Abstract
Two new species-group are established for the Palaearctic species of the Geodromicus Redtenbacher, 1857: the bodemeyeri-group containing G. (s.str.) bodemeyeri Bernhauer, 1902 (Russia (Caucasus), Turkey), G. (s.str.) championi Cameron, 1924 (Pakistan, India, Nepal) and G. (s.str.) zwickianus Fagel, 1976 (Turkey, Iran); the convexicollis-group containing G. (s.str.) convexicollis Luze, 1903 (Middle Asia), G. (s.str.) kashmirensis Cameron, 1930 (Afghanistan, Pakistan, India), G. (s.str.) lebedevianus Roubal, 1929 (Kyrgyzstan), G. (s.str.) macrothorax Kashcheev, 1999 (Kazakhstan, Kyrgyzstan), G. (s.str.) malcolmi sp.n. (India: Uttarakhand) and G. (s.str.) medvedevi Shavrin, 2009 (Pakistan). All these species are (re-)described (except of G. (s.str.) medvedevi) and illustrated. The neotype for G. lebedevianus is designated. Keys to species of the bodemeyeri-group and the convexicollis-group are provided. Five species from the Himalayan Region are redescribed and illustrated: G. (s.str.) affinis Cameron, 1934 and G. (s.str.) alticola Cameron, 1945 (both from Pakistan, India), G. (s.str.) curvipes Cameron, 1924 (India, Nepal), G. (s.str.) ruficornis Cameron, 1941 (India: Kashmir) and G. (s.str.) subsimilis Cameron, 1941 (India, Nepal). Aedeagi for G. (s.str.) curvipes and G. (s.str.) subsimilis are figured for the first time. The following new synonymies are established: G. (s.str.) bodemeyeri = G. convexus Iablokoff-Khnzorian, 1962 syn.n., = G. (s.str.) rivularis Iablokoff-Khnzorian, 1962 syn.; G. (s.str.) championi = G. nepalicus Coiffait, 1977 syn.n.; G. (s.str.) convexicollis = G. cylindricus Tronquet, 1981 syn.n.; G. (s.str.) kashmirensis = G. similis Cameron, 1930 syn.n., = G. consors Cameron, 1945 syn.n., = G. afghanicus Scheerpeltz, 1961 syn.n., = G. karakorus Coiffait, 1977 syn.n.; G. (s.str.) affinis = G. torrentis Puthz, 1973 syn.n.; G. (s.str.) alticola = G. peraffinis Cameron, 1945 syn.n., = G. brancuccii Coiffait, 1984 syn.n. Lectotypes are designated for: G. championi, G. kashmirensis, G. similis and G. curvipes. Distribution maps for all species are provided. Geodromicus (s.str.) championi and G. (s.str.) subsimilis are recorded from Pakistan and Nepal respectively for the first time. Additionally, faunistic records of five species from the Eastern Palaearctic Region are provided. The habitus of G. (s.str.) medvedevi and G. (s.str.) smetanai Shavrin, 2018 are illustrated for the first time.
References
Bernhauer, M. (1902) Neue Staphyliniden der paläarktischen Fauna nebst Bemerkungen. Deutsche Entomologische Zeitschrift, 1901, 241–251.
Bernhauer, M. (1915) Neue Staphyliniden des paläarktischen Faunengebietes. Wiener Entomologische Zeitung, 34, 69–81. https://doi.org/10.5962/bhl.part.10619
Bordoni, A. (1984) Appunti per una revisione dei Geodromicus Redt. della regione Palearctica occidentale (Coleoptera, Staphylinidae). Redia, 67, 19–59.
Cameron, M. (1924). New species of Staphylinidae from India. Transactions of the Entomological Society of London, 1924, 160–198. https://doi.org/10.1111/j.1365-2311.1924.tb03357.x
Cameron, M. (1930) The fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Vol. 1. Taylor and Francis, London, xvii + 471 pp.
Cameron, M. (1934) Report on Coleoptera of the family Staphylinidae. Memoirs of the Connecticut Academy of Arts and Sciences, 10, 17–21.
Cameron, M. (1941) Descriptions of new Staphylinidae (Coleopt.). The Proceedings of the Royal Entomological Society of London (B), 10, 142–147.
