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Article
Published: 2021-10-12

John Berkeley James Wells—An appreciation of his contributions to harpacticoid diversity and systematics

Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, U.K.
Arthropoda

Abstract

John Wells was born in Hammersmith, a district in west London, where he spent most of his childhood and teenage life. It was a surprise to find out only recently that he had received a scholarship to Latymer Upper School on King Street which is literally one block away from where I used to live when I started working at the Natural History Museum in the early 1990s. The site has a long history and can be traced to a charity school founded in 1624 by the English merchant Edward Latymer, a wealthy lawyer and puritan, who left part of his wealth for the clothing and education of “eight poore boyes” from Hammersmith.

 

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    Kunz, H. (1962) Revision der Paramesochridae (Crust. Copepoda). Kieler Meeresforschungen, 18, 245–257.
    Kunz, H. (1974) Zwei neue afrikanische Paramesochridae (Copepoda Harpacticoidea) mit Darstellung eines Bewegungsmechanismus für die Furkaläste. Mikrofauna des Meeresbodens, 36, 1–20.
    Kunz, H. (1975) Copepoda Harpacticoida aus dem Litoral des südlichen Afrika. I. Teil. Kieler Meeresforschungen, 31, 179–212.
    Kunz, H. (1981) Beitrag zur Systematik der Paramesochridae (Copepoda, Harpacticoida) mit Beschreibung einiger neuen Arten. Mitteilungen aus dem zoologischen Museum der Universität Kiel, 1 (8), 2–33.
    Lang, K. (1936) Die Familie der Cletodidae Sars, 1909. Zoologische Jahrbücher für Systematik, 68, 445–480.
    Lang, K. (1948) Monographie der Harpacticiden. Vol. 1 & 2. Håkan Ohlsson, Lund, 1682 pp.
    Lang, K. (1965) Copepoda Harpacticoida from the Californian Pacific coast. Kungliga Svenska Vetenskapsakademiens Handlingar, Serier 4, 10 (2), 1–560.
    Lewis, M.H. (1972) Freshwater harpacticoid copepods of New Zealand. 1. Attheyella and Elaphoidella (Canthocamptidae). New Zealand Journal of marine and Freshwater Research, 6, 23–47. https://doi.org/10.1080/00288330.1977.9515408
    Löffler, H. (1968) Zur Harpacticiden fauna des Östlichen Nepal mit besonderer Berücksichtigung der Gattung Maraenobiotus. Archiv für Hydrobiologie, 65, 1–24.
    Lorenzen, S. (1969) Harpacticoiden aus dem lenitischen Watt und den Salzwiesen der Nordseeküste. Kieler Meeresforschungen, 25, 215–223.
    Ma, L., Liu, Q.H., Li, X.Z. & Huys, R. (2021) A new species of Stylicletodes Lang, 1936 (Copepoda: Harpacticoida: Cletodidae) from the East China Sea, including an updated key to species and synopsis of distribution records. Zootaxa, 5051 (1), 423–442. https://doi.org/10.11646/zootaxa.5051.1.17
    McIntyre, A.D. (1961) Quantitative differences in the fauna of boreal mud associations. Journal of the marine biological Association of the United Kingdom, 41, 599–616. https://doi.org/10.1017/S0025315400016180
    McIntyre, A.D. (1962) The class Kinorhyncha (Echinoderida) in British waters. Journal of the marine biological Association of the United Kingdom, 42, 503–510. https://doi.org/10.1017/S0025315400054217
    McIntyre, A.D. (1964) Meiobenthos of sub-littoral muds. Journal of the marine biological Association of the United Kingdom, 44, 665–674. https://doi.org/10.1017/S0025315400027843
    McIntyre, A.D. (1968) The meiofauna and macrofauna of some tropical beaches. Journal of Zoology, 156, 377–392. https://doi.org/10.1111/j.1469-7998.1968.tb04360.x
    McIntyre, A.D. (1969) Ecology of marine meiobenthos. Biological Reviews, 44, 245–288. https://doi.org/10.1111/j.1469-185X.1969.tb00828.x
    Marcus, A. (1971) Copépodes du mésopsammon de la zone médiolittorale de la mer Noire (littoral roumain). La description d’une nouvelle sous espèce Klieonychocamptus kliei ponticus n. ssp. Travaux du Muséum d’Histoire naturelle “Gr. Antipa”, 11, 71–82.
