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Published: 2021-10-12

Johnwellsia, a new intertidal genus of Parastenheliidae (Copepoda, Harpacticoida) from the Taiwan Strait, China, including a review of the family and key to genera

Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, U.K.
College of Marine Life Science, Ocean University of China, 5 Yushan Road, Qingdao 266003, China.
Crustacea Parastenhelia Microthalestris Thalestrella Paraleptomesochra Psammoleptomesochra Foweya Karaytugia gen. nov. Penicillicaris gen. nov. Porirualia gen. nov. sexual dimorphism identification keys species complex


A new genus of Parastenheliidae, Johnwellsia gen. nov., is proposed for its type and only species, J. bipartita sp. nov., collected from Dadeji Beach in Xiamen, Taiwan Strait, China. The intricate taxonomic history of the family is reviewed with special emphasis on its type genus Parastenhelia Thompson & Scott, 1903. It is concluded that P. hornelli Thompson & Scott, 1903 is the type of the genus and that the widely adopted previous designation of Harpacticus spinosus Fischer, 1860 as type species of Parastenhelia is invalid. The taxonomic concept of Parastenhelia is restricted to the hornelli­-group which includes four valid species: P. hornelli, P. similis Thompson & Scott, 1903, P. oligochaeta Wells & Rao, 1987, and P. willemvervoorti sp. nov.

                The currently accepted concept of Parastenhelia spinosa as a highly variable cosmopolitan species is rejected. The genus Microthalestris Sars, 1905 (type: Thalestris forficula Claus, 1863) is resurrected to accommodate most Parastenhelia species that were previously placed in the spinosa-group. Two species, Thalestris forficuloides Scott & Scott, 1894 and Parastenhelia antarctica Scott, 1912, are reinstated as valid members of the genus which further includes Parastenhelia gracilis Brady, 1910, Microthalestris littoralis Sars, 1911, P. costata Pallares, 1982, P. minuta Pallares, 1982, P. bulbosa Gee, 2006 and five new species: M. campbelliensis sp. nov.; M. polaris sp. nov.; M. santacruzensis sp. nov.; M. sarsi sp. nov. and M. variabilis sp. nov. Both the type species, Thalestris forficula, and Harpacticus spinosus are considered species inquirendae in Microthalestris. Three new genera are proposed to accommodate the remaining Parastenhelia species. Porirualia gen. nov. contains P. megarostrum Wells, Hicks & Coull, 1982 (type) and P. pyriformis Song, Kim & Chang, 2003, and is the sistergroup of Johnwellsia gen. nov. Parastenhelia aydini Kuru & Karaytuğ, 2015 is placed in the monotypic genus Karaytugia gen. nov. while all species with penicillate elements on the antenna and P1 are transferred to Penicillicaris gen. nov., including Thalestris pectinimana Car, 1884, which is removed from the synonyms of the Parastenhelia spinosa (Fischer, 1860) “complex”, and three new species: P. maldivensis sp. nov., P. penicillata sp. nov., and P. sewelli sp. nov.

                The genus Karllangia Noodt, 1964 (type: K. arenicola Noodt, 1964) is relegated to a junior subjective synonym of Thalestrella Monard, 1935a (type: T. ornatissima Monard, 1935a). New or updated diagnoses for each genus, and differential diagnoses for species where appropriate, are provided. A key to the ten currently recognized genera in the Parastenheliidae is presented as well as keys to species for Parastenhelia, Microthalestris, Thalestrella and Penicillicaris gen. nov.



  1. Alper, A. (2009) Datça-Bozburun özel çevre koruma bölgesi (Muğla, Türkiye) sahillerinin kumiçi ve fital Harpacticoida (Crustacea, Copepoda) faunasının belirlenmesi. Ph. D. dissertation, Balıkesir Üniversitesi, Balıkesir, xiii + 150 pp.
    Alper, A., Karaytuğ, S. & Sak, S. (2010) Interstitial and phytal Harpacticoida (Crustacea: Copepoda) inhabiting the mediolittoral zone of the Datça-Bozburun Peninsulas (Muğla, Turkey). SDU Journal of Science, 5, 16–28.
    Alper, A. & Sak, S. (2018) Sarımsaklı Sahili’nin (Balıkesir) Mediolittoral Bölgesinde Kumiçi Habitatta Yaşayan Harpaktikoid Kopepod (Crustacea, Copepoda, Harpacticoida) Faunasının Kalitatif ve Kantitatif Yönden İncelenmesi. [Qualitative and Quantitative Examination of the Harpacticoid Copepod (Crustacea, Copepoda, Harpacticoida) Fauna in the Mediolittoral Region of the Sarımsaklı Coast (Balıkesir)]. TÜBİTAK KBAG Project No. 215Z019, Progress Report, 133 pp. Available from: (accessed 15 August 2019) [in Turkish]
    Alper, A., Sönmez, S., Sak, S. & Karaytuğ, S. (2015) Marine harpacticoid (Copepoda, Harpacticoida) fauna of the Dilek Peninsula (Aydın, Turkey). Turkish Journal of Zoology, 39, 580–586.
    Anger, K. & Scheibel, W. (1976) Die benthische Copepodenfauna in einem ufernahen Verschmutzungsgebiet der westlichen Ostsee. Helgoländer wissenschaftlichen Meeresuntersuchungen, 28, 19–30.
    Apostolov, A. (1968) Neue und bemerkenswerte harpacticoide Ruderfußkrebse (Crustacea Copepoda) aus dem Küstengrundwasser Bulgariens. Zoologischer Anzeiger, 180, 395–402.
    Apostolov, A. (1973) Apport vers l’étude d’Harpacticoïdes pontiques habitant les algues marines. Zoologischer Anzeiger, 191, 263–281.
    Apostolov, A. (1975) Les Harpacticoïdes marins de la mer Noire. Description de quelques formes nouvelles. Vie et Milieu, (A), 25, 165–178.
    Apostolov, A. & Marinov, T.M. (1988) Copepoda Harpacticoida (morski kharpaktikoidi). [Copepoda, Harpacticoida (marine harpacticoids)]. Fauna Bolgarii [Fauna Bulgarica], 18, 1–384. [in Bulgarian]
    Apostolov, A. & Pandourski, I. (1999) Marine harpacticoids (Crustacea: Copepoda) from the littoral of the Livingston Island (The Antarctic). Bulgarian Antarctic Research, Life Sciences, 2, 68–82.
    Back, J. & Lee, W. (2013) Three new species of the genus Paramesochra T. Scott, 1892 (Copepoda: Harpacticoida: Paramesochridae) from Yellow Sea, Korea with a redescription of Paramesochra similis Kunz, 1936. Journal of natural History, 47, 769–803.
    Barroso, M.S., da Silva, B.J., Flores Montes, M.J. & Santos, P.J.P. (2018) Anthropogenic impacts on coral reef harpacticoid copepods. Diversity, 10, 32.
    Beckley, L.E. (1982) Studies on the littoral seaweed epifauna of St Croix Island. 3. Gelidium pristoides (Rhodophyta) and its epifauna. South African Journal of Zoology, 17, 3–10.
    Bick, A. & Arlt, G. (2013) Description of intertidal macro- and meiobenthic assemblages in Maxwell Bay, King George Island, South Shetland Islands, Southern Ocean. Polar Biology, 36, 673–689.
    Bigelow, H.B. (1926) Plankton of the offshore waters of the Gulf of Maine. Bulletin of the Bureau of Fisheries, Washington, D.C., 40 (2), 1–509.
    Blaschke, P., Woods, J. & Forsyth, F. (2010) The Porirua Harbour and its catchment: a literature summary and review. Report for Porirua City Council & Wellington City Council, 95 pp.
    Bodin, P. (1964) Recherches sur la systématique et la distribution des Copépodes Harpacticoïdes des substrats meubles des environs de Marseille. Recueil des Travaux de la Station marine d’Endoume, Faculté des Sciences de Marseille, 51 (= Bulletin 35), 107–183.
    Bodin, P. (1968) Copépodes Harpacticoïdes marins des substrats meubles des environs de la Rochelle. Note préliminaire. Recueil des Travaux de la Station marine d’Endoume, Faculté des Sciences de Marseille, 59 (= Bulletin 43), 279–283.
    Bodin, P. (1988) Results of ecological monitoring of three beaches polluted by the ‘Amoco Cadiz’ oil spill: development of meiofauna from 1978 to 1984. Marine Ecology Progress Series, 42, 105–123.
    Bodin, P. (1997) Catalogue of the new marine harpacticoid copepods (1997 edition). Documents de Travail, Institut royal des Sciences naturelles de Belgique, 89, 1–304.
    Bodin, P. & Boucher, D. (1983) Evolution à moyen terme du méiobenthos et des pigments chlorophylliens sur quelques plages polluées par la marée noire de l’«Amoco Cadiz». Oceanologica Acta, 6, 321–332.
    Boeck, A. (1865) Oversigt over de ved Norges Kyster iagttagne Copepoder henhørende til Calanidernes, Cyclopidernes og Harpactidernes Familier. Forhandlinger i Videnskabsselskabet i Kristiania, 1864, 226–282.
    Boeck, A. (1873) Nye Slægter og Arter af Saltvands-Copepoder. Forhandlinger i Videnskabsselskabet i Kristiania, 1872, 35–60.
    Božić, B. (1955) Copépodes Harpacticoïdes des sables des environs de Roscoff. Description de quelques formes nouvelles. Archives de Zoologie expérimentale et générale, 92 (Notes et Revue 1), 1–12.
