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Published: 2022-06-22

New taxonomic data on Zodariinae (Araneae: Zodariidae) of Azerbaijan, Iran, Afghanistan and Pakistan

Zoological Museum, Biodiversity Unit, University of Turku, FI-20014, Finland.
Institute for Biological Problems of the North RAS, Portovaya Str.18, Magadan, Russia. 3Department of Zoology & Entomology, University of the Free State, Bloemfontein 9300, South Africa.
Araneae new combination new record new species Indozodion Parazodarion Trygetus Zodarion


New taxonomic considerations on several species of the zodariid spider subfamily Zodariinae Thorell, 1881 occurring in Azerbaijan, Iran, Afghanistan and Pakistan are presented. Three species, Trygetus cyrus sp. n. (♀; Fars, south-central Iran), T. nakhchivanicus sp. n. (♂; Nakhchivan, Azerbaijan) and Zodarion sharurensis sp. n. (♂♀; Nakhchivan, Azerbaijan) are described as new, and the hitherto unknown male of Trygetus susianus Zamani & Marusik, 2021 is described based on topotype material from Khuzestan, southwestern Iran. Indozodion Ovtchinnikov, 2006 stat. rev. is rediagnosed and removed from the synonymy of Tropizodium Jocqué & Churchill, 2005, and here considered to comprise I. inayatullahi Ovtchinnikov, 2006 comb. rev. (type species) and I. lindbergi (Roewer, 1960) comb. n. (ex. Zodarion Walckenaer, 1826); the latter species, previously known only from Afghanistan, is redescribed and reported from Pakistan. One of the paratypes of I. lindbergi, reported as a subadult female in the original description, is found to be an adult female of Parazodarion raddei (Simon, 1889). An error in a previous publication (attributing figures of Acanthinozodium elburzicum Zamani & Marusik, 2021 to P. raddei) is also corrected here.



  1. Dankittipakul, P., Jocqué, R. & Singtripop, T. (2012) Two new species of the spider genus Tropizodium Jocqué & Churchill, 2005 with notes on the femoral organ clade from Southeast Asia (Araneae, Zodariidae, Zodariinae). Zootaxa, 3327 (1), 62–68.
    Denis, J. (1958) Araignées (Araneidea) de l'Afghanistan. I. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjøbenhavn, 120, 81–120.
    Jocqué, R. (1991) A generic revision of the spider family Zodariidae (Araneae). Bulletin of the American Museum of Natural History, 201, 1–160.
    Levy, G. (1992) The spider genera Palaestina, Trygetus, Zodarion and Ranops (Araneae, Zodariidae) in Israel with annotations on species of the Middle East. Israel Journal of Zoology, 38, 67–110.
    Marusik, Y.M. (2011) A new Trygetus species from Central Asia (Araneae: Zodariidae). Turkish Journal of Zoology, 35, 29–31.
    Marusik, Y.M. & Guseinov, E. (2003) Spiders (Arachnida: Aranei) of Azerbaijan. 1. New family and genus records. Arthropoda Selecta, 12, 29–46.
    Ono, H. & Jocqué, R. (1986) Two new species of the family Zodariidae from Saudi Arabia (Arachnida: Araneae). Fauna Saudi Arabia, 8, 6–11.
    Otto, S. (2022) Caucasian Spiders. A faunistic database on the spiders of the Caucasus. Version 02.2022. Available from: (accessed 7 June 2022)
    Ovtchinnikov, S.V. (2006) New genus and species of spiders of the subfamily Zodariinae (Araneae, Zodariidae) from Pakistan. Vestnik Zoologii, 40, 77–79.
    Ovtchinnikov, S.V., Ahmad, B. & Gurko, V.O. (2009) Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia. Vestnik Zoologii, 43, 471–474.
    Roewer, C.F. (1960) Solifugen und Opilioniden - Araneae Orthognathae, Haplogynae und Entelegynae (Contribution à l'étude de la faune d'Afghanistan 23). Göteborgs Kungliga Vetenskaps och Vitterhets Samhälles Handlingar, Series 6, B8 (7), 1–53.
    WSC (2022) World Spider Catalog. Version 23.0. Natural History Museum Bern, Bern. Available from: (accessed 2 June 2022)
    Zamani, A. & Marusik, Y.M. (2021) Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species. ZooKeys, 1035, 145–193.
    Zamani, A., Mirshamsi, O., Kashani, G.M. & Karami, L. (2018) New data on the spider fauna of Iran (Arachnida: Araneae), Part V. Iranian Journal of Animal Biosystematics, 13 (2), 183–197. [for 2017]