    Marcus, A. (1974) Contributions to the study of the genus Amphiascus Sars (part.), Copepoda, Harpacticoida from the Black Sea. Travaux du Muséum d’Histoire naturelle “Gr. Antipa”, 15, 111–122.
    Mare, M.F. (1942) A study of a marine benthic community with special reference to the micro-organisms. Journal of the marine biological Association of the United Kingdom, 25, 517–554. https://doi.org/10.1017/S0025315400055132
    Marinov, T.M. (1973) Quelques espèces de la famille Ameiridae (Copépodes, Harpacticoïdes) dans la Mer Noire. Crustaceana, 24, 231–241. https://doi.org/10.1163/156854073X00407
    Melville, R.V. (1985) Opinion 1356. Dactylopusia Norman, 1903 (Crustacea, Copepoda): type species designated. Bulletin of zoological Nomenclature, 42, 335–337.
    Mielke, W. (1973) Zwei neue Harpacticoidea (Crustacea) aus dem Eulitoral der Nordseeinsel Sylt. Mikrofauna des Meeresbodens, 17, 1–14.
    Mielke, W. (1975) Systematik der Copepoda eines Sandstrandes der Nordseeinsel Sylt. Mikrofauna des Meeresbodens, 52, 1–134.
    Mielke, W. (1984) Interstitielle Fauna von Galapagos. XXXI. Paramesochridae (Harpacticoida). Microfauna marina, 1, 63–147.
    Monard, A. (1926a) Sur les Harpacticus de Banyuls. Bulletin de la Société zoologique de France, 51, 419–434.
    Monard, A. (1926b) Description de quelques espèces nouvelles d’Harpacticides marins de la région de Banyuls. Revue suisse de Zoologie, 33, 619–628.
    Monard, A. (1928) Les Harpacticoïdes marins de Banyuls. Archives de Zoologie expérimentale et générale, 67, 259–443.
    Monard, A. (1935a) Étude sur la faune des Harpacticoïdes marins de Roscoff. Travaux de la Station biologique de Roscoff, 13, 5–88.
    Monard, A. (1935b) Les Harpacticoïdes marins de la région de Salammbô. Bulletin de la Station océanographique de Salammbô, 34, 1–94.
    Moore, C.G. (1976) The harpacticoid families Thalestridae and Ameiridae (Crustacea, Copepoda) from the Isle of Man. Journal of natural History, 10, 29–56. https://doi.org/10.1080/00222937600770041
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    Newell, I.M. (1970) Construction and use of tabular keys. Pacific Insects, 12, 25–37.
    Newell, I.M. (1972) Tabular keys: further notes on their construction and use. Transactions of the Connecticut Academy of Arts and Sciences, 44, 259–267.
    Nicholls, A.G. (1935) Copepods from the interstitial fauna of a sandy beach. Journal of the marine biological Association of the United Kingdom, New Series, 20, 379–405. https://doi.org/10.1017/S0025315400045306
    Nicholls, A.G. (1939) Some new sand-dwelling copepods. Journal of the marine biological Association of the United Kingdom, new series, 23, 327–341. https://doi.org/10.1017/S0025315400013928
    Nicholls, A.G. (1941) A revision of the families Diosaccidae Sars, 1906 and Laophontidae T. Scott, 1905 (Copepoda, Harpacticoida). Records of the South Australian Museum, 7, 65–110.
    Nicholls, A.G. (1944) Littoral Copepoda from the Red Sea. Annals and Magazine of natural History, Series 11, 11, 487–503. https://doi.org/10.1080/00222934408527451
    Nogueira, M.H. (1959) O gênero Elaphoidella (Harpacticoidea—Cop.—Crust.) nas águas do Paraná. Dusenia, 8, 61–68.