    Brady, G.S. (1872) Contributions to the study of the Entomostraca. No. VII. A list of the non-parasitic marine Copepoda of the north-east coast of England. Annals and Magazine of natural History, Series 4, 10, 1–17, pls. II–VI. [also in Natural History Transactions of Northumberland and Durham, 4, 423–445, pls. XVII–XXI]
    Brady, G.S. (1899) On the marine Copepoda of New Zealand. Transactions of the zoological Society of London, 15 (2), 31–54, pls. IX–XIII.
    Brady, G.S. (1904) Notes on Entomostraca found at the roots of Laminariæ. Transactions of the Natural History Society of Northumberland, Durham and Newcastle-upon-Tyn (new series), 1, 3–9, pls. I–II.
    Brady, G.S. (1910) Die marinen Copepoden der Deutschen Südpolar Expedition 1901–1903. I. Ueber die Copepoden der Stämme Harpacticoida, Cyclopoida, Notodelphyoida und Caligoida. Deutsche Südpolar-Expedition, 11 (= Zoologie 3), 497–594.
    Bravender, B.A., Levings, C.D. & Brown, T.J. (1993) A comparison of meiofauna available as fish food on Sturgeon and Roberts Banks, Fraser River Estuary, British Columbia. Canadian technical Report of Fisheries and aquatic Sciences (= Rapport technique canadien des Sciences halieutiques et aquatiques), 1904, i–viii + 1–40.
    Brian, A. (1917) Descrizione di una nuova specie di Laophonte (L. quaterspinata n. sp. mihi) proveniente dai materiali del Laboratorio marino di Quarto. Monitore zoologico italiano, 28, 174–178.
    Brian, A. (1921) I Copepodi Harpacticoidi del Golfo di Genova. Stabilimento Tipografico-Litografico, R. Istituto Sordomuti, Genova, 112 pp.
    Brian, A. (1923a) Nota di Copepodi marini bentonici raccolti in Sardegna durante l’escursione organizzata dal Touringclub Italiano nel maggio 1921. Atti della Società italiana di Scienze naturali e del Museo civico di Storia natural, 62, 176–178.
    Brian, A. (1923b) Elenco di Copepodi marini bentonici provenienti da Rovigno e descrizione di una n. varietà di Parathalestris clausi Norm. Monitore zoologico italiano, 34, 126–135.
    Brian, A. (1928a) Descrizione di specie nuove o poco conosciute di Copepodi bentonici del mare Egeo. (Nota preliminare). Bolletino dei Musei e Laboratorii di Zoologia e Anatomia comparata della R. Università di Genova, Series 2, 7 (18), 1–37.
    Brian, A. (1928b) I Copepodi bentonici marini. Archivio zoologico italiano, 12, 293–343. [also in: Ghigi, A. (Ed.), Ricerche faunistiche nelle isole italiane dell’Egeo, 47–97]
    Brotskaya, V.A. (1962) Materialy po faune Harpacticoida (Crustacea, Copepoda) Velikoi salmy i prilezheshchikh uchastkov Belogo Morya. Contributions to the fauna of Harpacticoidea (Crustacea: Copepoda) of the Velilaya Salma Strait and of adjacent region of the White Sea. In: Zenkewitch, L.A. (Ed.), Biology of the White Sea. Trudy Belomorskoi Biologicheskoi Stantsii, M.G.U. (= Reports of the White Sea Biological Station of the State University of Moscow), 1, 109–129. [in Russian with English summary]
    Bruce, J.R., Colman, J.S. & Jones, N.S. (1963) Marine fauna of the Isle of Man. Liverpool Marine Biological Committee Memoirs on typical British marine Plants and Animals, 36, i–ix + 1–307.
    Callens, M., Gheerardyn, H., Ndaro, S.G.M., De Troch, M. & A. Vanreusel, A. (2012) Harpacticoid copepod colonization of coral fragments in a tropical reef lagoon (Zanzibar, Tanzania). Journal of the marine biological Association of the United Kingdom, 92, 1535–1545.
    Car, L. (1884) Ein Beitrag zur Copepoden-Fauna des adriatischen Meeres. Archiv für Naturgeschichte, 50, 237–256, pls. XVII–XVIII.
    Carlton, J.T., Chapman, J.W., Geller, J.B., Miller, J.W., Carlton, D.A., McCuller, M.I., Treneman, N.C., Steves, B.P. & Ruiz, G.M. (2017) Tsunami-driven rafting: Transoceanic species dispersal and implications for marine biogeography. Science, 357, 1402–1406.
    Carlton, J.T., Cordell, J.R., Hansen, G.I. & Pleus, A. (2013) Report on the first tsunami vessel, the Sou-you (壮 洋), washed ashore in North America with living Japanese species. Biofouling community on Japanese vessel 20-210-42909 washed ashore June 15, 2012 on Benson Beach at Cape Disappointment State Park, Ilwaco, Washington, 9 pp.
    Caspers, H. (1951) Quantitative Untersuchungen über die Bodentierwelt des Scharzen Meeres in bulgarischen Küstenbereich. Archiv für Hydrobiologie, 45, 1–192.
    Ceccherelli, V.U. & Mistri, M. (1990) Ecological and zoogeographical study of some Mediterranean associations of brackish water harpacticoids. Bolletino di Zoologia, 57, 73–81.
    Chadha, S. & Kar, C.S. (1999) s.n. In: Bhitarkanika: Myth and Reality. Nataraj Publishers, Dehradun, pp. v–xxxii + 1–388.
    Chapman, G. (1955) Aspects of the fauna and flora of the Azores. VI. The density of animal life in the coralline alga zone. Annals and Magazine of natural History, Series 12, 8, 801–805.
    Chapman, G. & Santler, J.E. (1955) Aspects of the fauna and flora of the Azores. V. Crustacea. Annals and Magazine of natural History, Series 12, 8, 371–376.
    Chertoprud, E.S., Azovsky, A. & Sapozhnikov, F.V. (2005) Colonization of azoic sediments of different grain-size composition by littoral Harpacticoida: Copepoda. Oceanology, 45, 698–706. [also published in Okeanologiya, 45, 737–746]
    Chertoprud, E.S., Frenkel, S.E., Kim, K. & Lee, W. (2015) Harpacticoida (Copepoda) of the northern East Sea (the Sea of Japan) and the southern Sea of Okhotsk: diversity, taxocenes, and biogeographical aspects. Journal of natural History, 49, 2869–2890.
    Chertoprud, E.S., Gómez, S. & Gheerardyn, H. (2009) Harpacticoida (Copepoda) fauna and the taxocene diversity of the South China Sea. Oceanology, 49, 488–498. [also published in Okeanologiya, 49, 532–542]
    Chislenko, L.L. (1967) Garpaktitsidy (Copepoda Harpacticoida) Karelskogo poberezh’ya Belogo Morya. Copepoda Harpacticoida of the Karelian coast of the White Sea. Issledovaniya Fauny Morei, 7 (15), 48–196. [in Russian]
    Chislenko, L.L. (1977) Garpaktitsidy (Copepoda, Harpacticoida) s gubok zemli Frantsa-Iosifa. Harpacticids (Copepoda Harpacticoidea) from sponges of Franz Josef Land. In: Biotsenosy shel’fa zemli Frantsa-Iosifa i fauna sopredel’nykh akuatorii. Issledovaniya Fauny Morei, 14 (22), 237–276. [in Russian with English summary]
    Claus, C. (1862) Untersuchungen über die Organisation und Verwandtschaft der Copepoden. (Im Auszuge zusammengestellt). Verhandlungen der physikalisch-medizinischen Gesellschaft zu Würzburg, 3, 51–103.
    Claus, C. (1863) Die freilebenden Copepoden mit besonderer Berücksichtigung der Fauna Deutschlands, der Nordsee und des Mittelmeeres. Wilhelm Engelmann, Leipzig, 230 pp., XXXVII pls.
    Claus, C. (1866. Die Copepoden-Fauna von Nizza. Ein Beitrag zur Charakteristik der Formen und deren Abänderungen “im Sinne Darwin’s”. Schriften der Gesellschaft zur Beförderung der gesammten Naturwissenschaften zu Marburg, Supplement 1, 1–34, pls. I–V.
    Colangelo, M.A., Bertasi, F., Dall’Olio P. & Ceccherelli, V.U. (2001) Meiofaunal biodiversity on hydrothermal seepage off Panarea (Aeolian Islands, Thyrrenian Sea). In: Farranda, F.M., Guglielmo, L. & Spezie, G. (Eds.), Mediterranean Ecosystems: Structures and Processes. Springer-Verlag Italia, Milano, pp. 353–359.
    Colman, J. (1940) On the faunas inhabiting intertidal seaweeds. Journal of the marine biological Association of the United Kingdom, New Series, 24, 129–183.
    Cordell, J.R. (2000) Chapter 9C5. Focal taxonomic collections: Copepoda Crustaceans. In: Hines, A.H., Ruiz, G.M., Chapman, K., Hansen, G.I., Carlton, J.T., Foster, N.R. & Feder, H.M. (Eds.), Biological Invasions of Cold-water Ecosystems: Ballast-mediated Introductions in Port Valdez/Prince William Sound, Alaska. Final Project Report, 2000, pp. 1–8.
    Cordell, J.R. (2018) Harpacticoid copepods associated with Japanese tsunami debris along the Pacific coast of North America. In: Carlton, J.T. & Fowler, A.E. (Eds.), Transoceanic Dispersal of Marine Life from Japan to North America and the Hawaiian Islands as a Result of the Japanese Earthquake and Tsunami of 2011. Aquatic Invasions, 13, pp. 113–124.
