Abstract
Papagona Ball, 1935 was originally described based on two species from the USA (Arizona). Both species of Papagona (P. papoosa Ball, 1935, type species of genus, and P. succinea Ball, 1935) are redescribed herein based on type specimens, including their previously unknown internal male genitalia. A new species from Brazil (Roraima) is described herein including the male and female terminalia. A taxonomic key to all included species is provided and additional diagnostic characters for this genus are proposed.
References
Amyot, C.J.B. & Audinet-Serville, J.G. (1843) Deuxième partie. Homoptères. Homoptera Latr. In: Histoire Naturelle des insectes. Hemiptères. Librairie encyclopédique de Roret, Paris, pp. 1–676. https://doi.org/10.5962/bhl.title.8471
Ball, E.D. (1935) Some new Issidae, with notes on others (Homoptera-Fulgoridae). Bulletin of the Brooklyn Entomological Society, 30, 37–41.
Berg, C. (1883) Addenda et emendanda ad Hemiptera Argentina. (Continuatio). Anales de la Sociedad Cientifica Argentina, 16, 180–191.
Bourgoin, T. (1987) A new interpretation of the homologies of the Hemiptera male genitalia, illustrated by the Tettigometridae (Hemiptera, Fulgoromorpha). 6th Auchenorrhyncha Meeting, Turin, Italy, 1987, 113–120.
Bourgoin, T. (1993) Female genitalia in Hemiptera Fulgoromorpha, Morphological and Phylogenetic Data. Annales de la Société Entomologique de France, 29, 225–244.
Bourgoin, T. (2021) FLOW (Fulgoromorpha Lists on The Web): a world knowledge base dedicated to Fulgoromorpha. Version 8. Last update 11 June 2021. Available from: http://hemiptera-databases.org/flow/ (accessed 12 June 2021)
Breddin, G. (1897) Hemipteren. Ergebnisse der Hamburger Magalhaensischen Sammelreise. Herausgegeben vom der Hamburger naturhistorisches Museum, 2 (4), 1–36.
Burmeister, H.C.C. (1835) Schnabelkerfe. Rhynchota. Fascicule 1. In: Handbuch der Entomologie, 1835, 99–183.
Caldwell, J.S. (1945) Notes on Issidae from Mexico (Homoptera, Fulgoroidea). Annals of the Entomological Society of America, 38 (1), 89–120. https://doi.org/10.1093/aesa/38.1.89
Doering, K.C. (1939) A contribution to the taxonomy of the subfamily Issinae in America north of Mexico (Fulgoridae, Homoptera). Part II. The University of Kansas Science Bulletin, 25, 447–575. https://doi.org/10.5962/bhl.part.1709
Doering, K.C. (1940) A contribution to the taxonomy of the subfamily Issinae in America north of Mexico (Fulgoridae, Homoptera). Part III. The University of Kansas Science Bulletin, 26, 83–167. [https://www.biodiversitylibrary.org/page/10705393]
Doering, K.C. (1941) A contribution to the taxonomy of the subfamily Issinae in America north of Mexico (Fulgoridae, Homoptera). The University of Kansas Science Bulletin, 27, 185–233. [https://www.biodiversitylibrary.org/page/4393771]
Emeljanov, A.F. (1999) Notes on delimitation of families of the Issidae group with description of a new species of Caliscelidae belonging to a new genus and tribe (Homoptera, Fulgoroidea). Zoosystematica Rossica, 8, 61–72.
Emeljanov, A.F. (2008) New species of the genus Peltonotellus Puton (Homoptera, Caliscelidae) from Kazakhstan, Middle and Central Asia. Tethys Entomological Research, 16, 5–12.
Emeljanov, A.F. (2015) A subgeneric subdivision of the genus Caliscelis Lap. with description of new species (Homoptera, Caliscelidae). Entomologicheskoe Obozreine, 94, 684–697. [in Russian, English translation published in Entomological Review, 95, 918–930] https://doi.org/10.1134/S001387381507009X
Fennah, R.G. (1954) The higher classification of the family Issidae (Homoptera: Fulgoroidea) with descriptions of new species. Transactions of the Royal Entomological Society of London, 105, 455–474. https://doi.org/10.1111/j.1365-2311.1954.tb00772.x
Freitas, A.S, Dietrich, C.H. & Takiya, D.M. (2020) Five new species of Caliscelidae (Insecta, Hemiptera) from Mexico and Panama, with additional redescriptions of little-know species. European Journal of Taxonomy, 717 (1), 27–69. https://doi.org/10.5852/ejt.2020.717.1097
Gnezdilov, V.M. & Wilson, M.R. (2006) Systematic notes on tribes in the family Caliscelidae (Hemiptera: Fulgoroidea) with the description of new taxa from Palaearctic and Oriental Regions. Zootaxa, 1359, 1–30.
