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Type: Article
Published: 2022-07-27
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Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae)

Arachnology, Senckenberg Research Institute, Mertonstraße 17–21, 60325 Frankfurt. Institute of Ecology, Diversity and Evolution, Campus Riedberg, Goethe University, 60438 Frankfurt am Main, Germany;
Arachnology, Senckenberg Research Institute, Mertonstraße 17–21, 60325 Frankfurt.
Araneae Huntsman spiders taxonomy copulatory organs morphology Southeast Asia


When examining material of Sparassidae Bertkau, 1872 four new genera were identified as new to science, all belonging to the subfamily Heteropodinae Thorell, 1873. Three new genera from Malaysia and Brunei, and one new genus from Papua New Guinea are described in this paper: Borniella gen. nov. with its type species Borniella parva spec. nov. from Malaysia: Sarawak and Brunei (male, female); Menarik gen. nov. with its type species Menarik kecil spec. nov. from Malaysia: Sarawak (male, female); Micropoda gen. nov. with its type species Micropoda daviesae spec. nov. from Papua New Guinea: New Britain (male, female); Tiomaniella gen. nov. with its type species Tiomaniella ladam spec. nov. from Malaysia: Tioman Island (male, female). All are diagnosed by their copulatory organs, which partly exhibit extraordinary character states. Most females of Menarik kecil spec. nov. exhibited scars at their epigynes that might be connected to the special apophyses in the male palp.