    Coull, B.C. (1970a) Harpacticoid copepods from Barbados and Jamaica, W.I., with descriptions of two new species. Caribbean Journal of Science, 10, 129–135.
    Coull, B.C. (1970b) Shallow water meiobenthos of the Bermuda platform. Oecologia, 4, 325–357.
    Coull, B.C. (1971a) Meiobenthic Harpacticoida (Crustacea, Copepoda) from St. Thomas, U.S. Virgin Islands. Transactions of the American microscopical Society, 90, 207–218.
    Coull, B.C. (1971b) Meiobenthic Harpacticoida (Crustacea, Copepoda) from the North Carolina continental shelf. Cahiers de Biologie marine, 12, 195–237.
    Coull, B.C. & Herman, S.S. (1970) Zoogeography and parallel level-bottom communities of the meiobenthic Harpacticoida (Crustacea, Copepoda) of Bermuda. Oecologia, 5, 392–399.
    Coull, B.C. & Wells, J.B.J. (1983) Refuges from fish predation: experiments with phytal meiofauna from the New Zealand rocky intertidal. Ecology, 64, 1599–1609.
    Cristoni, C., Colangelo, M.A. & Ceccherelli, V.U. (2004) Spatial scale and meiobenthic copepod recolonisation: testing the effect of disturbance size in a seagrass habitat. Journal of experimental marine Biology and Ecology, 298, 49–70.
    Crothers, J.H. (1966) Dale Fort Marine Fauna (2nd edition). Supplement to Field Studies Journal, 2, 1–169.
    Dahl, E. (1948) On the Smaller Arthropoda of Marine Algae, especially in the Polyhaline Waters off the Swedish West Coast. Undersökningar över Öresund XXXV. Ph. D. dissertation, University of Lund, 193 + 12 (appendix) pp.
    Dahms, H.-U. (1993) Copepodid development in Harpacticoida (Crustacea, Copepoda). Microfauna Marina, 8, 195–245.
    Dahms, H.-U. & Hicks, G.R.F. (1996) Naupliar development of Parastenhelia megarostrum (Copepoda: Harpacticoida), and its bearing on phylogenetic relationships. Journal of natural History, 30, 11–22.
    De Troch, M. (2001) Ecologie van Harpacticoide Copepoden: Structurele Biodiversiteit in Tropische Zeegrasvelden. Ecology of Harpacticoid Copepods: Structural Biodiversity in Tropical Seagrass Beds. Ph. D. dissertation, Universiteit Ghent, Ghent, 254 pp.
    Dev Roy, M.K. & Venkataraman, K. (2018) Catalogue on copepod fauna of India. Part 2. Cyclopoida and Harpacticoida (Arthropoda: Crustacea). Journal of Environment and Sociobiology, 15, 109–194.
    Dommasnes, A. (1969) On the fauna of Corallina officinalis L. in western Norway. Sarsia, 38, 71–86.
    Dvoretsky, V.G. & Dvoretsky, A.G. (2010) Checklist of fauna found in zooplankton samples from the Barents Sea. Polar Biology, 33, 991–1005.
    Farran, G.P. (1913) Marine Entomostraca. In: A biological survey of Clare Island in the county of Mayo, Ireland, and of the adjoining district. Proceedings of the royal Irish Academy, (B), 31 (45), 1–20, pls. I–III.
    Farran, G.P., Stelfox, A.W., Johnson, W.F., Halbert, J.N. & Tattersall, W.M. (1915) Addenda and corrigenda to section 2. In: A Biological Survey of Clare Island in the County of Mayo, Ireland, and of the Adjoining district. Proceedings of the royal Irish Academy, (A), 31 (Sections 1–3), 1911–1915, vii–xi.
    Fiers, F. (1990) Abscondicola humesi n. gen, n. sp. from the gill chambers of land crabs and the definition of the Cancrincolidae n. fam. (Copepoda, Harpacticoida). Bulletin de l’Institut royal des Sciences naturelles de Belgique, Biologie, 60, 69–103.
    Fischer, S. (1860) Beiträge zur Kenntnis der Entomostraceen. Abhandlungen der bayerischen Akademie der Wissenschaften, 8 (Abt. 3), 645–680, tabs. 1–3 (XX–XXII).
    Fraser, J.H. (1936) The distribution of rock pool Copepoda according to tidal level. Journal of animal Ecology, 5, 23–28.
    Fricke, H., Giere, O., Stetter, K., Alfredsson, G.A., Kristjansson, J.K., Stoffers, P. & Svavarsson, J. (1989) Hydrothermal vent communities at the shallow subpolar Mid-Atlantic ridge. Marine Biology, 102, 425–429.
    Geddes, D.C. (1972) The Copepoda Harpacticoida of Anglesey and the North Wales coast. The Naturalist, Hull, 921, 61–76.
    Gee, J.M. (2006) Parastenheliidae (Copepoda: Harpacticoida) from the Isles of Scilly. Journal of natural History, 40, 2611–2651.
    Gee, J.M. (2009) Some new and rare species of Ameiridae (Copepoda: Harpacticoida) from the Isles of Scilly, UK. Journal of natural History, 43, 2809–2851.
    Gee, J.M. & Warwick, R.M. (1994) Metazoan community structure in relation to the fractal dimensions of marine macroalgae. Marine Ecology Progress Series, 103, 141–150.
    Gheerardyn, H. (2007) Biodiversity and Taxonomy of Harpacticoid Copepods Associated with Coral Substrates of Tropics and Deep Sea. Biodiversiteit en Taxonomie van Harpacticoide Copepoden Geassocieerd met Koraalsubstraten van Tropen en Diepzee. Ph. D. Dissertation, University of Ghent, Ghent, xxvi + 251 pp.
    Gheerardyn, H., De Troch, M., Ndaro, S.G.M., Raes, M., Vincx, M. & Vanreusel, A. (2008) Community structure and microhabitat preferences of harpacticoid copepods in a tropical reef lagoon (Zanzibar Island, Tanzania). Journal of the marine biological Association of the United Kingdom, 88, 747–758.
    Gillman, R. (2018) Ecological Consequences of Marine Debris: Understanding Large-Scale Species Transport on Tsunami Debris and Research Priorities in Oregon. M. Sc. dissertation, University of Oregon, Eugene, Oregon, 114 pp.
    Grandori, R. (1912) I Copepodi. In: Carazzi, D.C. & Grandori, R. (Eds.), Ricerche sul Plancton della Laguna Veneta. Premiata Società Coop. Tipografica, Padova, pp. 1–40, foldout.
    Grandori, R. (1914) Studi biologici sul plancton della Laguna Veneta e de l’Alto Adriatico. Atti dell’Accademia scientifica veneto-trentino-istriana, Series 3, 6, 149–171.
    Gunnill, F.C. (1982) Effects of plant size and distribution on the numbers of invertebrate species and individuals inhabiting the brown alga Pelvetia fastigiata. Marine Biology, 69, 263–280.
    Gunnil, F.C. (1983) Seasonal variations in the invertebrate faunas of Pelvetia fastigiata (Fucacea): effects of plant size and distribution. Marine Biology, 73, 115–130.
    Gurney, R. (1927) XXXIII. Report on the Crustacea:—Copepoda (littoral and semi-parasitic). Zoological results of the Cambridge expedition to the Suez Canal, 1924. Transactions of the zoological Society of London, 22, 451–577, figs. 108–168, tab. 35.
    Haas, M.E., Simenstad, C.A., Cordell, J.R., Beauchamp, D.A. & Miller, B.S. (2002) Effects of large overwater structures on epibenthic juvenile salmon prey assemblages in Puget Sound, Washington. Final Research Report prepared for Washington State Transportation Commission, Department of Transportation and in cooperation with U.S. Department of Transportation, Federal Highway Administration. Published by Washington State Transportation Center (TRAC), University of Washington, Seattle, Washington, 114 pp.
    Hagerman, L. (1966) The macro- and microfauna associated with Fucus serratus L., with some ecological remarks Ophelia, 3, 1–43.
    Hartzband, D.J. & Hummon, W.D. (1973) Sub-community structure in subtidal meiobenthic Harpacticoida. Oecologia, 14, 37–51.
    Herdman, W.A. (1897) Tenth annual report of the Liverpool Marine Biology Committee and their Biological Station at Port Erin. Proceedings and Transactions of the Liverpool biological Society, 11, 7–56.
    Hicks, G.R.F. (1980) Structure of phytal harpacticoid copepod assemblages and the influence of habitat complexity and turbidity. Journal of experimental marine Biology and Ecology, 44, 157–192.
    Hicks, G.R.F. (1984) Spatio‑temporal dynamics of a meiobenthic copepod and the impact of predation‑disturbance. Journal of experimental marine Biology and Ecology, 81, 47–72.
    Hicks, G.R.F. (1985) Biomass and production estimates for an estuarine meiobenthic copepod, with an instantaneous assessment of exploitation by flatfish predators. New Zealand Journal of Ecology, 8, 125–127.
    Hicks, G.R.F. (1986) Distribution and behaviour of meiofaunal copepods inside and outside seagrass beds. Marine Ecology Progress Series, 31, 159–170.
    Hicks, G.R.F. (1988) Sediment rafting: a novel mechanism from small-scale dispersal of intertidal estuarine meiofauna. Marine Ecology Progress Series, 48, 69–80.
