Abstract
The genus Heraclia (Lepidoptera, Noctuidae, Agaristinae) and the species involved in the confused nomenclature and taxonomy of the type species, Phalaena euphemia Stoll, 1781 are investigated. The rediscovery of the type specimen of Noctua geryon Fabricius, 1781, a taxon that had previously been treated as the senior synonym of euphemia, justified the revival of the latter from synonymy with the former. The type material of euphemia could not be traced and a neotype is designated to fix the published name to a physical specimen. In addition to detailed diagnoses and re-descriptions, the revised taxonomy of Heraclia euphemia and Heraclia geryon is provided and the following new synonymies are proposed: Eusemia pallida Walker, 1854 and Eusemia niveosparsa Westwood, 1881 are synonymised with H. geryon and Eusemia nugatrix Westwood, 1881 is synonymised with H. euphemia. Distributional data for each of the species are presented.
References
Aurivillius, C. (1892) Verzeichniss einer vom Herrn Fritz Theorin aus Gabun und dem Gebiete des Camerunflusses heimgebrachten Schmetterlings-Sammlung. II. Heterocera. Entomologisk Tidskrift, 13, 181–200.
Aurivillius, C. (1925) s.n. In: Wissenschaftliche Ergebnisse der zweiten Deutschen Zentral-Afrika-Expedition, 1910–1911 unter Führung Adolf Friedrichs, Herzog zu Mecklenburg. Vol. 1. Zoologie. Lieferung 18. Lepidoptera. Part 4. Klinkhardt & Biermann, Leipzig, pp. 1243–1359, pls 49–50.
Boisduval, J.B. (1874) Monographie des Agaristidées (Lépidoptères). Revue et magasin de zoologie pure et appliquée, 3 (2), 26–110.
Butler, A.G. (1875a) Notes on certain Genera of Agaristidae, with descriptions of new species. The Annals and Magazine of Natural History, Series 4, 15 (86), 135–144. https://doi.org/10.1080/00222937508681041
Butler, A.G. (1875b) Revision of the lepidopterous genus Eusemia, with descriptions of new species. Entomologist’s Monthly Magazine, 12, 116–125.
Butler, A.G. (1877) Illustrations of typical specimens of Lepidoptera Heterocera in the collection of the British Museum. Part I. Printed by order of the Trustees of the British Museum, London, 62 pp.
Chainey, J.E. (2005) The species of Papilionidae and Pieridae (Lepidoptera) described by Cramer and Stoll and their putative type material in the Natural History Museum in London. Zoological Journal of the Linnean Society, 145, 283–337. https://doi.org/10.1111/j.1096-3642.2005.00184.x
Edge, D.A. (2013) Family Lycaenidae. In: Mecenero, S., Ball, J.B., Edge, D.A., Hamer, M.L., Henning, G.A., Krüger, M., Pringle, E.L., Terblanche, R.F. & Williams, M.C. (Eds.), Conservation Assessment of Butterflies of South Africa, Lesotho and Swaziland: Red List and Atlas. Saftronics (Pty) Ltd. and the Animal Demography Unit, Johannesburg, pp. 335–610.
Fabricius, J.C. (1781) Species Insectorum, Exhibentes Eorum Differentias Specificas, Synonyma Auctorum, Loca Natalia, Metamorphosin Adiectis, Observationibus, Descriptionibus 2. Impensis C.E. Bohnii, Hamburgi et Kilonii, 517 pp. https://doi.org/10.5962/bhl.title.36509
Fabricius, J.C. (1792) Entomologia systematica emandata et aucta. Secundum classes, ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus. Tome III. Impensis C.G. Proft, fil et soc, Hafniae, 349 pp. https://doi.org/10.5962/bhl.title.125869
Godart, J.B. (1824) Genre Polyommate. In: Latreille, P.A. (Ed.), Encyclopedie Methodique: Histoire Naturelle: Entomologie, 9 (2), 595–704.
Guillaumin, M. (1976) Participation des Euphaedra et des Bebearia (Lep. Nymphaidae) aux associations mimétiques de a forêt équatoriale Africaine. I.—Associations ayant des hétérocères a moeurs diurnes comme modèles. Bulletin de la Société Zoologique de France, 101 (4), 603–611.
Hampson, G.F. (1901) Catalogue of the Lepidoptera Phalænae in the British Museum. Volume III. Catalogue of the Acrtiadæ (Arctianæ) and Agaristidæ in the collection of the British Museum. Trustees of the British Museum (Natural History), London, 690 pp., pls. 36–54.
Hampson, G.F. (1920) Catalogue of the Lepidoptera Phalænæ in the British Museum. Supplement Volume II. Catalogue of the Lithosiadæ (Arctianæ) and Phalænoididae in the collection of the British Museum. Trustees of the British Museum (Natural History), London, 619 pp., pls. 42–71.
Hancock, E.G. (2015) The shaping role of Johann Christian Fabricius: William Hunter’s insect collection and entomology in 18th-century London. In: Hancock, E.G., Pearce, N. & Campbell, M. (Eds.), William Hunter’s World: The Art and Science of Eighteenth-Century Collecting. Ashgate, Burlington, Vermont, pp. 151–163.
Hübner, J. ([1820]) s.n. In: Verzeichniss bekannter Schmettlinge [sic]. bey dem Verfasser zu Finden, Augsburg, pp. 12–13 + 177–208.
