trombiculidae del género Hannemania Oudemans, endoparásitos de anfibios. Revista de la Sociedad Entomológica Argentina, 44, 111–116.

    Bleaker, P. (1861) Guntheria, Proceedings of the Zoological Society of London, p. 635. In: Jordan, D.S. & Snyder, J.O. (1902) A review of the labroid fishes and related forms found in the waters of Japan. Proceedings of the United States National Museum, 24, 595–662.

    Boshell, J. & Kerr, J.A. (1942) Veinticinco especies nuevas de Trombidiideos en Colombia. Revisita de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales, 5, 110–127.

    Brennan, J.M. (1946a) A new genus and species of chigger, Chatia setosa (Trombiculidae, Acarina) from northwestern United States. Journal of Parasitology, 32, 132–135.

    Brennan, J.M. (1946b) Two new species of Trombicula: T. montanensis and T. aplodontiae (Acarina, Trombiculidae) from northwestern United States. Journal of Parasitology, 32, 441–444.

    Brennan, J.M. (1947) New species of chiggers (Acarina, Trombiculidae) from bats of the Nearctic region. Journal of Parasitology, 33, 245–252.

    https://doi.org/10.2307/3273557

    Brennan, J.M. (1948) New North American chiggers (Acarina, Trombiculidae). Journal of Parasitology, 34, 465–478.

    https://doi.org/10.2307/3273313

    *Brennan, J.M. (1949) Tracheation in chiggers with special reference to Acomatacarus arizonensis Ewing (Acarina, Trombiculidae). Journal of Parasitology, 35, 467–471. (cited in Wharton and Fuller 1952)

    https://doi.org/10.2307/3273649

    Brennan, J.M. (1951) Two new species of Neoschoengastia with a key to the species of the world (Acarina: Trombiculidae). Journal of Parasitology, 37, 577–582.

    https://doi.org/10.2307/3273352

    Brennan, J.M. (1952a) The genus Pseudoschoengastia Lipovsky, 1951, with the description of two new species, and a key to the world species, also Neoschoengastia paenitens, new name for Neoschoengastia kohlsi, Brennan, 1951, preoccupied (Acarina: Trombiculidae). Proceedings of the Entomological Society of Washington, 54, 133–137.

    Brennan, J.M. (1952b) Trombicula cynos Ewing, 1937, and three related new species (Acarina: Trombiculidae). Wasmann Journal of Biology, 10, 55–65.

    Brennan, J.M. (1952c) Two new Venezuelan chiggers (Acarina: Trombiculidae). Journal of Parasitology, 38, 143–146.

    https://doi.org/10.2307/3273833

    Brennan, J.M. (1953) A note on the chiggers of Jamaica (Acarina: Trombiculidae). Journal of Parasitology, 39, 292–295.

    https://doi.org/10.2307/3273952

    Brennan, J.M. (1956) Neoschoengastia moucheti, n. sp., and N. brennani Crossley and Loomis from Africa. (Acarina: Trombiculidae). Journal of Parasitology, 42, 650–652.

    https://doi.org/10.2307/3274885

    Brennan, J.M. (1957) Revival of Crotiscus Ewing, 1944, and descriptions of four new species of the genus from Peruvian rodents (Acarina: Trombiculidae). Journal of Parasitology, 43, 673–680.

    https://doi.org/10.2307/3286564

    Brennan, J.M. (1958) Beamerella and Kaaia, two new genera of chiggers from South America (Acarina: Trombiculidae). Journal of the Kansas Entomological Society, 31, 71–75.

    Brennan, J.M. (1959) Synonymy of Odontacarus Ewing, 1929, and Acomatacarus Ewing, 1942, with redescriptions of O. dentatus (Ewing) and O. australis (Ewing), also descriptions of three new species from southern United States (Acarina: Trombiculidae). Annals of the Entomological Society of America, 52, 1–6.

    https://doi.org/10.1093/aesa/52.1.1

    Brennan, J.M. (1960a) Ectonyx, a new Neotropical genus of chiggers (Acarina: Trombiculidae). Acarologia, 2, 88–91.

    Brennan, J.M. (1960b) Eight new species of Pseudoschoengastia from Mexico and Panama with a revised key to the species (Acarina: Trombiculidae). Acarologia, 2, 480–492.