    Noodt, W. (1952) Marine Harpacticiden (Cop.) aus dem eulitoralen Sandstrand der Insel Sylt. Abhandlungen der mathematisch-naturwissenschaftlichen Klasse. Akademie der Wissenschaften und der Literatur in Mainz, 1952 (3), 105–142.
    Noodt, W. (1955) Harpacticiden (Crust. Cop.) aus dem Sandstrand der französischen Biscaya-Küste. Kieler Meeresforschungen, 11, 86–109, pls. 27–38.
    Noodt, W. (1958) Die Copepoda Harpacticoidea des Brandungsstrandes von Teneriffa (Kanarische Inseln). Abhandlungen der mathematisch-naturwissenschaftlichen Klasse. Akademie der Wissenschaften und der Literatur in Mainz, 1958 (2), 53–116.
    Norman, A.M. & Scott, T. (1905) Crustacea Copepoda new to Science from Devon and Cornwall. Annals and Magazine of natural History, Series 7, 15, 284–300. https://doi.org/10.1080/03745480509443044
    Norman, A.M. & Scott, T. (1906) The Crustacea of Devon and Cornwall. William Wesley & Son, London, xv + 232 pp.
    Özdikmen, H. (2008) Nomenclatural changes for nine crustacean genera (Crustacea: Copepoda). Munis Entomology & Zoology, 3, 265–274.
    Pallares, R.E. (1973) El género Harpacticus en la Ría Deseado (Crustacea, Copepoda). Physis, Buenos Aires, (A), 32, 275–288.
    Pallares, R.E. (1982) Copépodos harpacticoides marinos de Tierra del Fuego (Argentina). IV. Bahía Tethis. Contribuciónes científica del Centro de Investigación de Biología marina, Buenos Aires, 186, 3–39.
    Pennak, R.W. (1942) Harpacticoid copepods from some intertidal beaches near Woods Hole, Massachusetts. Transactions of the American microscopical Society, 61, 274–285. https://doi.org/10.2307/3222599
    Perkins, E.J. (1956a) Microarthridion fallax, a species of harpacticid copepod new to science from Whitstable, Kent. Annals and Magazine of natural History, Series 12, 9, 108–111. https://doi.org/10.1080/00222935608655733
    Perkins, E.J. (1956b) The harpacticoid genus Tetanopsis Brady, with a description of Tetanopsis smithi, sp. nov. and Tetanopsis medius, sp. nov. Annals and Magazine of natural History, Series 12, 9, 497–504. https://doi.org/10.1080/00222935608655845
    Pesta, O. (1935) Marine Harpacticiden aus dem Hawaiischen Inselgebiet. Zoologische Jahrbücher, Abteilung für Systematik, 66, 363–379.
    Petkovski, T.K. (1955) Zweiter Beitrag zur Kenntnis der Harpacticidenfauna unserer Meeresküste. Fragmenta balcanica, 1, 125–139.
    Petkovski, T.K. (1986) Zur Taxonomie des Genus Neocyclops Gurney 1927 (Crustacea, Copepoda Cyclopoida). Acta Musei macedonici Scientiarum naturalium, 18 (2/148), 27–46.
    Petkovski, T.K. & Brancelj, A. (1985) Zur Copepodenfauna (Crustacea) der Grotten Škocjanske Jame in Slowenien. Acta Musei macedonici Scientiarum naturalium, 18 (1), 1–25.
    Philippi, A. (1840) Zoologische Bemerkungen (Fortsetzung). IV. Kurze Charakteristik mehrerer neuer Genera aus der Familie der Copepoden. Archiv für Naturgeschichte, 6, 188–190.
    Pleşa, C. (1961) New cyclopoids (Crustacea, Copepoda) of the interstitial fauna from the beaches of Ghana. Journal of the West African Science Association, 7 (1), 1–13.
    Por, F.D. (1964) A study of Levantine and Pontic Harpacticoida (Crustacea, Copepoda). Zoölogische Verhandelingen, Leiden, 64, 1–128.