  1. Banks, N. (1929) Spiders from Panama. Bulletin of the Museum of Comparative Zoology, 69, 53–96.
    Bertkau, P. (1872) Über die Respirationsorgane der Araneen. Archiv für Naturgeschichte, 38, 208–233.
    Chrysanthus, P. (1965) Spiders from south New Guinea VII. Nova Guinea, Zoology, 34, 345–369.
    Davies, V.T. (1994) The huntsman spiders Heteropoda Latreille and Yiinthi gen. nov. (Araneae: Heteropodidae) in Australia. Memoirs of the Queensland Museum, 35, 75–122.
    Bruyn, M. de, Stelbring, B., Morley, R.J., Hall, R., Carvalho, G.R., Cannon, C.H, Bergh, G. van den, Meijaard, E., Metcalfe, I., Boitani, L., Maiorano, L., Shoup, R. & Rintelen, T. von (2014) Borneo and Indochina are major evolutionary hotspots for Southeast Asia biodiversity. Systematic Biology, 63, 879–901.
    Grall, E. & Jäger, P. (2020) Forty-seven new species of Sinopod from Asia, with a considerable extension of the distribution range to the South and description of a new species group (Sparassidae: Heteropodinae). Zootaxa, 4797 (1), 1–101.
    Hall, R. (2009) Southeast Asia’s changing palaeogeography. Blumea, 54, 148–161.
    Jäger, P. (1998) First results of a taxonomic revision of the SE Asian Sparassidae (Araneae). In: Selden, P. (Ed.), Proceedings of the 17th European Colloquium of Arachnology, Edinburgh. 1997. British Arachnological Society, Burnham Beeches, Bucks, pp. 53–59.
    Jäger, P. (1999) Sinopoda, a new genus of Heteropodinae (Araneae, Sparassidae) from Asia. Journal of Arachnology, 27, 19–24.
    Jäger, P. (2000) Two new heteropodine genera from southern continental Asia (Araneae: Sparassidae). Acta Arachnologica, 49, 61–71.
    Jäger, P. (2001) Diversität der Riesenkrabbenspinnen im Himalaya. Über eine Radiation zweier Gattungen in den Schneetropen (Araneae: Sparassidae: Heteropodinae). Courier Forschungsinstitut Senckenberg, 232, 1–136.
    Jäger, P. (2002) Heteropodinae: Transfers and synonymies (Arachnida: Araneae: Sparassidae). Acta arachnologica, 51, 33–61.
    Jäger, P. (2006) Martensopoda gen. nov. from southern Indian mountain ranges, the first genus of huntsman spiders with a cymbial spur (Araneae: Sparassidae: Heteropodinae). Zootaxa, 1325 (1), 335–345.
    Jäger, P. (2012) Revision of the genus Sinopoda Jäger, 1999 in Laos with discovery of the first eyeless huntsman spider species (Sparassidae: Heteropodinae). Zootaxa, 3415 (1), 37–57.
    Jäger, P. (2020a) Platnickopoda gen. nov., a new genus of huntsman spiders from Tanzania (Araneae: Sparassidae: Heteropodinae). Arachnology, 18, 499–506.
    Jäger, P. (2020b) Thunberga gen. nov., a new genus of huntsman spiders from Madagascar (Araneae: Sparassidae: Heteropodinae). Zootaxa, 4790 (2), 245–260.
    Jäger, P. (2021a) Enigmatic “love bites” and an indirect mating plug: revision of Thunberga Jäger, 2020 with description of new species (Araneae: Sparassidae: Heteropodinae). Arachnology, 18, 718–765.
    Jäger, P. (2021b) Two new enigmatic genera of huntsman spiders from Madagascar (Araneae: Sparassidae). In: Jäger, P., Schwendinger, P. & Shear, B. (Eds.) ,A Festschrift honouring Prof. Dr. Jochen Martens on occasion of his 80th birthday. Zootaxa, 4984 (1), 335–346.
    Jäger, P. & Krehenwinkel, H. (2015) May gen. nov. (Araneae: Sparassidae): a unique lineage from southern Africa supported by morphological and molecular features. African Invertebrates, 56, 365–392.
    Jäger, P. & Minn, M.Z. (2015) New species in the family Ctenidae Keyserling, 1877 from high altitude habitats in Myanmar, with the first case of penetration of the female’s cuticle by a male in the RTA-clade (Arachnida: Araneae: Ctenidae). Zootaxa, 3994 (2), 235–252.
    Jäger, P. & Ono, H. (2002) The Sparassidae from Japan. II. First Pseudopoda species and new Sinopoda species (Araneae: Sparassidae: Heteropodinae). Acta Arachnologica, 51, 109–124.
    Jäger, P. & Vedel, V. (2007) Sparassidae of China 4. The genus Pseudopoda (Araneae: Sparassidae) in Yunnan Province. Zootaxa, 1623 (1), 1–38.
    Koch, L. (1875) Die Arachniden Australiens, nach der Natur beschrieben und abgebildet. Erster Theil. Lieferung 12–16. Bauer & Raspe, Nürnberg, pp. 577–740, pls. 44–65.
    Latreille, P.A. (1804) Tableau methodique des Insectes. Nouveau Dictionnaire d‘Histoire Naturelle Paris, 24, 129–295.
    Lessert, R. de (1921) Araignées du Kilimandjaro et du Merou (suite). 4. Clubionidae. Revue Suisse de Zoologie, 28, 381–442.
    Liu, J., Li. S. & Jäger, P. (2008) New cave-dwelling Sinopoda spiders (Araneae: Sparassidae) from southern China. Zootaxa, 1857 (1), 1–20.
    Merian, P. (1911) Die Spinnenfauna von Celebes. Beiträge zur Tiergeographie im Indoaustralischen Archipel. Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere, 31, 165–354.
    Metcalfe, I. (1998) Palaeozoic and Mesozoic geological evolution of the SE Asia region: multidisciplinary constraints and implications for biogeography. In: Hall, R. & Holloway, J.D. (Eds.), Biogeography and Geological Evolution of SE Asia. Backhuys Publishers, Leiden, pp. 25–41.
    Metcalfe, I. (2011a) Tectonic framework and Phanerozoic evolution of Sundaland. Gondwana Research, 19, 3–21.
    Metcalfe, I. (2011b) Palaeozoic-Mesozoic history of SE Asia. In: Hall, R., Cottam, M. A. & Wilson, M.E.J. (Eds.), The SE Asian Gateway: History and Tectonics of the Australia—Asia Collision. Geological Society Special Publications 355. Geological Society, London, pp. 7–35.
    Moradmand, M. (2013) The stone huntsman spider genus Eusparassus (Araneae: Sparassidae): systematics and zoogeography with revision of the African and Arabian species. Zootaxa, 3675 (1), 1–108.
    Myers, N. (1988) Threatened biotas: “Hotspots” in tropical forests. Environmentalist, 8, 1–20.
    Myers, N., Mittermeier, R.A., Mittermeier, C.G., da Fonseca, G.A.B. & Kent, J. (2000) Biodiversity hotspots for conservation priorities. Nature, 403, 853–858.
    Rheims, C.A. (2010) A new genus of huntsman spiders from the Neotropical region (Araneae: Sparassidae: Heteropodinae). Zootaxa, 2650 (1), 33–46.
    Rheims, C.A. (2021) The Neotropical genera Guadana Rheims, 2010 and Sparianthina Banks, 1929 (Araneae: Sparassidae: Heteropodinae). Zootaxa, 5061 (3), 401–431.
    Simon, E. (1880) Révision de la famille des Sparassidae (Arachnides). Actes de la Société Linnéenne de Bordeaux, 34 (2/3/4), 223–351.
    Simon, E. (1897) Etudes arachnologiques. 27e Mémoire. XLII. Descriptions d’espèces nouvelles de l’ordre des Araneae. Annales de la Société Entomologique de France, 65, 465–510.
    Simon, E. (1903) Descriptions d’arachnides nouveaux. Annales de la Société Entomologique de Belgique, 47, 21–39.
    Simon, E. (1909) Arachnides recueillis par L. Fea sur la côte occidentale d’Afrique. 2e partie. Annali del Museo Civico di Storia Naturale di Genova, 44, 335–449.
    Thorell, T. (1873) s.n. In: Remarks on synonyms of European spiders. Part IV. C. J. Lundström, Uppsala, pp. 375–645.
    Thorell, T. (1881) Studi sui Ragni Malesi e Papuani. III. Ragni dell’Austro Malesia e del Capo York, conservati nel Museo civico di storia naturale di Genova. Annali del Museo Civico di Storia Naturale di Genova, 17, 1–720.
    Thorell, T. (1890) Diagnoses aranearum aliquot novarum in Indo-Malesia inventarum. Annali del Museo Civico di Storia Naturale di Genova, 30, 132–172.
    Walckenaer, C.A. (1837) Histoire naturelle des insectes. Aptères. Tome premier. Roret, Paris, 682 pp., 15 pls.
    World Spider Catalog (2022) World Spider Catalog, Version 23.0. Natural History Museum Bern. Available from: (accessed 15 April 2022)