Gnezdilov, V.M. & Bourgoin T. (2009) First record of the family Caliscelidae (Hemiptera, Fulgoroidea) from Madagascar, with description of new taxa from the Afrotropical Region and biogeographical notes. Zootaxa, 2020 (1), 1–36. https://doi.org/10.11646/zootaxa.2020.1.1
Gnezdilov, V.M. (2014a) A modern classification of the family Caliscelidae Amyot et Serville (Homoptera, Fulgoroidea). Entomological Review, 94 (2), 211–214. Original published in Zoologicheskiï Zhurnal, 92 (11), 1309–1311. [in Russian] https://doi.org/10.1134/S0013873814020092
Gnezdilov, V.M. (2014b) First record of the genus Issopulex (Hemiptera: Fulgoroidea: Caliscelidae) from Madagascar. Zoosystematica Rossica, 23, 234–237. https://doi.org/10.31610/zsr/2014.23.2.234
Gnezdilov, V.M. (2015) Madagascan Caliscelidae (Hemiptera, Fulgoroidea): current knowledge and description of a new genus and species. African Invertebrates, 56: 739–746. https://doi.org/10.5733/afin.056.0316
Melichar, L. (1906) Monographie der Issiden. Abhandlungen der Kaiserlich-Königliche Zoologisch-Botanische Gesellschaft in Wien, 3 (4), 13–27 + 329.
Metcalf, Z.P. (1958) Part 15: Issidae. In: General Catalogue of the Homoptera. Fascicle IV. Fulgoroidea. North Carolina State College, Raleigh, North Carolina, pp. 1–562.
O’Brien, L.B. & Wilson, S.W. (1985) Planthopper systematics and external morphology. In: Nault, L.R. & Rodriquez, J.G. (Eds.), The Leafhoppers and Planthoppers. Wiley-Interscience, New York, pp. 61–102.
O’Brien, L.B. (2002) The wild wonderful world of Fulgoromorpha. Denisia, 4 (176), 83–102.
Schmidt, E. (1910) Die Issinen des Stettiner Museums. (Hemiptera-Homoptera). Stettiner Entomologische Zeitung, 71, 146–220. https://www.biodiversitylibrary.org/page/9012707
Schmidt, E. (1927) Neue Zikaden-Gattungen und Arten. Archiv für Naturgeschichte, 91, 147–160.
Schmidt, E. (1932) Neue und bekannte Zikadengattungen und Arten der neuen Welt . (Hemipt.-Homopt.). Stettiner Entomologische Zeitung, 93, 35–54.
Song, N. & Liang, A.P. (2013) A preliminary molecular phylogeny of planthoppers (Hemiptera: Fulgoroidea) based on nuclear and mitochondrial DNA sequences. PLoS ONE, 8 (3), e58400, 1–11. https://doi.org/10.1371/journal.pone.0058400
Tishechkin, D.Y. (1998) Acoustic signals of Issidae (Homoptera, Cicadinea, Fulgoroidea) compared with signals of some other Fulgoroidea with notes on the taxonomic status of the subfamily Caliscelinae. Entomological Review, 78, 884–892.
Tishechkin, D.Y. (2003) Vibrational communication in Cercopoidea and Fulgoroidea (Homoptera:Cicadina) with notes on classification of higher taxa. Russian Entomological Journal, 12, 129–181. [in Russian]
Urban, J.M. & Cryan, J.R. (2007) Evolution of the planthoppers (Insecta: Hemiptera: Fulgoroidea). Molecular Phylogenetics and Evolution, 42, 556–572. https://doi.org/10.1016/j.ympev.2006.08.009
Wang, M.L., Zhang, Y.L. & Bourgoin, T. (2016) Planthopper family Issidae (Insecta: Hemiptera: Fulgoromorpha): Linking molecular phylogeny with classification. Molecular Phylogenetics and Evolution, 105, 224–234. https://doi.org/10.1016/j.ympev.2016.08.012
Yeh, W.B., Yang, C.T. & Hui, C.F. (1998) Phylogenetic relationships of the Tropiduchidae-group (Homoptera: Fulgoroidea) of planthoppers inferred through nucleotide sequences. Zoological Studies, 37 (1), 45–55. [http://zoolstud.sinica.edu.tw/Journals/37.1/45.pdf]
Yeh, W.B., Yang, C.T. & Hui, C.F. (2005) A molecular phylogeny of planthoppers (Hemiptera: Fulgoroidea) inferred from mitochondrial 16S rDNA sequences. Zoological Studies, 44 (4), 519–535. [http://zoolstud.sinica.edu.tw/Journals/44.4/519.pdf]