    Brennan, J.M. (1962) Four new chiggers from Mexico. Journal of Parasitology, 48, 618–620.

    https://doi.org/10.2307/3274924

    Brennan, J.M. (1965a) Five new chiggers from southwestern United States (Acarina: Trombiculidae). Journal of Parasitology, 51, 108–113.

    https://doi.org/10.2307/3275656

    Brennan, J.M. (1965b) A small collection of chiggers (Acarina: Trombiculidae) from the north central Pacific. Journal of Parasitology, 51, 888–892.

    https://doi.org/10.2307/3276181

    Brennan, J.M. (1965c) Two new species and other records of chiggers from Texas (Acarina: Trombiculidae). Acarologia, 7, 79–83.

    Brennan, J.M. (1966) New records of chiggers (Acarina: Trombiculidae) from Baja California and islands of the Gulf of California. Journal of Parasitology, 52, 772–775.

    https://doi.org/10.2307/3276453

    Brennan, J.M. (1967) New records of chiggers from the West Indies. Studies on the Fauna of Curaçao and other Caribbean Islands, 24, 146–156.

    Brennan, J.M. (1968a) A collection of chiggers (Acarina: Trombiculidae) from rodents in southwestern Colombia. Journal of Parasitology, 54, 679–685.

    https://doi.org/10.2307/3277019

    Brennan, J.M. (1968b) Endoparasitic chiggers: IV. Revision of Kymocta Yunker and Brennan, 1962 (Acarina: Trombiculidae), with description of a new species. Journal of Parasitology, 54, 613–615.

    https://doi.org/10.2307/3277095

    Brennan, J.M. (1969a) Five new species of the unique genus Polylopadium (Acarina: Trombiculidae). Journal of Parasitology, 55, 866–871.

    https://doi.org/10.2307/3277233

    Brennan, J.M. (1969b) New bat chiggers of the genus Perissopalla from Venezuela and northeastern Brazil (Acarina: Trombiculidae). Journal of Medical Entomology, 6, 427–431.

    https://doi.org/10.1093/jmedent/6.4.427

    Brennan, J.M. (1969c) Three new species of subgenus Parasecia Loomis (genus Fonsecia) from northeastern Brazil and a key to the included species (Acarina: Trombiculidae). Journal of Parasitology, 55, 662–666.

    https://doi.org/10.2307/3277312

    Brennan, J.M. (1970a) Chiggers from the Bolivian-Brazilian border (Acarina: Trombiculidae). Journal of Parasitology, 56, 807–812.

    https://doi.org/10.2307/3277728

    Brennan, J.M. (1970b) Colicus, a new Neotropical genus with descriptions of two new species and key to included species (Acarina: Trombiculidae). Journal of Medical Entomology, 7, 271–272.

    https://doi.org/10.1093/jmedent/7.2.271

    Brennan, J.M. (1970c) A review of Neoschoengastia colombiae Boshell and Kerr 1942, with descriptions of two new species misidentified with it (Acarina: Trombiculidae: Colicus). Journal of Medical Entomology, 7, 489–493.

    https://doi.org/10.1093/jmedent/7.4.489

    Brennan, J.M. (1970d) A small collection of chiggers from Surinam (Acarina: Trombiculidae). Bulletin of the Southern California Academy of Sciences, 69, 32–37.

    Brennan, J.M. (1970e) Two new genera and a new species of Neotropical chiggers (Acarina: Trombiculidae). Annals of the Entomological Society of America, 63, 1694–1696.

    https://doi.org/10.1093/aesa/63.6.1694

    Brennan, J.M. (1971a) Anomalaspis ambiguus Brennan, 1952 redescribed and the questionable association of South American species with Pseudoschoengastia (Acarina: Trombiculidae). Journal of Medical Entomology, 8, 602–604.

    https://doi.org/10.1093/jmedent/8.5.602

    Brennan, J.M. (1971b) Endoparasitic chiggers: VI. Neoschoengastia esorhina sp. n. (Acarina: Trombiculidae), a second intranasal chigger of birds. Journal of Parasitology, 57, 666–667.

    https://doi.org/10.2307/3277936

    Brennan, J.M. (1971c) Three new chiggers from spiny rat, Proechimys guyannensis, in northern Brazil (Acarina: Trombiculidae). Annals of the Entomological Society of America, 64, 212–215.

    https://doi.org/10.1093/aesa/64.1.212

    Brennan, J.M. (1972a) Buclypeus, new genus for Trombicula ignota Brennan, T. rotundiscutata Fauran, and two new species (Acarina: Trombiculidae). Journal of Parasitology, 58, 1178–1181.

    https://doi.org/10.2307/3278161

    Brennan, J.M. (1972b) A new genus and two new species of Venezuelan chiggers with eyes in the scutum (Acarina: Trombiculidae). Journal of Medical Entomology, 9, 16–18.