    Hicks, G.R.F. (1989) Does epibenthic structure negatively affect meiofauna? Journal of experimental marine Biology and Ecology, 133, 39–55.
    Hicks, G.R.F. (1991) Monitoring with meiofauna: a compelling option for evaluating environmental stress in tidal inlets. In: Coastal Engineering—Climate for Change, Proceedings of the 10th Australasian Conference on Coastal and Ocean Engineering, December 2–6, 1991. Water Quality Centre, DSIR Marine and Freshwater, Hamilton, New Zealand. Water Quality Centre Publications, 21, 387–391.
    Hicks, G.R.F. (1992) Tidal and diel fluctuations in abundance of meiobenthic copepods on an intertidal estuarine sandbank. Marine Ecology Progress Series, 87, 15–21.
    Hines, A.H. & Ruiz, V. (2001) Marine invasive species and biodiversity of South Central Alaska. Report submitted to Regional Citizens’ Advisory Council of Prince William Sound, Anchorage, Alaska, 75 pp.
    Huys, R. (1987) Some morphological observations on the Neobradyidae Olofsson, 1917 (Copepoda, Harpacticoida) including the redescription of Antarcticobradya tenuis (Brady, 1910) comb. nov. Bulletin de l’Institut royal des Sciences naturelles de Belgique, Biologie, 57, 133–148.
    Huys, R. (1990) A new harpacticoid copepod family collected from Australian sponges and the status of the subfamily Rhynchothalestrinae Lang. Zoological Journal of the Linnean Society, 99, 51–115.
    Huys, R. (2009) Unresolved cases of type fixation, synonymy and homonymy in harpacticoid copepod nomenclature (Crustacea: Copepoda). Zootaxa, 2183 (1), 1–99.
    Huys, R. (2016) Harpacticoid copepods—their symbiotic associations and biogenic substrata: a review. In: Huys, R. (Ed.), Recent Developments in Taxonomy and Biodiversity of Symbiotic Copepoda (Crustacea)—A Volume in Celebration of the Career of Prof. Il-Hoi Kim. Zootaxa, 4174, 448–729.
    Huys, R. & Boxshall, G.A. (1991) Copepod Evolution. The Ray Society, London, 468 pp.
    Huys, R., Gee, J.M., Moore, C.G. & Hamond, R. (1996) Marine and Brackish Water Harpacticoids, Part 1. In: Barnes, R.S.K. & Crothers, J.H. (Eds.), Synopses of the British Fauna, New Series, 51, 1–352.
    Huys, R., MacKenzie-Dodds, J. & Llewellyn-Hughes, J. (2009) Cancrincolidae (Copepoda, Harpacticoida) associated with land crabs: a semiterrestrial leaf of the ameirid tree. Molecular Phylogenetics and Evolution, 51, 143–156.
    ICZN [International Commission on Zoological Nomenclature] (1999) International Code of Zoological Nomenclature. 4th Edition. The International Trust for Zoological Nomenclature, London, xxx + 306 pp.
    Iwasaki, N. (1993) Distribution of meiobenthic copepods from various habitats in Pauatahanui Inlet, New Zealand. New Zealand Journal of marine and freshwater Research, 27, 399–405.
    Iwasaki, N. (1994) Preliminary investigation of meiobenthic harpacticoid copepods on Heron Reef, Great Barrier Reef (Extended Abstract). Bulletin of marine Sciences and Fisheries, Kochi University, 14, 43–45.
    Iwasaki, N. (1995a) Short-term variation of meiobenthic harpacticoid copepods in a sea bream nursery pond: effects of environmental changes and predation. Benthos Research, 48, 1–12.
    Iwasaki, N. (1995b) Small-scale spatial patterns and intra- and interspecific relationships of meiobenthic harpacticoid copepods. Benthos Research, 49, 39–50.
    Iwasaki, N. (1999) Horizontal zonation of meiobenthic copepods in the Pauatahanui Inlet, New Zealand. New Zealand Natural Sciences, 24, 89–99.
    Jakubisiak, S. (1933) Notatka o Harpaktikoidach Adrjatyku. Notes sur les Harpacticoïdes de l’Adriatique (île de Šolta, Dalmatie). Fragmenta faunistica, 2, 113–116. [in French with Polish summary]
    Jakubisiak, S. (1936) Materialy do fauny Harpaktikoidów okolic Roscoff (brzegi bretońskie Francji). Matériaux à la faune des Harpacticoïdes de Roscoff (côtes bretonnes, France). Fragmenta faunistica, 2, 315–321. [in French with Polish summary]
    Jarvis, S.C. & Seed, R. (1996) The meiofauna of Ascophyllum nodosum (L.) Le Jolis: characterization of the assemblages associated with two common epiphytes. Journal of experimental marine Biology and Ecology, 199, 249–267.
    Jayabarathi, R. (2016) Spatial Heterogeneity and Biodiversity of Harpacticoid Copepods and other Meiofaunal Associates within the Seagrass Patches of South Andaman. Ph. D. dissertation, Pondicherry University, Port Blair, vii + 205 pp.
    Jayabarathi, R., Padmavati, G. & Anandavelu, I. (2012) Abundance and species composition of harpacticoid copepods from a sea grass patch of South Andaman, India. Current Research Journal of biological Sciences, 4, 717–724.
    Jayabarathi, R., Padmavati, G. & Anandavelu, I. (2015) Spatial heterogeneity of benthic copepods: a comparative aspect on composition, abundance, and correlation. Zoological Studies, 54, 51.
    Jayalakshmi, T., Nandakumar, R., Balaji Prasath, B. & Santhanam, P. (2016) Effect of acidification on fatty acids profiling of marine benthic harpacticoid copepod Parastenhelia sp. Annals of agrarian Science, 14, 278–282.
    Jenkins, G.P., Walker-Smith, G.K. & Hamer, P.A. (2002) Elements of habitat complexity that influence harpacticoid copepods associated with seagrass beds in a temperate bay. Oecologia, 131, 598–605.
    Jespersen, P. (1940) Non-parasitic Copepoda. In: Friðriksson, A. & Tuxen, S. (Eds.), The Zoology of Iceland, 3 (33), pp. 1–116.
    Johansen, J.A. & Reis, K.E.M. (1994) Fraser River Basin Benthic Invertebrate Catalogue: Continental and Oceanographic Data Information System (CODIS). Fraser River Action Plan (Canada). Environment Canada, Pacific and Yukon Region, iii + 104 pp.
    Johnson, S.C. & Scheibling, R.E. (1986) Species compostion and abundance of harpacticoid copepods on intertidal macroalgae, Fucus vesiculosus and Ascophyllum nodosum, off Nova Scotia, Canada. In: Schriever, G., Schminke, H.K. & Shih, C.-T. (Eds.), Proceedings II. International Conference on Copepoda, Ottawa, Canada, 13–17 August, 1984. Syllogeus, 58, pp. 567–574.
    Johnson, S.C. & Scheibling, R.E. (1987a) Structure and dynamics of epifaunal assemblages on intertidal macroalgae Ascophyllum nodosum and Fucus vesiculosus in Nova Scotia, Canada. Marine Ecology Progress Series, 37, 209–227.
    Johnson, S.C. & Scheibling, R.E. (1987b) Reproductive patterns of harpacticoid copepods on intertidal macroalgae (Ascophyllum nodosum and Fucus vesiculosus) in Nova Scotia, Canada. Canadian Journal of Zoology, 65, 129–141.
    Karanovic, T. (2004) Subterranean copepods (Crustacea, Copepoda) from arid Western Australia. Crustaceana Monographs, 3, 1–366.
    Karaytuğ, S. & Sak, S. (2006) A contribution to the marine harpacticoid (Crustacea, Copepoda) fauna of Turkey. E.U. Journal of Fisheries and aquatic Sciences (E.Ü. Su Ürünleri Dergisi), 23, 403–405.
    Kask, B.A., Sibert, J.R. & Windecker, B. (1982) A check list of marine and brackish water harpacticoid copepods from the Nanaimo estuary, southwestern British-Columbia. Syesis, 15, 25–38.
    Kim, D.-S. & Shirayama, Y. (1996) Seasonal fluctuation of meiobenthos inhabiting subtidal coarse sand in Otsuchi Bay, with special reference to free-living nematodes. Benthos Research, 51, 9–20.
    Klie, W. (1929) Die Copepoda Harpacticoida der südlichen und westlichen Ostsee mit besonderer Berücksichtigung der Sandfauna der Kieler Bucht. Zoologische Jahrbücher, Systematik, 57, 329–386.
    Klie, W. (1941) Marine Harpacticoiden von Island. Kieler Meeresforschungen, 5, 1–44.
    Klie, W. (1949) Harpacticoida (Cop.) aus dem Bereich von Helgoland und der Kieler Bucht. 1. Kieler Meeresforschungen, 6, 1–40.
    Kornev, P.N. & Chertoprud, E.S. (2008) Veslonogie rakoobraznye otryada Harpacticoida Belogo Morya: morfologiya, sistematika, ecologiya. [Copepod Crustaceans of the Order Harpacticoida of the White Sea: Morphology, Systematics, Ecology]. Tovarischchestvo Nauchnikh Izdanii KMK, Moscow, 379 pp.
    Krishna Murty, P.V.M. (1983) Distribution of phytal harpacticoid copepods along Visakhapatnam Coast. Mahasagar – Bulletin of the national Institute of Oceanography, 16, 47–54.
    Krishnaswamy, S. (1953) Pelagic Copepoda of the Madras coast. Journal of the zoological Society of India, 5, 64–75.