    Por, F.D. (1967) Level bottom Harpacticoida (Crustacea, Copepoda) from Elat (Red Sea), part I. Israel Journal of Zoology, 16, 101–165. https://doi.org/10.1080/00212210.1967.10688252
    Raffaelli, D.G. & Mason, C.F. (1981) Pollution monitoring with meiofauna, using the ratio of nematodes to copepods. Marine Pollution Bulletin, 12, 158–163. https://doi.org/10.1016/0025-326X(81)90227-7
    Rao, G.C. (1967) On the life history of a new sand dwelling harpacticoid copepod. Crustaceana, 13, 129–136. https://doi.org/10.1163/156854067X00297
    Rao, G.C. (1972) Some new interstitial harpacticoid copepods from Andhra coast, India. Cahiers de Biologie marine, 13, 305–319. https://doi.org/10.21411/CBM.A.6C3F88E8
    Rao, G.C. (1975) The interstitial fauna in the intertidal sands of Andaman and Nicobar groups of islands. Journal of the marine biological Association of India, 17, 116–128.
    Rao, G.C. (1980) On the zoogeography of the interstitial meiofauna of the Andaman and Nicobar Islands, Indian Ocean. Records of the Zoological Survey of India, 77, 153–178.
    Rao, G.C. & Ganapati, P.N. (1969a) Some new interstitial copepods from Waltair coast. Proceedings of the Indian Academy of Sciences, (B), 69, 1–14.
    Rao, G.C. & Ganapati, P.N. (1969b) On some interstitial copepods from the beach sands of Waltair coast. Proceedings of the Indian Academy of Sciences, (B), 70, 262–286.
    Remane, A. (1927) Halammohydra, ein eigenartiges Hydrozoon der Nord- und Ostsee. Zeitschrift für Morphologie und Ökologie der Tiere, 7 (4), 643–677. https://doi.org/10.1007/BF00540535
    Roe, K.M. (1955) Genus Paraphyllopodopsyllus Lang (Copepoda Harp.). Two new species—P. hibernicus and P. hardingi. Proceedings of the Royal Irish Academy, (B), 57, 131–139.
    Roe, K.M. (1958) The littoral harpacticids of the Dalkey (Co. Dublin) area with descriptions of six new species. Proceedings of the Royal Irish Academy, (B), 59, 221–255.
    Roe, K.M. (1960) Some harpacticids from Lough Ine, with descriptions of two new species. Proceedings of the Royal Irish Academy, (B), 60, 277–289.
    Sak, S., Huys, R. & Karaytuğ, S. (2008a) Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae). Zoological Journal of the Linnean Society, 152, 409–458. https://doi.org/10.1111/j.1096-3642.2007.00373.x
    Sak, S., Karaytuğ, S. & Huys, R. (2008b) A review of Pseudoleptomesochrella Lang, 1965 (Copepoda, Harpacticoida, Ameiridae), including a redescription of P. halophila (Noodt, 1952) from the Black Sea and a key to species. Zootaxa, 1758 (1), 45–60. https://doi.org/10.11646/zootaxa.1758.1.3
    Sars, G.O. (1903) Copepoda Harpacticoida. Parts I & II, Misophriidæ, Longipediidæ, Cerviniidæ, Ectinosomidæ (part). An Account of the Crustacea of Norway, with short Descriptions and Figures of all the Species, 5, 1–28, pls. I–XVI.
    Sars, G.O. (1905) Pacifische Plankton-Crustaceen. (Ergebnisse einer Reise nach dem Pacific. Schauinsland 1896–1897). II. Brackwasser-Crustaceen von den Chatham-Inseln. Zoologische Jahrbücher für Systematik, 21 (4), 371–414.
    Sars, G.O. (1906) Copepoda Harpacticoida. Parts XIII & XIV. Diosaccidæ (continued). An Account of the Crustacea of Norway, with short Descriptions and Figures of all the Species, 5, 157–172, pls. XCVII–CXII.