    Brennan, J.M. (1973) Revival of Vanidicus, Brennan and Jones, 1961, descriptions of two new species of the genus from Venezuela and notes on related taxa (Acarina: Trombiculidae). Journal of Medical Entomology, 10, 107–109.

    Brennan, J.M. & Amerson, A.B. (1971) Six new species and additional records of chiggers from the central Pacific (Acarina: Trombiculidae). Journal of Parasitology, 57, 1311–1317.

    Brennan, J.M. & Beck, D.E. (1955) The chiggers of Utah (Acarina: Trombiculidae). Great Basin Naturalist, 15, 1–26.

    Brennan, J.M. & Dalmat, H.T. (1960) Chiggers of Guatemala (Acarina: Trombiculidae). Annals of the Entomological Society of America, 53, 183–191.

    Brennan, J.M. & Goff, M.L. (1977a) Keys to the genera of chiggers of the Western Hemisphere (Acarina: Trombiculidae). Journal of Parasitology, 63, 554–566.

    Brennan, J.M. & Goff, M.L. (1977b) The Neotropical genus Hoffmannina: four new species and other records from Mexico, Panama and Venezuela (Acarina: Trombiculidae). Journal of Parasitology, 63, 908–914.

    Brennan, J.M. & Goff, M.L. (1978a) Three new monotypic genera of chiggers (Acari: Trombiculidae) from South America. Journal of Medical Entomology, 14, 541–544.

    Brennan, J.M. & Goff, M.L. (1978b) Two new species of Crotiscus (Acari: Trombiculidae) from northern South America. Journal of Medical Entomology, 14, 569–572.

    Brennan, J.M. & Goff, M.L. (1979a) A new species of Atelepalme (Acari: Trombiculidae) from Venezuela. Acarologia, 20, 241–243.

    Brennan, J.M. & Goff, M.L. (1979b) A new species of Teratothrix (Acari: Trombiculidae) from Venezuela. Journal of Medical Entomology, 15, 272–273.

    Brennan, J.M. & Jones, E.K. (1954) A report on the chiggers (Acarina: Trombiculidae) of the Frances Simes Hastings Natural History Reservation, Monterey County, California. Wasmann Journal of Biology, 12, 155–194.

    Brennan, J.M. & Jones, E.K. (1959a) Keys to the chiggers of North America with synonymic notes and descriptions of two new genera (Acarina: Trombiculidae). Annals of the Entomological Society of America, 52, 7–16.

    Brennan, J.M. & Jones, E.K. (1959b) Pseudoschoengastia and four new Neotropical species of the genus (Acarina: Trombiculidae). Journal of Parasitology, 45, 421–429.

    Brennan, J.M. & Jones, E.K. (1960) Chiggers of Trinidad, B.W.I. (Acarina: Trombiculidae). Acarologia, 2, 493–540.

    Brennan, J.M. & Jones, E.K. (1961a) Chiggers of Peru (Acarina: Trombiculidae). Acarologia, 3, 172–205.

    Brennan, J.M. & Jones, E.K. (1961b) New genera and species of chiggers from Panama (Acarina: Trombiculidae). Journal of Parasitology, 47, 105–124.

    Brennan, J.M. & Jones, E.K. (1964a) Five new species of chiggers from South America (Acarina: Trombiculidae). Journal of Medical Entomology, 1, 307–310.

    Brennan, J.M. & Jones, E.K. (1964b) Four new chiggers from Argentina (Acarina: Trombiculidae). Journal of Parasitology, 50, 698–702.

    Brennan, J.M. & Loomis, R.B. (1959) A review of the reptile chiggers, genus Fonsecia (Acarina: Trombiculidae), with descriptions of two new American species. Journal of Parasitology, 45, 53–64.

    Brennan, J.M. & Lukoschus, F.S. (1971) Parasitic mites of Surinam, VIII. A new genus and species of chigger, Fauranius atecmartus, and additional records of species (Acarina: Trombiculidae). Bulletin of the Southern California Academy of Sciences, 70, 42–45.

    Brennan, J.M. & Reed, J.T. (1972a) Loomisia gen. n. with descriptions of three new Venezuelan species (Acarina: Trombiculidae). Journal of Parasitology, 58, 796–800.

    Brennan, J.M. & Reed, J.T. (1972b) Two new Venezuelan chiggers of the genus Polylopadium (Acarina: Trombiculidae). Journal of Medical Entomology, 9, 461–463.

    Brennan, J.M. & Reed, J.T. (1973a) More new genera and species of chiggers (Acarina: Trombiculidae) from Venezuela. Journal of Parasitology, 59, 706–710.