    Krishnaswamy, S. (1957) Studies on the Copepoda of Madras. Ph. D. dissertation, University of Madras, Chennai, 168 pp.
    Kunz, H. (1935) Zur Oekologie der Copepoden Schleswig-Holsteins und der Kieler Bucht. Schriften des naturwissenschaftlichen Vereins für Schleswig-Holstein, 21, 84–132.
    Kunz, H. (1951) Marine Harpacticoiden aus dem Küstensand von Südwestafrika. Kieler Meeresforschungen, 8, 76–81, pls. 23–26.
    Kunz, H. (1963) Weitere Harpacticoiden (Crustacea Copepoda) von der südwestafrikanischen Küste. Zoologischer Anzeiger, 171, 33–51
    Kunz, H. (1975) Copepoda Harpacticoida aus dem Litoral des südlichen Afrika. I. Teil. Kieler Meeresforschungen, 31, 179–212, pls. 1–30 (unpaginated).
    Kuru, S. & Karaytuğ, S. (2015) A new species of Parastenhelia Thompson & A. Scott, 1903 (Copepoda, Harpacticoida, Parastenheliidae) from Turkey. Biharean Biologist, 9, 121–127.
    Lang, K. (1934) Marine Harpacticiden von der Campbell-Insel und einigen anderen südlichen Inseln. Acta Universitatis lundensis, New Series, Avd. 2, 30 (14), 1–56.
    Lang, K. (1936a) Copepoda Harpacticoida. Further zoological Results of the Swedish Antarctic Expedition 1901–1903 under the Direction of Dr Otto Nordenskjöld, 3 (3), 1–68.
    Lang, K. (1936b) Undersökningar över Öresund. Untersuchungen aus dem Öresund. XX. Harpacticiden aus dem Öresund. Acta Universitatis lundensis, New Series, Avd. 2, 31 (8), 1–52.
    Lang, K. (1944) Monographie der Harpacticiden (Vorläufige Mitteilung). Almqvist & Wiksells Boktryckeri Ab, Uppsala, 39 pp.
    Lang, K. (1948) Monographie der Harpacticiden. Vols. 1–2. Håkan Ohlsson, Lund, 1682 pp.
    Lang, K. (1965) Copepoda Harpacticoida from the Californian Pacific coast. Kungliga Svenska Vetenskapsakademiens Handlingar, Seris 4, 10 (2), 1–560.
    Leduc, D. & Probert, P.K. (2011) Small-scale effect of intertidal seagrass (Zostera muelleri) on meiofaunal abundance, biomass, and nematode community structure. Journal of the marine biological Association of the United Kingdom, 91, 579–591.
    Leduc, D., Probert, P.K. & Duncan, A. (2009) A multi-method approach for identifying meiofaunal trophic connections. Marine Ecology Progress Series, 383, 95–111.
    Le Guellec, C. (1988) Colonisation d’un sable azoïque exogène par la méiofaune. Cahiers de Biologie marine, 29, 469–481.
    Lintas, C. & Seed, R. (1994) Spatial variation in the fauna associated with Mytilus edulis on a wave-exposed rocky shore. Journal of molluscan Studies, 60, 65–174.
    Mahadi, M., Sazlina, S., Aileen, T.S.H. & Zulfigar, Y. (2014) Benthic harpacticoid copepod assemblages in Merambong shoal, Johor. Malayan Nature Journal, 66, 52–58.
    Marcus, A. & Pór, F. (1960) Die Copepoden einer Probe aus dem Felsbiotop von Yalta (Krimhalbinsel). Travaux du Muséum d’Histoire naturelle “Gr. Antipa”, 2, 145–163.
    Marine Biological Association (1931) Plymouth Marine Fauna, Second Edition. Marine Biological Association, Plymouth, 371 pp., 1 chart.
    Marine Biological Association (1957) Plymouth Marine Fauna. 3rd Edition. Marine Biological Association, Plymouth, 457 pp., charts 1–4.
    Marinov, T.M. (1974) Dop’lnenie k’m izuchavaneto na kharpaktikoidnata fauna ot b’garskoto Cherno morsko kraibrezhie. Supplement to the study of the harpacticoid fauna from the Bulgarian Black Sea coast. Izvestiya na Instituta Okeanografiya i Ribno Stopanstvo, Varna, 13, 77–92. [in Bulgarian with English and Russian summaries]
    Mielke, W. (1974) Eulitorale Harpacticoidea (Copepoda) von Spitzbergen. Mikrofauna des Meeresbodens, 37, 1–52
    Mielke, W. (1990) A Parastenhelia species from Bahía Lapataia, Ushuaia, Tierra del Fuego (Argentina). Microfauna marina, 6, 157–167.
    Mielke, W. (1994a) A new interstitial copepod species related to the “Leptomesochra complex” (Copepoda, Ameiridae) from Chile. Microfauna marina, 9, 251–259.
    Mielke, W. (1994b) Two co-occuring new Karllangia species (Copepoda: Ameiridae) from the Caribbean coast of Costa Rica. Revista de Biología tropical, 42, 141–153.
    Mielke, W. (1997) Interstitial Fauna of Galapagos. XL. Copepoda, part 8. Microfauna Marina, 11, 153–192.
    Mielke, W. (2003) Meiobenthic copepods of Galápagos: species composition, distribution, colonization, speciation. Bulletin of Marine Science, 73, 557–568.
    Miklucho-Maclay, N.N. (1870) Über einige Schwämme des nördlichen Stillen Oceans und des Eismeeres, welche im Zoologischen Museum der Kaiserlichen Akademie der Wissenschaften in St. Petersburg aufgestellt sind. Ein Beitrag zur Morphologie und Verbreitung der Spongien. Mémoires de l’Académie impériale des Sciences de St. Petersbourg, 15 (3), 1–24.
    Miller, J.A., Gillman, R., Carlton, J.T., Clarke Murray, C., Nelson, J.C., Otani, M. & Ruiz, G.M. (2018) Trait-based characterization of species transported on Japanese tsunami marine debris: Effect of prior invasion history on trait distribution. Marine Pollution Bulletin, 132, 90–101.
    Monard, A. (1927) Synopsis universalis generum harpacticoidarum. Zoologische Jahrbücher für Systematik, 54, 139–176.
    Monard, A. (1928) Les Harpacticoïdes marins de Banyuls. Archives de Zoologie expérimentale et générale, 67, 259–443.
    Monard, A. (1935a) Étude sur la faune des Harpacticoïdes marins de Roscoff. Travaux de la Station biologique de Roscoff, 13, 5–88, figs. 1–235.
    Monard, A. (1935b) Les Harpacticoïdes marins de la région de Salammbô. Bulletin de la Station océanographique de Salammbô, 34, 1–94.
    Monard, A. (1936) Note préliminaire sur la faune des Harpacticoïdes marins d’Alger. Bulletin des travaux publiés par la Station d’Aquiculture et de Pêche de Castiglione, 1935 (1), 45–85.
    Monard, A. (1937) Les Harpacticoïdes marins de la région d’Alger et de Castiglione. Bulletin des travaux publiés par la Station d’Aquiculture et de Pêche de Castiglione, 1935 (2), 9–93.
    Moore, H.B. (1932) Faunistic records for the Manx region. In: Johnstone, J. & Daniel, R.J. (Eds.), Report for 1931 (no. 40) on the Lancashire Sea-Fisheries Laboratory at the University of Liverpool and the Annual Report of the Marine Biological Station (no. 45) at Port Erin. Proceedings and Transactions of the Liverpool biological Society, 46, appendix, pp. 152–159.
    Moore, P.G. (1973) The kelp fauna of northeast Britain. II. Multivariate classification: turbidity as an ecological factor. Journal of experimental marine Biology and Ecology, 13, 127–164.
    Moore, P.G. (1974) The kelp fauna of northeast Britain. III. Qualitative and quantitative ordinations, and the utility of a multivariate approach. Journal of experimental marine Biology and Ecology, 16, 257–300.
    Nicholls, A.G. (1945) Marine Copepoda from Western Australia. III.—Littoral harpacticoids from Port Denison. Journal of the Royal Society of Western Australia, 29, 1–16.
    Noli, N., Sbrocca, C., Sandulli, R., Balsamo, M. & Semprucci, F. (2018) Contribution to the knowledge of meiobenthic Copepoda (Crustacea) from the Sardinian coast, Italy. Arxius de Miscel·lània Zoològica, 16, 121–133.
    Noodt, W. (1955a) Marmara denizi Harpacticoid’leri (Crust. Cop.). [Marine Harpacticoiden (Crust. Cop.) aus dem Marmara Meer]. Istanbul Universitesi Fen Fakültesi Mecmuasi, (B), 20 (1–2), 49–94.
    Noodt, W. (1955b) Copepoda Harpacticoidea von Teneriffa (Kanarische Inseln). Zoologischer Anzeiger, 154, 200–222.
    Noodt, W. (1964) Copepoda Harpacticoidea aus dem Litoral des Roten Meeres. Kieler Meeresforschungen, 20, Sonderheft, 128–154.
    Norman, A.M. & Brady, G.S. (1909) The Crustacea of Northumberland and Durham. Transactions of the natural History Society of Northumberland, Durham and Newcastle-upon-Tyne, New Series, 3, 252–417, pls. VIII–IX, IX(A).
    Norman, A.M. & Scott, T. (1905) Crustacea Copepoda new to Science from Devon and Cornwall. Annals and Magazine of natural History, Series 7, 15, 284–300.
    Norman, A.M. & Scott, T. (1906) The Crustacea of Devon and Cornwall. William Wesley & Son, London, xv + 232 pp., pls. I–XXIV.