    Sars, G.O. (1909) Copepoda Harpacticoida. Parts XXVII & XXVIII. Cletodidæ (concluded), Anchorabolidæ, Cylindropsyllidæ, Tachidiidæ (part). An Account of the Crustacea of Norway, with short Descriptions and Figures of all the Species, 5, 305–336, pls. 209–224.
    Sars, G.O. (1911) Copepoda Harpacticoida. Parts XXXV & XXXVI. Supplement (concluded), index, etc. An Account of the Crustacea of Norway, with short Descriptions and Figures of all the Species, 5, 421–449 + i–xiv (text) + i–xii (plates), title of text and of plates, supplement pls. 43–54.
    Sars, G.O. (1920) Copepoda Supplement. Parts III & IV. Harpacticoida (continued). An Account of the Crustacea of Norway, with short Descriptions and Figures of all the Species, 7, 25–52, pls. XVI–XXXII.
    Sars, G.O. (1921) Copepoda Supplement. Parts IX & X. Harpacticoida (concluded), Cyclopoida. An Account of the Crustacea of Norway, with short Descriptions and Figures of all the Species, 7, title page, preface, 93–121, pls. LXV–LXXVI.
    Schäfer, H.W. (1936) Cyclopiniden (Crustacea Copepoda) aus der deutschen Nordsee. Zoologischer Anzeiger, 114, 225–234.
    Schizas, N.V., Dahms, H.-U., Kangtia, P., Corgosinho, P.H.C. & Galindo Estronza, A.M. (2015) A new species of Longipedia Claus, 1863 (Copepoda: Harpacticoida: Longipediidae) from Caribbean mesophotic reefs with remarks on the phylogenetic affinities of Polyarthra. Marine Biology Research, 11, 789–803. https://doi.org/10.1080/17451000.2015.1013556
    Schmeil, O. (1894) Einige neue Harpacticiden Formen des Süsswassers. Zeitschrift für Naturwissenschaften, 67, 341–350.
    Schminke, H.K. (2008) First report of groundwater fauna from Papua New Guinea: Kinnecaris Jakobi, 1972 redefined (Copepoda, Harpacticoida, Parastenocarididae), and description of a new species. Crustaceana, 81, 1241–1253. https://doi.org/10.1163/156854008X374568
    Scott, A. (1902) On some Red Sea and Indian Ocean Copepoda. Proceedings and Transactions of the Liverpool biological Society, 16, 397–428, pls. I–III.
    Scott, A. (1909) The Copepoda of the Siboga Expedition. Part I. Free-swimming, littoral and semi-parasitic Copepoda. Siboga-Expeditie. Uitkomsten op zoologisch, botanisch, oceanographisch en geologisch gebied verzameld in Nederlandsch Oost-Indië 1899–1900 aan boord H.M. Siboga onder commando van Luitenant ter Zee 1e kl. F. Tydeman, 29a, 1–323.
    Scott, T. (1894a) Additions to the fauna of the Firth of Forth. Part VI. Reports of the Fishery Board for Scotland, Edinburgh, 12 (3), 231–271, pls. V–X.
    Scott, T. (1894b) Report on Entomostraca from the Gulf of Guinea, collected by John Rattray, B.Sc. Transactions of the Linnean Society of London, Zoology, Series 2, 6, 1–161, pls. I–XV. https://doi.org/10.1111/j.1096-3642.1894.tb00660.x
    Scott, T. (1895) Additions to the fauna of the Firth of Forth. Part VII. Reports of the Fishery Board for Scotland, Edinburgh, 13 (3), 165–173, pls. III–IV.
    Scott, T. (1902) Notes on gatherings of Crustacea collected by the Fishery Steamer “Garland” and the steam trawlers “Star of Peace” and “Star of Hope”, of Aberdeen, during the year 1901. Reports of the Fishery Board for Scotland, Edinburgh, 20 (3), 447–485, pls. XXII–XXV.
    Scott, T. (1905) On some new and rare Crustacea from the Scottish seas. Reports of the Fishery Board for Scotland, Edinburgh, 23 (3), 141–153, pls. X–XIII.
