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Published: 2021-09-24

Middle Ordovician (Darriwilian) cheirurid trilobites from the Table Cove Formation, western Newfoundland, Canada

Department of Earth and Environmental Sciences, University of Iowa, 115 Trowbridge Hall, Iowa City, IA 52242, USA
Institute of Earth Sciences, University of Lausanne, Géopolis, CH-1015 Lausanne, Switzerland
Arthropoda Newfoundland Laurentia silicified taxonomy trilobites Cheiruridae


A diverse mid-Darriwilian trilobite fauna from the Table Cove Formation, western Newfoundland, has long been known on the basis of calcareous specimens from the west coast of the Great Northern Peninsula. Discovery of silicified faunas in localities on the east coast provides additional morphological information for previously known species, and also reveals the presence of multiple new genera and species. Many of these species are important, as they represent some of the earliest Laurentian members of the diversifying Whiterock Fauna, and seem phylogenetically near to the base of their respective clades. The concept of Sphaerexochinae is restricted to the genus Sphaerexochus itself, with the possible inclusion of Newfoundlandops n. gen. (type species N. karimae n. sp.), which shares with Sphaerexochus potential synapomorphies including the structure of the hypostome and the presence of fine granular sculpture on the librigenal border and field. Most of the Early and Middle Ordovician taxa with three pygidial segments previously classified as Sphaerexochinae by many authors are reassigned to Acanthoparyphinae on the basis of multiple putative synapomorphies. Other new cheirurid genera from the Table Cove Formation are the pilekiine Harebayaspis n. gen. (type species H. plurima n. sp.), and the deiphonine Mainbrookia n. gen. (type species M. becki n. sp.). Other species revised or described include the cheirurine Laneites polydorus (Billings, 1865), and the acanthoparyphines Cydonocephalus tiffanyae n. sp., Kawina stougei n. sp., and Kawina? sp.



  1. Adrain, J.M. (1998) Systematics of the Acanthoparyphinae (Trilobita), with species from the Silurian of Arctic Canada. Journal of Paleontology, 72, 698–718.
    Adrain, J.M. & Fortey, R.A. (1997) Ordovician trilobites from the Tourmakeady Limestone, western Ireland. Bulletin of the Natural History Museum, London, Geology Series, 53, 79–115.
    Adrain, J.M. & Karim, T.S. (2019) Revision of the Early Ordovician (late Tremadocian; Stairsian) cheirurid trilobite Tesselacauda Ross, with species from the Great Basin, western USA. Zootaxa, 4661 (2), 201–255.
    Adrain, J.M., Karim, T.S. & McAdams, N.E.B. (2020) Atlanticalymene, a new genus of Middle Ordovician (Darriwilian) calymenine trilobites, and revision of the calymenoidean genus Protocalymene Ross. Zootaxa, 4859 (1), 1–55.
    Ancygin, N.Â. (1977) [Trilobites of the Karakol-Mikhailovsk horizon of the Lower Ordovician of the southern Urals]. In: Sapelnikov, V.P. & Cuvosov, B.I. (Eds.), Materialy po Paleontologij Srednego Paleozoâ Urala i Sibiri. Trudy Instituta Geologij i Geohimij, Academiya Nauk SSSR, Ural,skij Naucnyj Centr., 126, 68–95. [in Russian]
    Angelin, N.P. (1854) Palaeontologica Scandinavica. Pars II, Crustacea formationis transitionis. Academiae Regiae Scientarum Suecanae, Holmiae, 25–92.
    Barrande, J. (1850) (Bohemian Trilobites). Bericht über die Mitteilungen von Freunden der Naturwissenschaften in Wien, 7, 4–7.
    Barton, D.C. (1915) A revision of the Cheirurinae, with notes on their evolution. Washington University Studies, 3, 102–152.
    Beyrich, E. (1845) Ueber einige böhmische Trilobiten. s.n., Berlin, 47 pp.
    Billings, E. (1865a) s.n. In: Palaeozoic fossils. Vol. I (4). Geological Survey of Canada, Montreal, pp. 168–345.
    Billings, E. (1865b) s.n. In: Palaeozoic fossils. Vol. I (5). Geological Survey of Canada, Montreal, pp. 344–426.
    Bradley, J.H., Jr. (1930) Fauna of the Kimmswick Limestone of Missouri and Illinois. Contributions from Walker Museum, 2, 219–290.