    Brennan, J.M. & Reed, J.T. (1973b) The Neotropical genus Aitkenius: Three new species and other records from Venezuela (Acarina: Trombiculidae). Journal of Parasitology, 59, 531–535.

    Brennan, J.M. & Reed, J.T. (1974a) Endoparasitic chiggers: VIII. The intradermal genus Intercutestrix Brennan and Yunker, 1966, and three new species (Acarina: Trombiculidae). Journal of Parasitology, 60, 185–187.

    Brennan, J.M. & Reed, J.T. (1974b) The genus Eutrombicula in Venezuela (Acarina: Trombiculidae). Journal of Parasitology, 60, 699–711.

    Brennan, J.M. & Reed, J.T. (1975) A list of Venezuela chiggers, particularly of small mammalian hosts (Acarina: Trombiculidae). Brigham Young University Science Bulletin, Biological Series, 20, 45–75.

    Brennan, J.M. & van Bronswijk, J.E.M.H. (1973) Endoparasitic chiggers VII. A redescription of Kymocta Yunker and Brennan, 1962, a new species from Venezuela, and other records (Acarina: Trombiculidae). Journal of Medical Entomology, 10, 449–451.

    Brennan, J.M. & van Bronswijk, J.E.M.H. (1975) Parasitic mites of Surinam XXI. New records of Surinam and certain French Guiana chiggers with the description of a new species of Loomisia Brennan and Reed, 1972 (Acarina: Trombiculidae). Journal of Medical Entomology, 12, 243–249.

    Brennan, J.M. & Vercammen-Grandjean, P.H. (1970) Revival of the monotypic genus Boshellia Ewing, 1950 (Acarina: Trombiculidae). Bulletin of the Southern California Academy of Sciences, 69, 100–103.

    Brennan, J.M. & Wharton, G.W. (1950) Studies on North American chiggers. No. 3. The subgenus Neotrombicula. American Midland Naturalist, 44, 153–197.

    Brennan, J.M. & White, J.S. (1960) New records and descriptions of chiggers (Acarina: Trombiculidae) on bats in Alabama. Journal of Parasitology, 46, 346–350.

    Brennan, J.M. & Yunker, C.E. (1964) A new species of Euschoengastia of potential veterinary importance (Acarina: Trombiculidae). Journal of Parasitology, 50, 311–312.

    Brennan, J.M. & Yunker, C.E. (1966a) The chiggers of Panama (Acarina: Trombiculidae). In: Wenzel, R.L. and Tipton, V.J. (Eds.) Ectoparasites of Panama. Field Museum of Natural History, Chicago, Illinois, pp. 221–266.

    Brennan, J.M. & Yunker, C.E.M. (1966b) Endoparasitic chiggers: III. Euschoengastia velata, n. sp. (Acarina), an intradermal parasite of boreal rodents in Montana. Journal of Medical Entomology, 3, 338–339.

    Brennan, J.M. & Yunker, C.E. (1969) Endoparasitic chiggers: V. New genera, species and records from Venezuela and Brazil (Acarina: Trombiculidae). Journal of Medical Entomology, 6, 299–304.

    Brown, J.H. & Brennan, J.M. (1952) A note on the chiggers (Trombiculidae) of Alberta. Canadian Journal of Zoology, 30, 338–343.

    Brown, W.A. (1990) A new genus and species of chigger (Acari: Trombiculidae) from the Philippine Islands. International Journal of Acarology, 16, 23–25.

    Brown, W.A. (1992) Description of six new species of the genus Leptotrombidium from Mindanao Island and notes on the medically important genera of chiggers (Acari: Trombiculidae) of the Philippine Islands. Journal of Medical Entomology, 29, 284–292.

    Brown, W.A. (1997a) A new species of chigger in the genus Siseca (Acari: Trombiculidae) from Camiguin Island, Mindanao, Philippines. Silliman Journal, 38, 170–175.

    Brown, W.A. (1997b) Ten new species of chiggers (Acari: Trombiculidae) from bat hosts from the Philippines. International Journal of Acarology, 23, 147–166.

    https://doi.org/10.1080/01647959708683558

    Brown, W.A. (1997c) Two new species of chiggers (Acari: Trombiculidae) from the Philippine Islands. International Journal of Acarology, 23, 27–32.

    https://doi.org/10.1080/01647959708684116

    Brown, W.A. (1998a) A new species of chigger in the genus Chiroptella from a bat host in the Philippines collection of the B.P. Bishop Museum. Silliman Journal, 39, 79–83.