    Scott, T. & Scott, A. (1893) Notes on Copepoda from the Firth of Forth: Longipedia coronata, Claus; and a preliminary description of an apparently new genus and species. Annals of Scottish natural History, 1893, 89–94, pl. II.
    Scott, T. & Scott, A. (1896) Revision of the British species belonging to the genera Bradya, Boeck, and Ectinosoma, Boeck. Transactions of the Linnean Society of London, Zoology, Series 2, 6, 419–446, pls. 35–38. https://doi.org/10.1111/j.1096-3642.1896.tb00483.x
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    Vervoort, W. & Holthuis, L.B. (1983) Dactylopusia Norman, 1903 (Crustacea, Copepoda): proposed designation of type species. Z.N.(S.)1517. Bulletin of zoological Nomenclature, 40, 56–57.
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  6. APPENDIX 1. Publications of J.B.J. Wells (and co-workers) on harpacticoid and other copepods arranged in chronological order.

  7. Wells, J.B.J. (1961a) The Ecological Distribution of the Microfauna of the Exe estuary with especial Reference to the Copepoda. Ph.D. dissertation, University of Exeter.
    Wells, J.B.J. (1961b) Interstitial copepods from the Isles of Scilly. Crustaceana, 2, 262–274. https://doi.org/10.1163/156854061X00392
    Wells, J.B.J. (1963a) Copepoda from the littoral region of the estuary of the River Exe (Devon, England). Crustaceana, 5, 10–26. https://doi.org/10.1163/156854063X00020
    Wells, J.B.J. (1963b) On some new and rare Crustacea from Northern Ireland. Annals and Magazine of natural History, Series 13, 6, 85–96. https://doi.org/10.1080/00222936308651328
    Wells, J.B.J. (1964) Some additions to the Dale Fort marine fauna. Proceedings of the zoological Society of London, 142, 453–458. https://doi.org/10.1111/j.1469-7998.1964.tb04509.x
    Wells, J.B.J. (1965a) Two new genera of harpacticoid copepods of the family Ectinosomidae. Revista de Biologia, Lisboa, 5, 30–35.
    Wells, J.B.J. (1965b) Copepoda (Crustacea) from the meiobenthos of some Scottish marine sub-littoral muds. Proceedings of the Royal Society of Edinburgh, (B), 69 (1), 1–33. https://doi.org/10.1017/S0080455X00010110
    Wells, J.B.J. (1965c) Six new species of Parastenocaris (Crustacea, Copepoda) from southern Rhodesia. Annals and Magazine of natural History, Series 13, 7, 193–204. https://doi.org/10.1080/00222936408651458
    Wells, J.B.J. & Clark, M.E. (1965) The interstitial Crustacea of two beaches in Portugal. Revista de Biologia, Lisboa, 5(1/2), 87–108.
    Wells, J.B.J. (1967) The littoral Copepoda (Crustacea) of Inhaca Island, Mozambique. Transactions of the Royal Society of Edinburgh, 67, 189–358. https://doi.org/10.1017/S0080456800024017
    Wells, J.B.J. (1968) New and rare Copepoda Harpacticoida from the Isles of Scilly. Journal of natural History, 2, 397–424. https://doi.org/10.1080/00222936800770391
    Wells, J.B.J. (1970a) The marine flora and fauna of the Isles of Scilly. Crustacea: Copepoda: Harpacticoida. Journal of natural History, 4, 255–268. https://doi.org/10.1080/00222937000770251
    Wells, J.B.J. (1970b) Copepoda.—I. Sub-order Harpacticoida. Fiches d’Identification du Plancton, 133, 1–7.
    Lasker, R., Wells, J.B.J. & McIntyre, A.D. (1970) Growth, reproduction, respiration and carbon utilization of the sand-dwelling harpacticoid copepod Asellopsis intermedia. Journal of the marine biological Association of the United Kingdom, 50, 147–160. https://doi.org/10.1017/S0025315400000679
    Wells, J.B.J. (1971a) The Harpacticoida (Crustacea: Copepoda) of two beaches in south-east India. Journal of natural History, 5, 507–520. https://doi.org/10.1080/00222937100770381
    Wells, J.B.J. (1971b) A brief review of methods of sampling the meiobenthos. In: Hulings, N.C. (Ed.), Proceedings of First International Conference on Meiofauna. Smithsonian Contributions to Zoology, 76, 183–186.