    Brögger, W.C. (1882) Die silurischen Etagen 2 und 3 im Kristianiagebiet und auf Eker. Universitäts-Programm, 32, Semester 1882, I–VIII, 1–376.
    Bruton, D.L. & Harper, D.A.T. (1981) Brachiopods and trilobites of the early Ordovician serpentine Otta Conglomerate, south central Norway. Norsk Geologisk Tidsskrift, 61, 153–181.
    Bruton, D.L., Hoel, O.A., Beyene, L.T. & Ivantsov, A.Y. (1997) Catalogue of the trilobites figured in Friedrich Schmidt’s “Revision der ostbaltischen silurischen Trilobiten” (1881-1907). Contributions form the Palaeontological Museum University of Oslo, 403, 1–117.
    Chatterton, B.D.E. (1980) Ontogenetic studies of Middle Ordovician trilobites from the Esbataottine Formation, Mackenzie Mountains, Canada. Palaeontographica, Abteilung A, 171, 1–74.
    Chatterton, B.D.E. & Ludvigsen, R. (1976) Silicified Middle Ordovician trilobites from the South Nahanni River area, District of Mackenzie, Canada. Palaeontographica, Abteilung A, 154, 1–106.
    Chatterton, B.D.E. & Perry, D.G. (1984) Silurian cheirurid trilobites from the Mackenzie Mountains northwestern Canada. Palaeontographica, Abteilung A, 184, 1–78.
    Chatterton, B.D.E., Siveter, D.J., Edgecombe, G.D. & Hunt, A.S. (1990) Larvae and relationships of the Calymenina (Trilobita). Journal of Paleontology, 64, 255–277.
    Chen, R.-Y. & Zhou, Z.-Y. (2002) An Ordovician (early Llanvirn) trilobite faunule from Nanzheng, southern Shaanxi. Acta Palaeontologica Sinica, 41, 219–231.
    Congreve, C.R. & Lieberman, B.S. (2011) Phylogenetic and biogeographic analysis of sphaerexochine trilobites. PLoS ONE, 6 (6), e21304.
    Cooper, B.N. (1953) Trilobites from the Lower Champlainian Formations of the Appalachian Valley. Geological Society of America Memoir, 55, 1–69.
    Čugaeva, M.N. (1973) [Trilobites]. In: Čugaeva, M.N., Ivanova, V.A., Oradovskaya, M.N. & Yakovlev, U.N. (Eds.), [Biostratigraphy of the lower part of the Ordovician in the North-east of the USSR, and biogeography of the uppermost Lower Ordovician]. Trudy Geologicheskogo Instituta. Akademiya Nauk SSSR, 213, 43–122. [in Russian]
    Čugaeva, M.N., Rozman, K. & Ivanova, V.A. (1964) Comparative biostratigraphy of Ordovician deposits in the north-east of the USSR. Trudy Geologicheskogo Instituta. Akademiya Nauk SSSR. Leningrad, 106, 1–226. [in Russian with English table of contents]
    Curtis, M.L.K. (1961) Ordovician trilobites from the Valongo area, Portugal. Cheiruridae, Pliomeridae and Dionidae. Boletim da Sociedade Geológica de Portugal, 14, 1–16.
    Dean, W.T. (1971) The trilobites of the Chair of Kildare Limestone (upper Ordovician) of eastern Ireland. Part 1. Monographs of the Palaeontographical Society, 531, 1–60.
    Edgecombe, G.D., Chatterton, B.D.E., Vaccari, N.E. & Waisfeld, B.G. (1999) Ordovician cheirurid trilobites from the Argentine Precordillera. Journal of Paleontology, 73, 1155–1175.
    Eichwald, K.E. (1843) Neuer Beitrag zur Geognosie Esthlands und Finnlands. Beiträge zur Kenntniss des Russischen Reiches und der angränzenden Länder Asiens, 8, 1–138.
    Esker, G.C. (1964) New species of trilobites from the Bromide Formation (Pooleville Member) of Oklahoma. Oklahoma Geology Notes, 24, 195–209.
    Esker, G.C. & Levin, H. (1964) Pandaspinapyga, a new trilobite genus from the Kimmswick Limestone (Ordovician) of Missouri. Journal of Paleontology, 38, 776–778.
    Esmark, M.T. (1833) Om nogle nye Arter af Trilobiter. Magazin for Naturvidenskaberne, Christiania, 11, 268–270.
    Evitt, W.R. (1951) Some Middle Ordovician trilobites of the families Cheiruridae, Harpidae and Lichidae. Journal of Paleontology, 25, 587–616.