    Brown, W.A. (1998b) Three new species of chiggers of the genera Cheladonta, Gahrliepia, and Walchiella (Acari: Trombiculidae) from rodent hosts from the Philippines. International Journal of Acarology, 24, 107–112.

    https://doi.org/10.1080/01647959808684137

    Brown, W.A. (1999) Two new species of Microtrombicula (Acari: Trombiculidae) from the Philippines. International Journal of Acarology, 25, 19–22.

    https://doi.org/10.1080/01647959908683607

    Brown, W.A. (2004) Two new species of chiggers (Acari: Trombiculidae) in Ascoschoengastia and Microtrombicula from hyrax (Hyracoidea) in Serengeti, Tanzania, Africa. International Journal of Acarology, 30, 41–44.

    https://doi.org/10.1080/01647950408684367

    *Brown, W.A. (2006a) An African chigger, Schoengastia pitheciagambiae n. sp. (Acari: Trombiculidae, Schoengastiini), from primate monkey hosts, Chorocebus sabaeus and Papio papio (L.) from Gambia. International Journal of Acarology, 32, 283–286. (cited in Stekolnikov 2018b)

    https://doi.org/10.1080/01647950608684469

    *Brown, W.A. (2006b) Two new species of Apoloniinae (Acari: Trombiculidae: Leeuwenhoekiidae) from African small mammals, with a key to the species of the world. Folia Parasitologica, 53, 217–222. (cited in Stekolnikov 2018b)

    Brown, W.A. (2007) A new taxon of African chigger, Tanautarsala callithrixa gen. nov., sp. nov. (Acari: Trombidioidea: Trombiculidae: Trombiculini), from the callithrix monkey Chlorocebus sabaeus (L.) from the Gambia. Systematic & Applied Acarology, 12, 223–228.

    https://doi.org/10.11158/saa.12.3.7

    Brown, W.A. (2008) Aboriginesia imlilica sp. nov. (Acari: Trombiculidae) from small mammal hosts from Marrikech Sector, Morocco. Systematic & Applied Acarology, 13, 145–149.

    https://doi.org/10.11158/saa.13.2.8

    Brown, W.A. (2009) Anasuscuta, a new genus of chigger (Acari: Leeuwenhoekiidae: Apoloniinae) described to accommodate Liuella monosetosa Brown. Systematic & Applied Acarology, 14, 248–250.

    https://doi.org/10.11158/saa.14.3.11

    Brown, W.A. & Goff, M.L. (1988a) Chigger mites (Acari: Trombiculidae) of Leyte Island, Philippine Islands with descriptions of five new species and key to the genera and species. Journal of Medical Entomology, 25, 214–223.

    https://doi.org/10.1093/jmedent/25.4.214

    Brown, W.A. & Goff, M.L. (1988b) Three new species of chiggers (Acari: Trombiculidae) from the Philippine Islands. International Journal of Acarology, 14, 115–119.

    https://doi.org/10.1080/01647958808683503

    Brown, W.A., Goff, M.L. & Nadchatram, M. (1988) The bat-infesting genus Rudnicula (Acari: Trombiculidae), with description of a new species from the Philippine Islands and a key to the species. Journal of Medical Entomology, 25, 17–19.

    https://doi.org/10.1093/jmedent/25.1.17

    Brown, W.A., Udagama-Randeniya, P.V. & Seneviratne, S.S. (2003) Two new species of chiggers (Acari: Leeuwenhoekiidae and Trombiculidae) from bats (Chiroptera) collected in the Kanneliya Forest Reserve of Sri Lanka. International Journal of Acarology, 29, 69–73.

    https://doi.org/10.1080/01647950308684323

    Brumpt, E. (1910) Précis de Parasitologie (2ème édition). Masson, Paris, 915 pp.

    *Bruyant, L. (1909) Essai de determination spécifique des rougets de l’homme (Leptus autumnalis Latr.). Comptes Rendus des Séances de la Société de Biologie, Paris, 67, 207–209. (cited in Wharton and Fuller 1952)

    Bruyant, L. & Joyeux, C.H. (1913) Un nouveau rouget des animaux en Guinée Française (Microtrombidium guineense n. sp.). Bulletin de la Société de Pathologie Exotique, 6, 202–205.

    Buffa, P. (1899) Trombidium canestrinii n. sp. Prespetto dell’Acarofauna Italiana, 8, 975–976.

