Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2021-06-15
Page range: 423–431
Abstract views: 387
PDF downloaded: 19

Two new species of Neotropical Leptomorphus Curtis (Diptera: Mycetophilidae)

Pacific Northwest Diptera Research Lab, 1460 SW Allen St., Corvallis, OR, 97333 USA.
systematics, taxonomy, fungus gnats, Sciophilinae Diptera


Two new species of Leptomorphus Curtis, L. guatemalensis sp. n. and L. juxtafurcatus sp. n., are described from Guatemala, figured, and compared with congeners. The new species belong to the clade including the “furcatus” and “walkeri” species-groups as defined by Borkent & Wheeler (2012). L. guatemalensis sp. n. can be recognized by the gonocoxite bulbous and crescent-shaped with the apex asetose and spatula-like, but remains without a clear sister-species. Based on the structure of the male terminalia, particularly the presence of submedian, pointed lobes on sternite nine, L. juxtafurcatus sp. n. is suggested to be the sister-species to L. furcatus Borkent from the southwestern USA and northern Mexico; these species can be distinguished by the form of the gonocoxites.


  1. Borkent, C.J. & Wheeler, T.A. (2012) Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae). Zootaxa, 3549 (1), 1–117.
  3. Borkent, C.J. & Wheeler, T.A. (2013) Phylogeny of the tribe Sciophilini (Diptera: Mycetophilidae: Sciophilinae). Systematic Ento-mology, 38, 407–427.
  5. Cumming, J.M. & Wood, D.M. (2017) Adult Morphology and Terminology. In: Kirk-Spriggs, A.H. & Sinclair, B.J. (Eds.), Manual of Afrotropical Diptera. Volume 1. Introductory chapters and keys to Diptera families. Suricata. Vol. 4. South African Na-tional Biodiversity Institute, Pretoria, pp. 89–133.
  6. Henao-Sepúlveda, C., Wolff, M. & Amorim, D.S. (2020) Four new Neotropical species of Eudicrana Loew (Diptera, Mycetophi-lidae, Sciophilinae) from the Colombian high Andean ecosystems, with comments on the genus. ZooKeys, 988, 129–150.
  8. Johannsen, O.A. (1909) Fascicule 93. Diptera, Fam. Mycetophilidae. In: Wytsman, P. (Ed.), Genera Insectorum. Desmet-Verteneuil, Bruxelles, pp. 1–141.
  9. Johannsen, O.A. (1910) The fungus gnats of North America. The Mycetophilidae of North America. Part II. Bulletin of the Maine Agricultural Experiment Station, 180, 125–191.
  11. Kaspřák, D., Borkent, C. & Wahab, R. (2017) Leptomorphus sevciki sp. nov., a remarkable new wasp-mimicking fungus gnat from Brunei (Diptera: Mycetophilidae). Acta Entomologica Musei Nationalis Pragae, 57, 195–203.
  13. Kaspřák, D., Kerr, P., Sýkora, V., Tóthova, A. & Ševčík, J. (2019) Molecular phylogeny of the fungus gnat subfamilies Gnoristi-nae and Mycomyinae, and their position within Mycetophilidae (Diptera). Systematic Entomology, 44, 128–138.
  15. Matile, L. (1991) Diptera Mycetophiloidea Nouvelle-Calédonie. 4. Mycetophilidae Mycomyinae Sciophilinae et Gnoristinae. In: Chazeau, J. & Tillier, S. (Eds.), Zoologia Neocaledonica. Vol. 2. Muséum national d’Histoire Naturelle, Paris, pp. 233–250.
  16. Papp, L. & Ševčík, J. (2011) Eight new Oriental and Australasian species of Leptomorphus (Diptera: Mycetophilidae). Acta Zoo-logica Academiae Scientiarum Hungaricae, 57 (2), 139–159.
  17. Ševčík, J., Kaspřák, D. & Tóthová, A. (2013) Molecular phylogeny of fungus gnats (Diptera: Mycetophilidae) revisited: position of Manotinae, Metanepsiini, and other enigmatic taxa as inferred from multigene analysis. Systematic Entomology, 38, 654–660.
  19. Vockeroth, J.R. (2009) Mycetophilidae (fungus gnats). In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 1. NRC Research Press, Ottawa, Ontario, pp. 1–714.