    Wells, J.B.J. & McKenzie, K.G. (1973) Report on a small collection of benthic copepods from marine and brackish waters of Aldabra, Indian Ocean. Crustaceana, 25, 133–146. https://doi.org/10.1163/156854073X00786
    Wells, J.B.J., Kunz, H. & Rao, G.C. (1975) A review of the mechanisms for movement of the caudal furca in the family Paramesochridae (Copepoda Harpacticoida), with a description of a new species of Kliopsyllus Kunz. Mikrofauna des Meeresbodens, 53, 1–16.
    Wells, J.B.J. (1976) Keys to Aid in the Identification of Marine Harpacticoid Copepods. Department of Zoology, University of Aberdeen, Aberdeen, 215 pp.
    Wells, J.B.J. & Rao, G.C. (1976) The relationship of the genus Schizopera Sars within the family Diosaccidae (Copepoda: Harpacticoida). Zoological Journal of the Linnean Society, 58, 79–90. https://doi.org/10.1111/j.1096-3642.1976.tb00821.x
    Wells, J.B.J. (1977) Protopsammotopa wilsoni n. sp. (Copepoda, Harpacticoida) and the fate of the type material of Goffinella stylifer C.B. Wilson. Crustaceana, 32, 200–202. https://doi.org/10.1163/156854077X00584
    Munro, A.L.S., Wells, J.B.J. & McIntyre, A.D. (1978) Energy flow in the flora and meiofauna of sandy beaches. Proceedings of the Royal Society of Edinburgh, 76B, 297–315. https://doi.org/10.1017/S0269727000002943
    Wells, J.B.J. (1978a) Keys to aid in the identification of marine harpacticoid copepods. Amendment Bulletin no. 1. Zoological Publications Victoria University of Wellington, 70, 1–11.
    Wells, J.B.J. (1978b) A preliminary report on the littoral copepods of Moce Island, Lau group, Fiji. In: Lau-Tonga 1977. Report from the Royal Society of New Zealand Expedition. Bulletin of the Royal Society of New Zealand, 17, pp. 91–93.
    Wells, J.B.J. (1979) Keys to aid in the identification of marine harpacticoid copepods. Amendment Bulletin no. 2. Zoological Publications Victoria University of Wellington, 73, 1–8.
    Wells, J.B.J. (1980) A revision of the genus Longipedia Claus (Crustacea: Copepoda: Harpacticoida). Zoological Journal of the Linnean Society, 70, 103–189. https://doi.org/10.1111/j.1096-3642.1980.tb00850.x
    Coull, B.C., Hicks, G.R.F. & Wells, J.B.J. (1981) Nematode/copepod ratios for monitoring pollution: a rebuttal. Marine Pollution Bulletin, 12, 378–381. https://doi.org/10.1016/0025-326X(81)90408-2
    Coull, B.C. & Wells, J.B.J. (1981) Density of mud-dwelling meiobenthos from three sites in the Wellington region. New Zealand Journal of Marine and Freshwater Research, 15, 411–415. https://doi.org/10.1080/00288330.1981.9515933
    Wells, J.B.J. (1981) Keys to aid in the identification of marine harpacticoid copepods. Amendment bulletin no. 3. Zoological Publications Victoria University of Wellington, 75, 1–13.