    Fortey, R.A. (1980) The Ordovician trilobites of Spitsbergen. III. Remaining trilobites of the Valhallfonna Formation. Norsk Polarinstitutt Skrifter, 171, 1–163.
    Fortey, R.A. (2005) Opipeuterella, a replacement name for the trilobite Opipeuter Fortey, 1974, preoccupied. Journal of Paleontology, 79, 1036.[1036:OARNFT]2.0.CO;2
    Fortey, R.A. & Bruton, D.L. (2013) Lower Ordovician trilobites of the Kirtonryggen Formation, Spitsbergen. Fossils and Strata, 59, 1–116.
    Hammann, W. (1992) The Ordovician trilobites from the Iberian Chains in the province of Aragón, NE-Spain. I. The trilobites of the Cystoid Limestone (Ashgill Series). Beringeria, 6, 3–219.
    Hansen, T. (2005) A new trilobite species of Hemisphaerocoryphe from the Arenig of St. Petersburg area, Russia. Norwegian Journal of Geology, 85, 203–208.
    Hansen, T. & Nielsen, A.T. (2003) Upper Arenig trilobite biostratigraphy and sea-level changes at Lynna River near Volkhov, Russia. Bulletin of the Geological Society of Denmark, 50, 105–114.
    Hawle, I. & Corda, A.J.C. (1847) Prodrom einer Monographie der böhmischen Trilobiten. J. G. Calve, Prague, 176 pp.
    Hintze, L.F. (1953) Lower Ordovician trilobites from western Utah and eastern Nevada. Utah Geological and Mineralogical Survey Bulletin, 48, 1–249. [for 1952]
    Holliday, S. (1942) Ordovician trilobites from Nevada. Journal of Paleontology, 16, 471–478.
    Holloway, D.J. & Campbell, K.S.W. (1974) The Silurian trilobite Onycopyge Woodward. Palaeontology, 17, 409–421.
    Ingham, J.K., Curry, G.B. & Williams, A. (1986) Early Ordovician Dounans Limestone fauna, Highland Border Complex, Scotland. Transactions of the Royal Society of Edinburgh: Earth Sciences, 76, 481–513. [for 1985]
    Jell, P.A. & Adrain, J.M. (2003) Available generic names for trilobites. Memoirs of the Queensland Museum, 48, 331–553.
    Klikushin, V., Evdokimov, A. & Pilipyuk, A. (2009) Ordovician trilobites of the St. Petersburg Region, Russia. Petersburg Paleontological Laboratory, Griffon Enterprises, Master Fossil Japan, St. Petersburg, 541 pp.
    Kobayashi, T. & Hamada, T. (1976) A new Silurian trilobite from Ofunato, North Japan. Proceedings of the Japan Academy, 52, 367–370.
    Kobayashi, T. & Hamada, T. (1986) The second addition to the Silurian trilobite fauna of Yokokura-yama, Shikoku, Japan. Transactions and Proceedings of the Palaeontological Society of Japan, 143, 447–462.
    Krueger, H.-H. (1994) Über die mittelordovizische Trilobitengattung Hemisphaerocoryphe. Archiv für Geschiebekunde, 1 (8), 469–484.
    Krueger, H.-H. (1996) Nachtrag zu mittelordovizischen Trilotengattungen Hemisphaerocoryphe und Nieszkowskia. Archiv für Geschiebekunde, 2 (1), 21–26.
    Lane, P.D. (1971) British Cheiruridae (Trilobita). Monographs of the Palaeontographical Society, 530, 1–95.
    Lane, P.D. (1972) New trilobites from the Silurian of north-east Greenland, with a note on trilobite faunas in pure limestones. Palaeontology, 15, 336–364.
    Lane, P.D. (2002) The taxonomic position and coaptative structures of the Lower Ordovician trilobite Cyrtometopus. Special Papers in Palaeontology, 67, 153–169.
    Lee, D.-C. & Chatterton, B.D.E. (1997) Ontogenies of trilobites from the Lower Ordovician Garden City Formation of Idaho and their implications for the phylogeny of the Cheirurina. Journal of Paleontology, 71, 683–702.
    Li, Y.-X., Song, L.-S., Zhou, Z.-Q. & Yang, J.-Y. (1975) [Stratigraphical Gazetteer of the Lower Paleozoic, western Dabashan Mountains]. Geological Publishing House, Beijing, 372 pp.