    Ólafsson, E., Ingólfsson, A. & Steinarsdóttir, M.B. (2001) Harpacticoid copepod communities of floating seaweed: controlling factors and implications for dispersal. Hydrobiologia, 453, 189–200.
    Packmor, J. & George, K.H. (2018) Littoral Harpacticoida (Crustacea: Copepoda) of Madeira and Porto Santo (Portugal). Journal of the marine biological Association of the United Kingdom, 98, Special Issue 1 (Special Section: European Marine Biology Symposium Papers 2016), 171–182.
    Pallares, R.E. (1963) Los Copépodos planctónicos marinos de Puerto Deseado (Santa Cruz, Argentina). Ph. D. dissertation, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, 104 pp., 58 unpaginated pls.
    Pallares, R.E. (1968) Copépodos marinos de la Ría Deseado (Santa Cruz, Argentina). Contribución sistemática ecológica. I. Contribución científica del Centro de Investigación de Biología marina, Buenos Aires, 27, 1–125.
    Pallares, R.E. (1975) Copepodos harpacticoides marinos de Tierra del Fuego (Argentina) 1. Isla de Los Estados. Contribución científica del Centro de Investigación de Biología marina, Buenos Aires, 122, 1–35.
    Pallares, R.E. (1982) Copépodos harpacticoides marinos de Tierra del Fuego (Argentina). IV. Bahía Thetis. Contribución científica del Centro de Investigación de Biología marina, Buenos Aires, 186, 3–39, pls. I–XII.
    Pearse, A.S. (1934) VII. Inhabitants of certain sponges at Dry Tortugas. Papers from the Tortugas Laboratory of Carnegie Institution of Washington, 28, 117–124, pls. 1–2.
    Pesce, G.L. & Pandourski, I. (2002) Pseudocyclopina livingstoni sp. n. (Copepoda, Cyclopinidae) from the Livingston Island (Antarctica). Biologia, Bratislava, 57, 133–137.
    Pesta, O. (1920) Die Planktoncopepoden der Adria. Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere, 43, 471–660.
    Pesta, O. (1959) Harpacticoiden (Crust. Copepoda) aus submarinen Höhlen und den benachbarten Litoralbezirken am Kap von Sorrent (Neapel). Ergebnisse der österreichischen Tyrrhenia Expedition 1952. Teil: VI. Pubblicazioni della Stazione zoologica di Napoli, 30 (Supplement), 94–177.
    Petkovski, T.K. (1955) Zweiter Beitrag zur Kenntnis der Harpacticidenfauna unserer Meeresküste. Fragmenta balcanica, 1, 125–139.
    Pintea, M. & Klemens, K. (1957) Harpacticides collectionnés des touffes de Callithamnion de la zone littorale d’Agigea. Annales scientifiques de l’Université de Jassy, 1957. [unknown pagination]
    Pongener, L., Padmavati, G. & Jayabarathi, R. (2018) Meiofauna and microalgae associates on the pneumatophores of Avicennia marina from the coastal waters of South Andaman, India. International Journal of zoological Studies, 3, 203–212.
    Pór, F.D. (1960) Littorale Harpacticoiden der Nordwest-Küsten des Schwarzen Meeres. Travaux du Muséum d’Histoire naturelle “Gr. Antipa”, 2, 97–143.
    Por, F.D. (1964) A study of Levantine and Pontic Harpacticoida (Crustacea, Copepoda). Zoölogische Verhandelingen, Leiden, 64, 1–128.
    Preston, A. & Moore, P.G. (1988) The flora and fauna associated with Cladophora albida (Huds.) Kütz from rockpools on Great Cumbrae Island, Scotland. Ophelia, 29, 169–186.
    Preston, A. & Moore, P.G. (1990) Observations on the feeding habits of the commoner animals associated with Cladophora albida (Huds.) Kütz in rockpools. The Glasgow Naturalist, 21, 523–535.
    Rao, G.C. (1972) Some new interstitial harpacticoid copepods from Andhra coast, India. Cahiers de Biologie marine, 13, 305–319.
    Rao, G.C. (1975) The interstitial fauna in the intertidal sands of Andaman and Nicobar group of islands. Journal of the marine biological Association of India, 17, 116–128.
    Rao, G.C. (1980) On the zoogeography of the interstitial meiofauna of the Andaman and Nicobar Islands, Indian Ocean. Records of the Zoological Survey of India, 77, 153–178.
    Rao, G.C. (1989) Intertidal meiofauna. In: Jairajpuri, M.S. (Ed.), Fauna of Orissa. Part–II. State Fauna Series 2. Zoological Survey of India, Calcutta, pp. 1–77.
    Rao, G.C. (1991) Meiofauna. In: Jairajpuri, M.S. (Ed.), Fauna of Lakshadweep. State Fauna Series, 2. Zoological Survey of India, Calcutta, pp. 41–135.
    Rao, G.C. (1993) Littoral meiofauna of Little Andaman. Records of the Zoological Survey of India, Occasional Papers, 155, 1–120.
    Rao, G.C. & Misra, A. (1983) Meiofauna from Lakshadweep, Indian Ocean. Cahiers de Biologie marine, 24, 51–68.
    Renaud-Debyser, J. (1963a) Recherches écologiques sur la faune interstitielle des sables du Bassin d’Arcachon. Procès verbaux de la Société linnéenne de Bordeaux, 100, 75–82.
    Renaud-Debyser, J. (1963b) Recherches écologiques sur la faune interstitielle des sables. Bassin d’Arcachon – île de Bimini, Bahamas. Vie et Milieu, Supplement 15, 1–157.
    Roe, K.M. (1958) The littoral harpacticids of the Dalkey (Co. Dublin) area with descriptions of six new species. Proceedings of the Royal Irish Academy, (B), 59, 221–255.
    Roe, K.M. (1960) Some harpacticids from Lough Ine, with descriptions of two new species. Proceedings of the Royal Irish Academy, (B), 60, 277–289.
    Ruiz, G.M., Huber, T., Larson, K., McCann, L., Steves, B., Fofonoff, P. & Hines, A.H. (2006) Biological Invasions in Alaska’s Coastal Marine Ecosystems: Establishing a Baseline Final Report Submitted to Prince William Sound Regional Citizens’ Advisory Council & U.S. Fish & Wildlife Service. Smithsonian Environmental Research Center, Edgewater, Maryland, 112 pp.
    Sabater, F. (1986) Some interstitial species of the crustacean communities of the Ter and Ebre rivermouths (NE Spain). Miscellania Zoologica, Barcelona, 10, 113–119.
    Sarma, A.L.N. (1974a) The phytal fauna of Ulva fasciata off Visakhapatnam coast. Proceedings of the Indian Academy of Sciences, (B), 80, 147–161.
    Sarma, A.L.N. (1974b) Phytal fauna of Caulerpa taxifolia and C. racemosa off Visakhapatnam coast. Indian Journal of marine Science, 3, 155–164.
    Sarma, A.L.N. (1974c) The phytal fauna of Sargassum off Visakhapatnam coast. Journal of the marine biological Association of India, 16, 741–755.
    Sarma, A.L.N. & Ganapati, P.N. (1975) Meiofauna of the Visakhapatnam harbour in relation to pollution. Bulletin of the Department of marine Sciences, University of Cochin, 7, 243–255.
    Sarmento, V.C., Lage, L.M. & Santos, P.J.P. (2009) Distribuição espaço—temporal de Copepoda Harpacticoida em costão sob influência de ressurgência (Arraial do Cabo, RJ). Anais do III Congresso Latino Americano de Ecologia, 10 a 13 de Setembro de 2009, São Lourenço—MG, pp. 1–4.
    Sarmento, V.C., Lage, L.M. & Santos, P.J.P. (2012) Copepoda Harpacticoida community of a rocky shore under the influence of upwelling (Arraial do Cabo, southeastern Brazil). Journal of the marine biological Association of the United Kingdom, 92, 1117–1126.
    Sarmento, V.C. & Santos, P.J.P. (2012a) Trampling on coral reefs: tourism effects on harpacticoid copepods. Coral Reefs, 31, 135–146.
    Sarmento, V.C. & Santos, P.J.P. (2012b) Species of Harpacticoida (Crustacea, Copepoda) from the phytal of Porto de Galinhas coral reefs, northeastern Brazil. Check List, 8, 936–939.
    Sars, G.O. (1903) Copepoda Harpacticoida. Parts I & II, Misophriidæ, Longipediidæ, Cerviniidæ, Ectinosomidæ (part). An Account of the Crustacea of Norway, with short Descriptions and Figures of all the Species, 5, 1–28, pls. I–XVI.
    Sars, G.O. (1905) Copepoda Harpacticoida. Parts IX & X. Thalestridae (continued). An Account of the Crustacea of Norway, with short Descriptions and Figures of all the Species, 5, 109–132, pls. LXV–LXXX.
    Sars, G.O. (1909) Crustacea. Report of the second Norwegian Arctic Expedition in the “Fram” 1898–1902, 18, 1–47, pls. I–XII.
    Sars, G.O. (1911) Copepoda Harpacticoida. Parts XXXI & XXXII. Supplement (continued). An Account of the Crustacea of Norway, with short Descriptions and Figures of all the Species, 5, 369–396, supplement pls. 11–26.