    *Buitendijk, A.M. (1945) Voorloopige catalogus van de Acari in de Collectie Oudemans. Zoologische Mededelingen, 24, 281–391. (cited in Wharton and Fuller 1952)

    *Bunyea, H. & Wehr, E.E. (1941) Diseases and parasites of poultry. U.S. Department of Agriculture Farmers’ Bulletin, (1652), 1–82. (cited in Wharton and Fuller 1952)

    Chaisiri, K., Stekolnikov, A.A., Makepeace, B. & Morand, S. (2016) A revised checklist of chigger mites (Acari: Trombiculidae) from Thailand, with the description of three new species. Journal of Medical Entomology, 53, 321–342.

    https://doi.org/10.1093/jme/tjv244

    *Chang, M. & Wen, T. (1963) A new species of trombiculid mite belonging to the genus Shunsennia Jameson et Toshioka. Acta Zootaxonomica Sinica, 2, 254–257. (cited in Wen and Ma 1997)

    Chau, N.V. (2005) Three new chigger species (Acariformes: Trombiculidae) collected on mammal, bird and reptile from Vietnam. Tap Chi Sinh Hoc, 27, 8–15. (in Vietnamese)

    Chen, C. (1960) Preliminary survey of chigger mites in Hangchow area, Chekiang Province (Acariformes, Trombiculidae) with the description of new species. Acta Zoologica Sinica, 12, 31–44. (in Chinese with English summary)

    Chen, H.P. (1978) Survey of chigger mites on rats in southern Kiangsu Province with descriptions of two new species (Acariformes: Trombiculidae). Acta Entomologica Sinica 21, 197–203.

    Chen, H.T. & Hsu, P.K. (1955) Report on 12 species and varieties of chiggers in China, including six new species and two new varieties and the creation of a new genus. Acta Zoologica Sinica, 7, 101–145. (in Chinese with English summary)

    Chen, H.T. & Hsu, P.K. (1957) Scrub mites from mammals and birds, including the description of three new species. Acta Zoologica Sinica, 9, 379–404. (in Chinese with English summary)

    *Chen, H. T. & Hsu, P. K (1958a) Chinese species of trombiculid larvae and their distribution. Acta Zoologica Sinica, 10, 404–415.

    *Chen, H.T. & Hsu, P.K. (1958b) Studies on the trombiculid larvae of the genus Neoschoengastia from China. Acta Zoologica Sinica, 10, 377–384. (in Chinese with English summary) (cited in Chen and Hsu 1965a)

    Chen, H.T. & Hsu, P.K. (1959) Two new species of trombiculid mites belonging to the genus Whartonia (Acarina: Trombiculidae). Acta Zoologica Sinica, 11, 549–557. (in Chinese with English summary)

    Chen, H.T. & Hsu, P.K. (1963) Trombiculid mites infesting bats from Kwangtung Province. Acta Zoologica Sinica, 15, 597–606. (in Chinese with English summary)

    Chen, H.T. & Hsu, P.K. (1965a) Seven new species of trombiculid mites belonging to the genera Doloisia and Paradoloisia (Acarina: Trombiculidae). Acta Zoologica Sinica, 17, 283–297. (in Chinese with English summary)

    Chen, H.T. & Hsu, P.K. (1965b) Studies on the trombiculid mites of the genera Tragardhula and Trombicula (Acarina: Trombiculidae). Acta Zoologica Sinica, 17, 80–85. (in Chinese with English summary)

    Chen, H.T., Hsu, P.K. & Wang, T.C. (1956) Five species and varieties of trombiculid mites from China. Acta Zoologica Sinica, 8, 149–160.

    Chen, H.T., Hsu, P.K. & Wang, T.C. (1958) Studies on the trombiculid mites of the genus Euschoengastia, with a description of two new varieties. Acta Zoologica Sinica, 10, 388–398. (in Chinese with English summary)

    *Chen, S.B. (1975) Chigger mites (Acariformes, Trombiculidae) on bats in Kiangsu Province. Acta Entomologica Sinica, 18, 229–234. (in Chinese) (cited in Stekolnikov 2013)

    *Chen, S.B. (1980) Two new species of the genus Walchia (Acari: Trombiculidae) from China. International Journal of Acarology, 6, 55–58. (cited in Wen 2002)

    https://doi.org/10.1080/01647958008683194

    *Chen, S.B. & Fan, P.F. (1981) Two new species of trombiculid mites belonging to the genera Gahrliepia and Leptotrombidium from Anhui, China. Acta Zootaxonomica Sinica, 6, 145–148. (in Chinese with English summary) (cited in Stekolnikov 2013)

    Chen, S.B., Fan, P.F. & Chen, D.X. (1980) Two new species of chigger from Anhui Province. Acta Zootaxonomica Sinica, 5, 154–158. (in Chinese with English summary)