    Lindström, G. (1885) Förteckning på Gotlands Siluriska Crustacéer. Öfversigt af Kongl. Vetenskaps-Akademiens Förhandlingar. Stockholm, 6, 37–100.
    Lisogor, K.A. (1995) [Llanvirn trilobites from the Ordovician of Betpak-Dali (southern Kazakhstan)]. Vestnik kazakhskogo Natsional’nogo universiteta Geologo-Mineralogicheskikh nauk, Alma-ata, 4, 110–115. [in Russian]
    Lu, Y.-H. (1975) Ordovician trilobite faunas of central and southwestern China. Palaeontologia Sinica, New Series B, 11, 1–463. [in Chinese and English]
    Ludvigsen, R. (1977) The Ordovician trilobite Ceraurinus Barton in North America. Journal of Paleontology, 51, 959–972.
    Ludvigsen, R. (1979) A trilobite zonation of Middle Ordovician rocks, southwestern District of Mackenzie. Geological Survey of Canada Bulletin, 312, 1–99.
    Maletz, J., Egenhoff, S., Böhme, M., Asch, R., Borowski, K., Höntzsch, S., Kirsch, M. & Werner, M. (2011) A tale of both sides of Iapetus—upper Darriwilian (Ordovician) graptolite faunal dynamics on the edges of two continents. Canadian Journal of Earth Sciences, 48, 841–859.
    Männil, R. (1958) Trilobites of the families Cheiruridae and Encrinuridae from Estonia. Eesti NSV Teaduste Akadeemia Geoloogia Instituudi Uurimused, 3, 165–212. [in Russian with English summary]
    McAdams, N.E.B. & Adrain, J.M. (2009) Heckethornia, a new genus of dimeropygid trilobites from the Lower Ordovician (Ibexian; Tulean and Blackhillsian) of the Great Basin, western USA. Canadian Journal of Earth Sciences, 46, 875–914.
    Morris, S.F. & Fortey, R.A. (1985) Catalogue of the type and figured specimens of trilobites in the British Museum (Natural History). British Museum (Natural History), London, 183 pp.
    Nieszkowski, J. (1859) Zusätze zur Monographie der Trilobiten der Ostseeprovinzen, nebst der Beschreibung einiger neuen obersilurischen Crustacean. Archiv für die Naturkunde Liv-, Ehst-und Kurlands, 2, 345–384.
    Nikolaisen, F. (1961) The Middle Ordovician of the Oslo Region, Norway. 7. Trilobites of the suborder Cheirurina. Norsk Geologisk Tidsskrift, 41, 279–310.
    Öpik, A.A. (1937) Trilobiten aus Estland. Tartu Ülikooli Geoloogia-Intituudi Toimetused, 52, 1–163.
    Pärnaste, H. (2003) The Lower Ordovician trilobite Krattaspis: The earliest cyrtometopinid (Cheiruridae) from the Arenig of the East Baltic. In: Lane, P.D., Siveter, D.J. & Fortey, R.A. (Eds.), Trilobites and their relatives. Contributions from the Third International Conference, Oxford 2001. Special Papers in Palaeontology, 70, 241–257.
    Pärnaste, H. (2004a) Early Ordovician trilobites of the suborder Cheirurina in Estonia and Russia: systematics, evolution and distribution. PhD Thesis, University of Tartu, Tartu, 137 pp.
    Pärnaste, H. (2004b) Revision of the Ordovician cheirurid trilobite genus Reraspis with the description of the earliest representative. Proceedings of the Estonian Academy of Sciences, Geology, 53, 125–138.
    Peng, S.-C. (1990) Tremadoc stratigraphy and trilobite faunas of northwestern Hunan. 2. Trilobites from the Panjiazui Formation and the Madaoyu Formation in Jiangnan Slope Belt. Beringeria, 2, 55–171.
    Poulsen, C. (1946) Notes on Cambro-Ordovician fossils collected by the Oxford University Ellesmere Land Expedition 1934–5. Quarterly Journal of the Geological Society of London, 102, 299–337.
    Přibyl, A., Vaněk, J. & Pek, I. (1985) Phylogeny and taxonomy of family Cheiruridae (Trilobita). Acta Universitatis Palackianae Olomucensis Facultas rerum naturalium, 83, 107–193.
    Ramsköld, L. (1983) Silurian cheirurid trilobites from Gotland. Palaeontology, 26, 175–210.
    Raymond, P.E. (1905) The trilobites of the Chazy Limestone. Annals of the Carnegie Museum, 3, 328–386.