    Wells, J.B.J., Hicks, G.R.F. & Coull, B.C. (1982a) Common harpacticoid copepods from New Zealand harbours and estuaries. New Zealand Journal of Zoology, 9, 151–184. https://doi.org/10.1080/03014223.1982.10423847
    Wells, J.B.J., Hicks, G.R.F. & Coull, B.C. (1982b) Erratum. New Zealand Journal of Zoology, 9, 522. https://doi.org/10.1080/03014223.1982.10423885
    Bradford, J.M. & Wells, J.B.J. (1983) New calanoid and harpacticoid Copepoda from beneath the Ross Ice Shelf, Antarctica. Polar Biology, 2, 1–15. https://doi.org/10.1007/BF00258279
    Coull, B.C., Creed, E.L., Eskin, R.A., Montagna, P.A., Palmer, M.A. & Wells, J.B.J. (1983) Phytal meiofauna from the rocky intertidal at Murrells Inlet, South Carolina. Transactions of the American microscopical Society, 102, 380–389. https://doi.org/10.2307/3225851
    Coull, B.C. & Wells, J.B.J. (1983) Refuges from fish predation: experiments with phytal meiofauna from the New Zealand rocky intertidal. Ecology, 64, 1599–1609. https://doi.org/10.2307/1937513
    Wells, J.B.J. (1983) Keys to aid in the identification of marine harpacticoid copepods. Amendment Bulletin no. 4. Zoological Publications Victoria University of Wellington, 77, 1–9.
    Wells, J.B.J. (1985) Keys to aid in the identification of marine harpacticoid copepods. Amendment Bulletin no. 5. Zoological Publications Victoria University of Wellington, 80, 1–19.
    Wells, J.B.J. (1986a) Biogeography of benthic harpacticoid copepods of the marine littoral and continental shelf. In: Schriever, G., Schminke, H.K. & Shih, C.-T. (Eds.), Proceedings II. International Conference on Copepoda, Ottawa, Canada, 13–17 August 1984, Syllogeus, 58, 126–135.
    Wells, J.B.J. (1986b) Copepoda: Marine-interstitial Harpacticoida. In: Botosaneanu, L. (Ed.), Stygofauna Mundi, a Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial). E.J. Brill/Dr. W. Backhuys, Leiden, pp. 356–381.
    Wells, J.B.J. & Rao, G.C. (1987) Littoral Harpacticoida (Crustacea: Copepoda) from Andaman and Nicobar Islands. Memoirs of the zoological Survey of India, 16 (4), 1–385.
    Wells, J.B.J. (1988) Copepoda. In: Higgins, R.P. & Thiel, H. (Eds.), Introduction to the Study of Meiofauna. Smithsonian Institution Press, Washington, D.C. & London, pp. 380–388.
    Wells, J.B.J. (2007) An annotated checklist and keys to the species of Copepoda Harpacticoida (Crustacea). Zootaxa, 1568 (1), 1–872. https://doi.org/10.11646/zootaxa.1568.1.1
    Webber, W.R., Fenwick, G.D., Bradford-Grieve, J.M., Eagar, S.H., Buckeridge, J.S., Poore, G.C.B., Dawson, E.W., Watling, L., Jones, J.B., Wells, J.B.J., Bruce, N.L., Ahyong, S.T., Larsen, K., Chapman, M.A., Olesen, J., Ho, J.-s., Green, J.D., Shiel, R.J., Rocha, C.E.F., Lörz, A.-N., Bird, G.J. & Charleston, W.A. (2010) Phylum Arthropoda. Subphylum Crustacea: shrimps, crabs, lobsters, barnacles, slaters, and kin. In: Gordon, D.P. (Ed.), New Zealand Inventory of Biodiversity. Vol. 2. Kingdom Animalia—Chaetognatha, Ecdysozoa, Ichnofossils. Canterbury University Press, Christchurch, pp. 98–232.
    Charry, M.P., Wells, J.B.J., Keesing, V., Smith, K.F., Stringer, T.J. & Tremblay, L.A. (2019) Quinquelaophonte aurantius sp. nov., a new harpacticoid species (Copepoda: Harpacticoida: Laophontidae: Quinquelaophonte) from New Zealand. New Zealand Journal of Zoology, 46, 301–320.
    https://doi.org/10.1080/03014223.2018.1548496

How to Cite

HUYS, R. . (2021). John Berkeley James Wells—An appreciation of his contributions to harpacticoid diversity and systematics. Zootaxa, 5051(1), 11–40. https://doi.org/10.11646/zootaxa.5051.1.5