    Chen, X., Nian, F. & Duan, Y. (1996) Two new species of trombiculid mites of genus Chiroptella (Acari: Trombiculidae). Entomological Journal of East China, 39, 6–9. (in Chinese with English summary)

    Chu, F. (1964) Descriptions of two new species and a new subspecies of chigger mites collected in Chekiang Province. Acta Zootaxonomica Sinica, 1, 167–176. (in Chinese with English summary)

    Chung, L.H., Wu, W.J, Kuo, C.C. & Wang, H.C. (2015) A checklist of chigger mites (Acari: Trombiculidae and Leeuwenhoekiidae) from Taiwan, with descriptions of three new species. Journal of Medical Entomology, 52, 1241–1253.

    https://doi.org/10.1093/jme/tjv139

    Chung, L.H., Wu, W.J. & Wang, H.C. (2015) Chigger mite fauna of Taiwan (Acari: Trombiculidae and Leeuwenhökiidae [sic]). Centers for Disease Control, Ministry of Health and Welfare, Taipei, Taiwan, ix + 177 pp.

    Cooreman, J. (1951) Mission scientifique Suisse au Maroc (Août-Septembre 1950) Acari. Institut Royal des Sciences Naturelles de Belgique, 27, 1–4.

    Crossley, D.A., Jr. (1955) A new species of chigger mite from Colorado (Acarina, Trombiculidae). Journal of Parasitology, 41, 289–291.

    https://doi.org/10.2307/3274208

    *Crossley, D.A., Jr. (1960) Comparative external morphology and taxonomy of nymphs of the Trombiculidae (Acarina). University of Kansas Science Bulletin, 40, 135–321. (cited in Walters et al. 2011)

    Crossley, D.A., Jr. & Atyeo, W.T. (1972) A new species of chigger, Blankaartia pauli (Acarina: Trombiculidae), from the southeastern United States. Journal of Medical Entomology, 9, 253–254.

    https://doi.org/10.1093/jmedent/9.3.253

    Crossley, D.A., Jr. & Lipovsky, L.J. (1954) Two new chiggers from the central states (Acarina, Trombiculidae). Proceedings of the Entomological Society of Washington, 56, 240–246.

    Crossley, D.A., Jr. & Loomis, R.B. (1955) A new Neoschoengastia (Acarina: Trombiculidae) from western Kansas and eastern Colorado. Entomological News, 66, 113–118.

    Daniel, M. (1956) Eine neue Trombicula-larve aus der Sudslowakei. Zoologischer Anzeiger, 157, 210–216.

    Daniel, M. (1957) Euschoengastia ulcerofaciens n. sp., novy druh larev roztocu z celedi Trombiculidae zpusobujici vredy v pokozce hrabosu Microtus arvalis. Acta Societatis Entomologicae Čechosloveniae, 54, 172–179.

    Daniel, M. (1960) Rougets (Trombiculidae) recueillis en Albanie. Československá Parasitologie, 7, 21–36.

    Daniel, M. (1963) Cockingsia samsinaki, espèce nouvelle de Leeuwenhoekiidae (Trombidiformes). Acarologia, 5, 576–581.

    Daniel, M. & Brelih, S. (1959) The chigger larvae of the family Trombiculidae (Acari) - parasites upon small mammals in Slovenia. Acta Societatis Zoologicae Bohemoslovenicae, 23, 358–375.

    Daniel, M. & Dusbábek, F. (1959) Trombicula (Leptotrombidium) komareki, n. sp., un nouveau parasite des chiroptères en Tchécoslovaquie. Acarologia, 1, 421–425.

    *Daniel, M. & Heneberg, N. (1972) Chiggers parasitizing small mammals in the southern part of Yugoslavia (Acarina: Trombiculidae). Folia Parasitologica, 19, 133–138. (cited in Kudryashova 1998)

    Daniel, M. & Stekolnikov, A.A. (2002) New data on chigger mites of the subfamily Leeuwenhoekiinae (Acari: Trombiculidae) parasitizing bats in Cuba. Acarina, 10, 149–154.

    Daniel, M. & Stekolnikov, A.A. (2003a) Chigger mites (Acari: Trombiculidae) new to the fauna of Cuba, with the description of two new species. Folia Parasitologica, 50, 143–150.

    Daniel, M. & Stekolnikov, A.A. (2003b) To the fauna of chigger mites (Acari: Trombiculidae) parasitize bats in Cuba. Acarologia, 43, 29–37.