    Schander, C., Rapp, H.T., Kongsrud, J.A., Bakken, T., Berge, J., Cochrane, S., Oug, E., Byrkjedal, I., Todt, C., Cedhagen, T., Fosshagen, A., Gebruk, A., Larsen, K., Levin, L., Obst, M., Pleijel, F., Stöhr, S., Warén, A., Mikkelsen, N.T., Hadler-Jacobsen, S., Keuning, R., Heggøy Petersen, K., Thorseth, I.H. & Pedersen, R.B. (2010) The fauna of hydrothermal vents on the Mohn Ridge (North Atlantic). Marine Biology Research, 6, 155–171.
    Scott, T. (1892) Additions to the fauna of the Firth of Forth. Part IV. Reports of the Fishery Board for Scotland, Edinburgh, 10 (3), 244–272, pls. VII–XIII.
    Scott, T. (1894a) Report on Entomostraca from the Gulf of Guinea, collected by John Rattray, B.Sc. Transactions of the Linnean Society of London, Zoology, Series 2, 6, 1–161, pls. I–XV.
    Scott, T. (1894b) Additions to the fauna of the Firth of Forth. Part VI. Reports of the Fishery Board for Scotland, Edinburgh, 12 (3), 231–271, pls. V–X.
    Scott, T. (1897) The marine fishes and invertebrates of Loch Fyne. Reports of the Fishery Board for Scotland, Edinburgh, 15 (3), 107–174, pls. I–III.
    Scott, T. (1899) Report on the marine and freshwater Crustacea from Franz Josef Land, collected by Mr. William S. Bruce, of the Jackson Harmsworth Expedition. Journal of the Linnean Society, Zoology, 27, 60–126, pls. 3–9.
    Scott, T. (1900) Notes on some Crustacea from Fairlie and Hunterston, Firth of Clyde. Transactions of the natural History Society of Glasgow, New Series, 5 (3), 346–355.
    Scott, T. (1906) A catalogue of the land, fresh water and marine Crustacea found in the basin of the River Forth and its estuary. Proceedings of the Royal physical Society of Edinburgh, 16, 97–381. [Copepoda on 296–375]
    Scott, T. (1912) The Entomostraca of the Scottish National Antarctic Expedition, 1902–1904. Transactions of the Royal Society of Edinburgh, 48, 521–599, pls. I–XIV.
    Scott, T. & Scott, A. (1894) On some new and rare Crustacea from Scotland. Annals and Magazine of natural History, Series 6, 13, 137–149, pls. VIII–IX.
    Scott, T. & Scott, A. (1901) On some Entomostraca collected in the Arctic Seas in 1898 by William S. Bruce, F.R.S.G.S. Annals and Magazine of natural History, Series 7, 8, 337–356, pls. III–VI.
    Seifried, S. (2003) Phylogeny of Harpacticoida (Copepoda): Revision of “Maxillipedasphalea” and Exanechentera. Cuvillier Verlag, Göttingen, 259 pp.
    Şerban, M. (1959) Les Copépodes de la mer Noire. Note préliminaire sur les Harpacticides de la côte Roumaine. Lucrările ale Statiei Zoologice Maritime “Regele Ferdinand I” dela Agigea, Vol. Festival 1956, 259–302.
    Sergeeva, N.G. (2003) Meiobenthos of deep-water anoxic hydrogen sulphide zone of the Black Sea. In: Yilmaz, A. (Ed.), Oceanography of the Eastern Mediterranean and Black Sea. Similarities and differences of two interconnected basins. Tübitak Publications, Ankara, pp. 880–887.
    Sevastou, K. (2005) On the Ecology of Meiofauna of the Sandy Shores of Crete. Distribution Patterns of Meiobenthic Copepods over Space and Time. Ph. D. dissertation, University of Crete, Heraklio: vii + 233 pp. [in Greek with English summary]
    Sewell, R.B.S. (1940) Copepoda, Harpacticoida. The John Murray Expedition 1933–1934 Scientific Reports, 7 (2), 117–382.
    Shaffer, J.A., Munsch, S.H. & Cordell, J.R. (2020) Kelp forest zooplankton, forage fishes, and juvenile salmonids of the Northeast Pacific nearshore. Marine and Coastal Fisheries: Dynamics, Management, and Ecosystem Science, 12, 4–20.
    Shiells, G.M. (1983) A Study of the Influence of Mixed Industrial and Domestic Sewage Effluent on Marine Intertidal Meiobenthos on certain Contaminated Shores of Firth of Forth. Ph. D. dissertation, Napier College, Edinburgh. [unknown pagination]
    Sibert, J.R. (1981) Intertidal hyperbenthic populations in the Nanaimo estuary. Marine Biology, 64, 259–265.
    Sibert, J., Brown, T.J. & Kask, B.A. (1982) Meiofauna counts from the Fraser River, 1980 and 1981 Fraser foreshore core samples. Canadian Data Report of Fisheries and aquatic Sciences, 342, i–iv + 1–129.
    Simenstad, C.A., Cordell, J.R. & Weitkamp, L.A. (1991) Effects of substrate modification on littoral flat meiofauna: assemblage structure changes associated with adding gravel. FRI-UW-9124. Fisheries Research Institute, University of Washington, Seattle, Washinton, vi + 91 pp.
    Simenstad, C.A., Cordell, J.R., Wissmar, R.C., Fresh, K.L., Schroder, S., Carr, M. & Berg, M. (1988) Assemblage structure, microhabitat distribution, and food web linkages of epibenthic crustaceans in Padilla Bay National Estuarine Research Reserve, Washington. FRI-UW-8813. Fisheries Research Institute, University of Washington, Seattle, Washington, viii + 60 pp.
    Sloane, J.F., Bassindale, R., Davenport, E., Ebling, F.J. & Kitching, J.A. (1961) The ecology of Lough Ine. IX. The flora and fauna associated with undergrowth forming algae in the Rapids area. Journal of Ecology, 49, 353–368.
    Song, S.J., Kim, W. & Chang, C.Y. (2003) A new species of Parastenhelia (Copepoda: Harpacticoida: Parastenheliidae) from Korea. Zoological Science, 20, 221–228.
    Song, S.J., Park, J., Kwon, B.-O., Ryu, J. & Khim, J.S. (2012) Ecological checklist of the marine and brackish-water harpacticoid copepod fauna in Korean waters. Zoological Studies, 51, 1397–1410.
    Southern, R. (1915) Part 67. Marine ecology. In: A biological survey of Clare Island in the County of Mayo, Ireland and of the adjoining district. Proceedings of the Royal Irish Academy, (B)31(67), 1–110, pls. I–III.
    Soyer, J. (1971) Bionomie benthique du plateau continental de la côte catalane française. III. Les peuplements de Copépodes Harpacticoïdes (Crustacea). Vie et Milieu, (B), 21, 337–511, annexe.
    Spies, R.B., Hardin, D.D. & Toal, J.P. (1988) Organic enrichment or toxicity? A comparison of the effects of kelp and crude oil in sediments on the colonization and growth of benthic infauna. Journal of experimental marine Biology and Ecology, 124, 261–282.
    Steinarsdóttir, M.B. & Ingólfsson, A. (2004) Itunella muelleri (Harpacticoida: Canthocamptidae): a marine species with preferences for the upper part of the littoral fringe. Journal of crustacean Biology, 24, 440–446.
    Steinarsdóttir, M.B., Ingólfsson, A. & Ólafsson, E. (2003) Seasonality of harpacticoids (Crustacea, Copepoda) in a tidal pool in subarctic south-western Iceland. In: Jones, M.B., Ingólfsson, A., Ólafsson, E., Helgason, G.V., Gunnarsson, K. & Svavarsson, J. (Eds.), Migrations and Dispersal of Marine Organisms. Hydrobiologia, 503, pp. 211–221.
    Stephensen, K. (1929) Marine parasitic, semiparasitic and non pelagic Crustacea Copepoda. Zoology of the Faroes, 2 (1) (30), 1–18.
    Steuer, A. (1912) Phaoplanktonische Copepoden aus der südlichen Adria. Verhandlungen der zoologisch-botanischen Gesellschaft in Wien, 62, 64–69.
    Stringer, T.J., Glover, C.N., Keesing, V., Northcott, G.L. & Tremblay, L.A. (2012a) Development of a harpacticoid copepod bioassay: selection of species and relative sensitivity to zinc, atrazine and phenanthrene. Ecotoxicology and environmental Safety, 80, 363–371.
    Stringer, T.J., Korsman, J.C., Peralta, G., Keesing, V., Tremblay, L.A. & Glover, C.N. (2012b) Effects of environmental gradients on the distribution of harpacticoid copepods in an intertidal flat, Portobello Bay, Otago Harbour, New Zealand. New Zealand Journal of marine and freshwater Research, 46, 385–397.
    Suderman, K. & Thistle, D. (1998) Adult female harpacticoid copepods maintain their energy reserves by feeding while suspended during storms. Marine Ecology Progress Series, 164, 245–252.
    Thistle, D. (2003) Harpacticoid copepod emergence at a shelf site in summer and winter: implications for hydrodynamic and mating hypotheses. Marine Ecology Progress Series, 248, 177–185.
    Thistle, D., Weatherly, G.L. & Ertman, S.C. (1995a) Shelf harpacticoid copepods do not escape into the seabed during winter storms. Journal of marine Research, 53, 847–863.
    Thistle, D., Weatherly, G.L., Wonnacott, A. & Ertman, S.C. (1995b) Suspension by winter storms has an energetic cost for adult male benthic harpacticoid copepods at a shelf site. Marine Ecology Progress Series, 125, 77–86.