    Raymond, P.E. (1910) Trilobites of the Chazy formation in Vermont. Report of the Vermont State Geologist, 7, 213–248.
    Raymond, P.E. (1913) Subclass Trilobita. In: Eastman, C.R. (Ed.), Text-book of paleontology Vol. 1. 2nd Edition. MacMillan, New York, pp. 607–638.
    Reed, F.R.C. (1945) Revision of certain Lower Ordovician faunas from Ireland. 1. Trilobites. Geological Magazine, 82, 55–66.
    Reed, F.R.C. (1896) Woodwardian Museum notes. Notes on the evolution of the genus Cheirurus. Geological Magazine, (4)3, 117–123, 161–167.
    Ross, R.J. Jr. (1951) Stratigraphy of the Garden City Formation in northeastern Utah, and its trilobite faunas. Peabody Museum of Natural History, Yale University, Bulletin, 6, 1–161.
    Ross, R.J. Jr. (1958) Trilobites in a pillow-lava of the Ordovician Valmy Formation, Nevada. Journal of Paleontology, 32, 559–570.
    Ross, R.J. Jr. (1967) Some Middle Ordovician brachiopods and trilobites from the Basin Ranges, western United States. United States Geological Survey Professional Paper, 523-D, 1–43.
    Ross, R.J. Jr. (1972) Fossils from the Ordovician bioherm at Meiklejohn Peak, Nevada. United States Geological Survey Professional Paper, 685, 1–47.
    Schmidt, F. (1881) Revision der ostbaltischen silurischen Trilobiten. Abtheilung I: Phacopiden, Cheiruriden und Encrinuriden. Mémoires de l’Academie Impériale des Sciences de St.-Petersbourg, VII’e Série, 30, 1–237.
    Sdzuy, K. (1955) Die Fauna der Leimitz-Schiefer (Tremadoc). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 492, 1–74.
    Shaw, F.C. (1968) Early Middle Ordovician Chazy trilobites of New York. New York State Museum and Science Service Memoir, 17, 1–163.
    Shaw, F.C. (1974) Simpson Group (Middle Ordovician) trilobites of Oklahoma. Paleontological Society Memoir, 6, 1–54.
    Šnajdr, M. (1984) Revision of the trilobite type material of I. Hawle and A. J. C. Corda, 1847. Sborník Národního Muzea v Praze. Rada B. Prírodní Vedy, 39, 129–212. [for 1983]
    Stouge, S. (1984) Conodonts of the Middle Ordovician Table Head Formation, western Newfoundland. Fossils and Strata, 16, 1–145.
    Thomas, A.T. (1981) British Wenlock trilobites. Part 2. Monographs of the Palaeontographical Society, 559, 57–99.
    Thomson, C.W. (1857) On some species of Acidaspis from the Lower Silurian Beds of the south of Scotland. Quarterly Journal of the Geological Society of London, 13, 206–210.
    Thorslund, P. (1940) On the Chasmops Series of Jemtland and Sodermanland (Tvären). Sveriges Geologiska Undersökning, Afhandlingar och Uppsatser, Series C, 436, 1–191.
    Torsvik, T.H. & Cocks, L.R.M. (2017) Earth History and Palaeogeography. Cambridge University Press, Cambridge, 317 pp.
    Tremblay, J.V. & Westrop, S.R. (1991) Middle Ordovician (Whiterockian) trilobites from the Sunblood Formation, District of Mackenzie, Canada. Journal of Paleontology, 65, 801–824.
    Tripp, R.P. (1962) Trilobites from the “confinis” Flags (Ordovician) of the Girvan district, Ayrshire. Transactions of the Royal Society of Edinburgh, 65, 1–40.
    Tripp, R.P. (1967) Trilobites from the upper Stinchar Limestone (Ordovician) of the Girvan District, Ayrshire. Transactions of the Royal Society of Edinburgh, 67, 43–94.
    Tripp, R.P. (1979) Trilobites from the Ordovician Auchensoul and Stinchar Limestones of the Girvan District, Strathclyde. Palaeontology, 22, 339–361.
    Tripp, R.P., Rudkin, D.R. & Evitt, W.R. (1997) Silicified trilobites of the genus Sphaerocoryphe from the Middle Ordovician of Virginia. Canadian Journal of Earth Sciences, 34, 770–788.