International Commission on Zoological Nomenclature. (1958) Opinion 516. Determination under the plenary powers of the relative precedence to be assigned to certain works on the order Lepidoptera (Class Insecta) published in 1775, by Pieter Cramer, Michael Denis & Ignaz Schiffermuller, Johann Christian Fabricius, Johann Caspar Fuessly, and S.A. von Rottemburg, respectively. Opinions and Declarations Rendered by the International Commission on Zoological Nomenclature, 19, 1–43.
Jordan, K. (1913) Agaristidae. In: Seitz, A., Die Gross-Schmetterlinge der Erde. Vol. 15. Alfred Kernen Verlag, Suttgart, pp. 1–29, pls. 1–4.
Jordan, K. (1926) Some new Agaristidae with remarks on nomenclature. Novitates zoologicae, 33 (3), 371–378. https://doi.org/10.5962/bhl.part.21148
Kirby, W.F. (1891) On the genus Xanthospilopteryx Wallengren. Transactions of the Entomological Society of London, 39, 272–292. https://doi.org/10.1111/j.1365-2311.1891.tb01651.x
Kirby, W.F. (1892) A Synonymic Catalogue of Lepidoptera Heterocera (Moths). Vol. 1. Gurney & Jackson c/o Taylor and Francis, London, 951 pp. https://doi.org/10.5962/bhl.title.9152
Kiriakoff, S.G. (1977) Lepidoptera Noctuiformes. Agaristidae II. Ethiopian and Madagascan Species. Das Tierreich, 98, 1–165. https://doi.org/10.1515/9783112420980
Lafontaine, J.D. & Mikkola, K. (1987) Lock–and–key system in the inner genitalia of Noctuidae (Lepidoptera) as taxonomic character. Entomologiske Meddelelser, 55, 161–167.
Linnaeus, C. (1764) Museum S:ae R:ae M:tis Ludovicae Ulricae Reginae Svecorum, Gothorum, Vandalorumque, &c. In quo animalia rariora, exotica, imprimis. Insecta & Conchilia describuntur & Determinatur. Laur. Salvius, Holmiae, [viii] + 720 pp. https://doi.org/10.5962/bhl.title.119811
Mabille, P. (1890) Sur les Phalaena Euphemia Cram. Annales de la Socieìteì entomologique de France, Series 6, 10, cxxii–cxxvi.
Mabille, P. (1891) La note synonymique et les descriptions (1re partie). Bulletin des seìances et bulletin bibliographique de la Socieìteì entomologique de France, 5, clxxxii–clxxxv.
Nye, I.W.B. (1975) The Generic names of moths of the world. Vol. 1. Noctuoidea (part): Noctuidae, Agaristidae & Nolidae. British Museum (Natural History), London, 568 pp. https://doi.org/10.5962/bhl.title.119777
Pagenstecher, A. (1893) Lepidopteren gesammelt in Ost-Africa 1888/89 von Dr. Franz Stuhlmann. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 10 (2), 207–262.
Poole, R.W. (1989) Noctuidae. In: Heppner, J.B. (Ed.), Lepidopterorum Catalogus. New Series. Fascicule 118. Parts 1–3. E.J. Brill, Leiden, pp. 1–500 + 501–1013 + 1015–1314.
Pinhey, E.C.G. (1979) Moths of Southern Africa. A.A. Balkema, Rotterdam, 273 pp.
Rothschild, M., Smith, D.A.S. & Pierre, J. (1975) Cardenolide storage in Danaus chrysippus (L.) with additional notes on D. plexippus (L.). Proceedings of the Royal Society of London B, Biological Sciences, 190 (1098), 1–31. https://doi.org/10.1098/rspb.1975.0076
Sharpe, E.M. (1906) Lepidoptera (Butterflies and Moths). In: Johnston, H.H. Liberia. Vol. 2. Hutchinson & Co., London, pp. 872–883.
Stoll, C. (1781) s.n. In: De Uitlandsche Kapellen voorkomende in de drie Waereld-deelen, Asia, Africa en America. Vol. IV. S. J. Baalde, Amsterdam, pp. 91–182, pls. CCCXXXVII–CCCLXXXIV.
Varga, Z. & Ronkay, L. (2013) Structural constraints of secondary asymmetry in male external genitalia of Noctuidae. Insect Systematics & Evolution, 44, 349–372. https://doi.org/10.1163/1876312X-04402001
Walker, F. (1854) List of the specimens of lepidopterous insects in the collection of the British Museum. Part 1. Printed by order of the Trustees, London, 278 pp.
Wallengren, H.D.J. (1858) Nya fjärilslägten. Öfversigt af Kongliga vetenskaps-akademiens förhandlingar, 15, 75–84.
Wallengren, H.D.J. (1865) Heterocer-Fjärilae samlade i Kafferlandet af J.A. Wahlberg. Kongliga Svenska Vetenskaps-akademiens handlingar, 5 (4), 1–83. https://doi.org/10.5962/bhl.title.13861
Wallengren, H.D.J. (1857) Kafferlandets Dag-fjärilar, insamlade åren 1838-1845 af J.A. Wahlberg. Kongliga Svenska Vetenskaps Akademiens Handlingar, 2 (4), 1–55.
Westwood, J. O. (1881) Appendix IV. Entomology. In: Oates, F., Matabele land and the Victoria Falls. C. Kegan Paul, London, pp. 331–365.
Zilli, A. & Grishin, N.V. (2019) Unveiling one of the rarest ‘butterflies’ ever (Lepidoptera: Hesperiidae, Noctuidae). Systematic Entomology, 44, 384–395. https://doi.org/10.1111/syen.12330
Zimsen, E. (1964) The type material of I.C. Fabricius. Munksgaard, Copenhagen, 656 pp.