    Thompson, I.C. & Scott, A. (1903) Report on the Copepoda collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W.A. (Ed.), Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar, 1 (Supplement 7), pp. 227–307, pls. I–XX.
    Thomson, G.M. (1883) On the New Zealand Copepoda. Transactions and Proceedings of the New Zealand Institute, New Series, 15, 93–116, pls. V–XI.
    Thomson, G.M. (1913) The natural history of Otago Harbour and the adjacent sea, together with a record of the researches carried on at the Portobello marine fish-hatchery: Part I. Transactions and Proceedings of the New Zealand Institute, new series, 45, 225–251, pl. X.
    Toft, J. & Cordell, J.R. (2006) Olympic Sculpture Park: Results from pre-construction biological monitoring of shoreline habitats. Report prepared for Seattle Public Utilities, City of Seattle. School of Aquatic and Fishery Sciences, University of Washington, Seattle, Washington, v + 31 pp.
    Toft, J., Ogston, A., Heerhartz, S., Cordell, J.R., Armbrust, E. & Levy, C. (2010) Olympic Sculpture Park: year 3 monitoring of shoreline enhancements. Report (SAFS-UW-1002) prepared for Seattle Public Utilities, City of Seattle. School of Aquatic and Fishery Sciences, University of Washington, Seattle, Washington, 110 pp.
    Vanhöffen, E. (1897) Die Fauna und Flora von Grönland. In: Grönland Expedition der Gesellschaft für Erdkunde zu Berlin, 1891–1893, 2 (1), 1–383, pls. 1–8.
    Vasconcelos, D.M. (2008) Distribuição dos Copepoda Harpacticoida da Meiofauna em Área de Talude no litoral de Sergipe, Brasil. Ph. D. dissertation, Universidade Federal de Pernambuco, 99 pp.
    Vàtova, A. (1928) Compendio della flora e fauna del mare Adriatico presso Rovigno con la distribuzione geografica delle specie bentoniche. Memoria. R. Comitato talassografico italiano, 143, 1–614, maps 1–68.
    Ventham, D. (2011) Harpacticoid Copepods from the Sussex Coast (eastern Channel): Records 1992–1997. The Booth Museum of Natural History, Brighton, 133 pp.
    Vervoort, W. (1964) Freeliving Copepoda from Ifaluk Atoll in the Caroline Islands with notes on related species. Bulletin of the United States National Museum, 236, i–ix + 1–431.
    Villano, N. & Warwick, R.M. (1995) Meiobenthic communities associated with the seasonal cycle of growth and decay of Ulva rigida Agardh in the Palude della Rosa, Lagoon of Venice. Estuarine coastal, and Shelf Science, 41, 181–194.
    Walker-Smith, G.K. (2003) The Harpacticoid Fauna of Port Phillip Bay (Victoria, Australia) and their Contribution to the Diet of Juvenile King George Whiting (Sillaginodes punctata: Sillaginidae). Ph. D. dissertation, The Zoology Department, The University of Melbourne, 278 pp.
    Watkins, R.L. (1983) Harpacticoida (Crustacea: Copepoda) from the California coast. Ph. D. Dissertation, Arizona State University, Phoenix, Arizona, 312 pp.
    Webber, W.R., Fenwick, G.D., Bradford-Grieve, J.M., Eagar, S.H., Buckeridge, J.S., Poore, G.C.B., Dawson, E.W., Watling, L., Jones, J.B., Wells, J.B.J., Bruce, N.L., Ahyong, S.T., Larsen, K., Chapman, M.A., Olesen, J., Ho, J.-s., Green, J.D., Shiel, R.J., Rocha, C.E.F., Lörz, A.-N., Bird, G.J. & Charleston, W.A. (2010) Phylum Arthropoda. Subphylum Crustacea: shrimps, crabs, lobsters, barnacles, slaters, and kin. In: Dennis, P.G. (Ed.), New Zealand Inventory of Biodiversity. Vol. 2. Kingdom Animalia—Chaetognatha, Ecdysozoa, Ichnofossils. Canterbury University Press, Christchurch, pp. 98–232.
    Wells, J.B.J. (1961) Interstitial copepods from the Isles of Scilly. Crustaceana, 2, 262–274.
    Wells, J.B.J. (1963a) Copepoda from the littoral region of the estuary of the River Exe (Devon, England). Crustaceana, 5, 10–26.
    Wells, J.B.J. (1963b) On some new and rare Crustacea from Northern Ireland. Annals and Magazine of natural History, Series 13 (6), 85–96.
    Wells, J.B.J. (1967) The littoral Copepoda (Crustacea) of Inhaca Island, Mozambique. Transactions of the Royal Society of Edinburgh, 67 (7), 189–358.
    Wells, J.B.J. (1970) The marine flora and fauna of the Isles of Scilly. Crustacea: Copepoda: Harpacticoida. Journal of natural History, 4, 255–268.
    Wells, J.B.J. (1978) A preliminary report on the littoral copepods of Moce Island, Lau group, Fiji. In: Lau-Tonga 1977. Report from the Royal Society of New Zealand Expedition. Bulletin of the Royal Society of New Zealand, 17, pp. 91–93.
    Wells, J.B.J. (2007) An annotated checklist and keys to the species of Copepoda Harpacticoida (Crustacea). Zootaxa, 1568 (1), 1–872.
    Wells, J.B.J., Hicks, G.R.F. & Coull, B.C. (1982) Common harpacticoid copepods from New Zealand harbours and estuaries. New Zealand Journal of Zoology, 9, 151–184.
    Wells, J.B.J. & Rao, G.C. (1987) Littoral Harpacticoida (Crustacea: Copepoda) from Andaman and Nicobar Islands. Memoirs of the zoological Survey of India, 16 (4), 1–385.
    Wiborg, K.F. (1964) Marine copepods of Tristan da Cunha. Results of the Norwegian scientific Expedition to Tristan da Cunha 1937–1938, 51, 1–44.
    Wieser, W. (1952) Investigations on the microfauna inhabiting seaweeds on rocky coasts. (Untersuchungen über die algenbewohnende Mikrofauna mariner Hartböden). IV. Studies on the vertical distribution of the fauna inhabiting seaweeds below the Plymouth Laboratory. Journal of the marine biological Association of the United Kingdom, 31, 145–174.
    Willems, W.R., Curini-Galletti, M., Ferrero, T.J., Fontaneto, D., Heiner, I., Huys, R., Ivanenko, V.N., Kristensen, R.M. Kånneby, T., MacNaughton, M.O., Martínez Arbizu, P., Todaro, M.A., Sterrer, W. & Jondelius, U. (2008) Meiofauna of the Koster-area, results from a workshop at the Sven Lovén Centre for Marine Sciences (Tjärnö, Sweden). Meiofauna marina, 17, 1–34.
    Willen, E. (2000) Phylogeny of the Thalestridimorpha Lang, 1944 (Crustacea, Copepoda). Cuvillier Verlag, Göttingen, 233 pp.
    Willey, A. (1935) Harpacticoid Copepoda from Bermuda. Part II. Annals and Magazine of natural History, Series 10, 15, 50–100.
    Williams, G. (1954) Fauna of Strangford Lough and neighbouring coasts. Proceedings of the Royal Irish Academy. Section B: Biological, Geological, and Chemical Science, 56, 29–133.
    Wilson, C.B. (1932a) The copepod crustaceans of Chesapeake Bay. Proceedings of the United States national Museum, 80 (15), 1–54, pls. 1–5.
    Wilson, C.B. (1932b) The copepods of the Woods Hole region, Massachusetts. Bulletin of the United States national Museum, 158, 1–635, pls. 1–41.
    Yıldız, N.Ö. & Karaytuğ, S. (2018) Harpacticoida (Crustacea: Copepoda) of the three islands on Aegean Sea (Turkey) with eight new records. Mediterranean Fisheries and Aquaculture Research, 1, 57–65.
    Yu, O.H. & Suh, H.-L. (1983) Life history and secondary production of the amphilochid amphipod Gitanopsis japonica Hirayama, 1983, on a temperate sandy shore of Korea. Crustaceana, 85, 151–171.
    Yu, O.H., Suh, H.-L. & Shirayama, Y. (2003) Feeding ecology of three amphipod species Synchelidium lenorostralum, S. trioostegitum and Gitanopsis japonica in the surf zone of a sandy shore. Marine Ecology Progress Series, 258, 189–199.
    Zaleha, K., Hazwani, H.H., Syazleen, Z. & Nurul Huda, A.I. (2018) Report on five species of harpacticoid copepods from vegetative area of Sungai Pulai, Johor. Malaysian Journal of fundamental and applied Sciences, 14, 284–288.
    Zaleha, K., Nazia, A.K. & Nurul Huda, A.I. (2010) Species assemblages of benthic harpacticoid copepods on tide rock pool seaweeds of Pulau Besar, Melaka, Malaysia. Journal of tropical Biology and Conservation, 7, 1–10.
    Zaleha, K., Roswati, M.A. & Iwasaki, N. (2006) Distribution of some species of harpacticoid copepods in east coast of Peninsular Malaysia. Coastal marine Science, 30, 140–145.
    Zavodnik, D. (1969) La communauté à Acetabularia mediterranea Lamour dans l’Adriatique du Nord. Internationale Revue der gesamten Hydrobiologie, 54, 543–551.

How to Cite

HUYS, R. ., & MU, F. . (2021). <em>Johnwellsia</em>, a new intertidal genus of Parastenheliidae (Copepoda, Harpacticoida) from the Taiwan Strait, China, including a review of the family and key to genera. Zootaxa, 5051(1), 236–318.