    Daniel, M. & Stekolnikov, A.A. (2004) Chigger mites of the genus Eutrombicula Ewing, 1938 (Acari: Trombiculidae) from Cuba, with the description of three new species. Folia Parasitologica, 51, 359–366.

    Daniel, M. & Stekolnikov, A.A. (2005) Three new species and new records of chigger mites (Acari: Trombiculidae) from East Hindukush, Pakistan. Acarologia, 45, 273–285.

    Daniel, M. & Stekolnikov, A.A. (2006) Three new species and new records of chigger mites (Acari: Trombiculidae) from Cuba. Acarina, 14, 69–78.

    Daniel, M. & Stekolnikov, A.A. (2009) Chigger mites (Acari: Trombiculidae) from Makalu Region in Nepal Himalaya, with description of three new species. Journal of Medical Entomology, 46, 753–765.

    https://doi.org/10.1603/033.046.0405

    Daniel, M., Stekolnikov, A.A., Hakimitabar, M. & Saboori, A. (2010) Chigger mites (Acari, Trombiculidae) parasitizing small mammals in the Eastern Hindu Kush and some other Afghan areas. Parasitology Research, 107, 1221–1233.

    https://doi.org/10.1007/s00436-010-1992-x

    Daniel, M. & Vercammen-Grandjean, P.H. (1985) Holubicula toroensis g. n., sp. n. (Acari: Trombiculidae), parasitic on the bat Tadarida condylura in SW Uganda. Folia Parasitologica, 32, 93–95.

    Daniel, M. & Zanina, Z.I. (1966) Deux nouvelles larves de Neotrombicula (Acari: Trombiculidae) de Tadjikistan (U.S.S.R.). Acarologia, 8, 78–85.

    Davis, R.M. & Loomis, R.B. (1971) The intranasal chigger Microtrombicula merrihewi (Acarina: Trombiculidae) in the North American free-tailed bat, Tadarida brasiliensis. Southwestern Naturalist, 15, 437–458.

    De la Cruz, J. & Daniel, M. (1994) Chigger mites (Acarina: Leeuwenhoekiidae) from Cuba. Folia Parasitologica, 41, 71–74.

    *Derrick, E.H., Smith, D.J.W., Brown, H.E. & Freeman, M. (1939) The role of the bandicoot in the epidemiology of “Q” fever: a preliminary survey. Medical Journal of Australia, 1, 150–155. (cited in Wharton and Fuller 1952)

    Dohany, A.L. & Cromroy H.L. (1976) New records of chiggers (Acarina: Trombiculidae) from Florida. Florida Entomologist, 59, 183–190.

    https://doi.org/10.2307/3493969

    Domrow, R. (1957) Malaysian parasites XXXI. Nymphs of Ascoschoengastia Ewing and Laurentella Audy (Acarina, Trombiculidae). Studies from the Institute for Medical Research, Federation of Malaya, (28), 394–402.

    Domrow, R. (1958) Acarina from Australian bats. Proceedings of the Linnean Society of New South Wales, 83, 227–240.

    Domrow, R. (1960a) The genus Guntherana (Acarina: Trombiculidae). Pacific Insects, 2, 195–237.

    Domrow, R. (1960b) Malaysian parasites XLII. Oculicola, new subgenus of Ascoschoengastia Ewing (Acarina): Trombiculidae from eyes of mammals. Studies from the Institute for Medical Research, Federation of Malaya, (29), 177–184.

    Domrow, R. (1961) New and little known Laelaptidae, Trombiculidae, and Listrophoridae (Acarina) from Australasia mammals. Proceedings of the Linnean Society of New South Wales, 86, 60–95.

    Domrow, R. (1962a) The Asian species of Whartonia (Acarina, Trombiculidae). Treubia, 26, 1–9.

    Domrow, R. (1962b) Four new species of Laurentella from S.E. Asia (Acarina: Trombiculidae). Acarologia, 4, 586–594.

    Domrow, R. (1962c) The genus Schoutedenichia in S.E. Asia (Acarina: Trombiculidae). Acarologia, 4, 357–361.

    Domrow, R. (1962d) The genus Siseca in Australia (Acarina: Trombiculidae). Journal of the Entomological Society of Queensland, 1, 21–24.

    Domrow, R. (1962e) The genus Walchiella (Acarina, Trombiculidae). Proceedings of the Linnean Society of New South Wales, 87, 105–115.

    Domrow, R. (1962f) Seven new Oriental-Australasian chiggers (Acarina, Trombiculidae). Treubia, 26, 39–56.

    Domrow, R. (1964) Five new chiggers of the genera Guntherana and Trombicula (Acarina, Trombiculidae). Acarologia, 6, 324–335.

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