    Vogdes, A.W. (1890) A bibliography of Paleozoic Crustacea from 1698 to 1890 including a list of North American species and a systematic arrangement of genera. United States Geological Survey Bulletin, 63, 1–177.
    Vogdes, A.W. (1893) A classed and annotated bibliography of the Palaeozoic Crustacea 1698-1892 to which is added a catalogue of North American species. Occasional Papers of the California Academy of Sciences, 4, 1–412.
    Waisfeld, B.G. & Vaccari, N.E. (2003) Trilobites. In: Benedetto, J.L. (Ed.), Ordovician Fossils from Argentina. Secretaría de Ciencia y Tecnología, Universidad Nacional de Córdoba, Cordoba, pp. 295–409.
    Warburg, E. (1925) The trilobites of the Leptaena Limestone in Dalarne with a discussion of the zoological position and the classification of the Trilobita. Bulletin of the Geological Institution of the University of Uppsala, 17, 1–446.
    Webby, B.D. (1974) Upper Ordovician trilobites from central New South Wales. Palaeontology, 17, 203–252.
    Whittard, W.F. (1966) The Ordovician trilobites of the Shelve Inlier, west Shropshire. Part 8. Monographs of the Palaeontographical Society, 508, 265–306.
    Whittington, H.B. (1963) Middle Ordovician trilobites from Lower Head, western Newfoundland. Bulletin of the Museum of Comparative Zoology, Harvard, 129, 1–118.
    Whittington, H.B. (1965) Trilobites of the Ordovician Table Head Formation, western Newfoundland. Bulletin of the Museum of Comparative Zoology, Harvard, 132, 275–442.
    Whittington, H.B. & Evitt, W.R. (1954) Silicified Middle Ordovician trilobites. Geological Society of America Memoir, 59, 1–137.
    Whittington, H.B. & Kindle, C.H. (1963) Middle Ordovician Table Head Formation, western Newfoundland. Geological Society of America Bulletin, 74, 745–758.
    Woodward, H. (1880) Description of a new genus of trilobites, Onycopyge liversidgei, from the Silurian of New South Wales. Geological Magazine, 7, 97–99.
    Yang, J.-L., Yu, S.-Y., Liu, G.-T., Su, N.-M., He, M.-H., Shang, J.G., Zhang, H.-Q., Zhu, H.-Y., Li, Y.-J. & Guo, G.-S. (1991) [Cambrian stratigraphy, lithofacies, palaeogeography and trilobite faunas of East Qinling-Dabashan Mountains, China]. China University of Geosciences, Wuhan, 246 pp.
    Zhang, W.-T. (1974) [Silurian Trilobita]. In: Nanking Institute of Geology and Paleontology, Academica Sinica (Ed.), [Handbook of the stratigraphy and paleontology of southwest China]. Science Press, Beijing, 173–187. [in Chinese; Chang, W.-T.]
    Zhao, D., Zhang, M.-S., Cheng, L.-R. & Zhu, H.-S. (1997) [Ordovician strata, trilobite fauna and its tectonic setting of Hinggan Region, China]. Jilin People’s Press, Changchun, 181 pp. [in Chinese with English abstract]
    Zhou, Z.-Y. & Zhou, Z.-Q. (2008) Latest Cambrian and Ordovician. In: Zhou, Z.-Y. & Zhen, Y.-Y. (Eds.), Trilobite Record of China. Science Press, Beijing, pp. 208–274.
    Zhou, Z.-Y., Yin, G.-Z. & Zhou, Z.-Q. (2014) Ordovician (Darriwilian-early Katian) trilobite faunas of northwestern Tarim, Xinjiang, China. Memoirs of the Association of Australasian Palaeontologists, 46, 1–142.
    Zhou, Z.-Y., Zhou, Z.-Q. & Yin, G.-Z. (2016) Ordovician trilobites from the uppermost Zhuozishan Formation (early Darriwilian) at Zhuozishan, Wuhai, Inner Mongolia. Memoirs of the Association of Australasian Palaeontologists, 49, 289–328.
    Zhou, Z.-Y., Dean, W.T., Yuan, W.-W. & Zhou, T.-R. (1998) Ordovician trilobites from the Dawangou Formation, Kalpin, Xinjiang, north-west China. Palaeontology, 41, 693–735.


How to Cite

ADRAIN, J. M. ., & PÉREZ-PERIS, F. . (2021). Middle Ordovician (Darriwilian) cheirurid trilobites from the Table Cove Formation, western Newfoundland, Canada. Zootaxa, 5041(1), 1–73.