Cameron, M. (1945) Descriptions of new Staphylinidae (Coleoptera). The Proceedings of the Royal Entomological Society of London (B), 14, 63–69.
Cheng, Z.-F., Shavrin, A.V. & Peng, Z. (2020) New species and records of Geodromicus Redtenbacher, 1857 from China (Coleoptera: Staphylinidae: Omaliinae: Anthophagni). Zootaxa, 4789 (1), 132–170. https://doi.org/10.11646/zootaxa.4789.1.4
Coiffait, H. (1970) Staphylinidae d’Ouzbekistan. L’Entomologiste, 26, 142–148.
Coiffait, H. (1977) Staphilinides récoltés au Népal par le Professeur Franz (2e partie). Bulletin de la Société d’Histoire Naturelle de Toulouse, 112 (1976), 243–275.
Coiffait, H. (1981) Staphylinides nouveaux du Moyen Orient appartenant au Musée Hongrois d’Histoire Naturelle (Coleoptera). Folia Entomologica Hungarica, 34 (1), 15–23.
Coiffait, H. (1984) Staphylinides (Col.) de la région himalayenne et de l’Inde. II. Tachyporinae, Oxytelinae et Aleocharinae. Entomologica Basiliensia, 9, 116–157.
Fagel, G. (1976) Contribution à la connaissance des Staphylinidae. CXVI. Les Geodromicus d’Anatolie. Revue suisse de Zoologie, 83 (2), 381–391.
Herman, L.H. (2001) Catalogue of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millenium. Bulletin of the American Museum of Natural History, 265 (1–7), i–vi + 1–421. https://doi.org/10.1206/0003-0090.265.1.1
Hlaváč, P., Kocian, M. & Nakládal, O. (2016) Updated list of the subfamily Omaliinae (Coleoptera: Staphylinidae) from Kazakhstan with some new records. Turkish Journal of Zoology, 40, 1−8. https://doi.org/10.3906/zoo-1505-29
Iablokoff-Khnzorian, S.M. (1989) Dva novykh Kavkaszkikh predstavitelia iz roda Geodromicus Redt. (Coleoptera Staphylinidae) [Two new Caucasian representatives of the genus Geodromicus (Coleoptera Staphylinidae)]. Doklady Akademii Armyanskoy SSR, 87 (1988], 134–139. [in Russian, with Abstract in Armenian]
Kashcheev, V.A. (1999) Novye vidy Oxytelinae (Coleoptera, Staphylinidae) fauny Kazakhstana [New species of Oxytelinae (Coleoptera, Staphylinidae) of the fauna of Kazakhstan]. Selevinia, 1996, 12–16. [in Russian]
Kashcheev, V.A & Ishkov, E.V. (2001) List of beetles (Coleoptera) of Aksu-Dzhabagly Natural Reserve. Tethys Entomological Research, 3, 99–108.
Khnzorian, S.M. (1962) Novye vidy zhestkokrylykh iz Zakavkazya [New species of Coleoptera from Transcaucasus (Insecta-Coleoptera)]. Zoologichesky Sbornik Zoologichesky Institut, Akademii Nauk Armyanskoy SSR, 12, 99–124. [in Russian]
Luze, G. (1903) Revision der paläarktischen Arten der Staphyliniden-Gattung Geodromicus Redtenb. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, 53, 103–117.
Luze, G. (1904) Beitrag zur Staphyliniden-Fauna von Russisch-Central-asien (Coleoptera). Horae Societatis Entomologicae Rossicae, 37, 74–115.
Newton, A.F. (2022) StaphBase: Staphyliniformia world catalog database (version October 2021): Staphylinoidea, Hydrophiloidea, Synteliidae, Sphaeritidae, Cretohisteridae. Catalogue of Life Checklist. October 2021. Species 2000: Naturalis, Leiden. Availabe from: http://www.catalogueoflife.org/ (accessed 18 June 2022).
Puthz, V. (1973) A new Geodromicus from Pakistan (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 80 (2), 517–519.
Roubal, J. (1929) Vier neue Coleopteren aus SSSR. Entomologische Blätter, 25 (1), 46–48.
Scheerpeltz, O. (1961) The 3rd Danish expedition to Central Asia. Zoological results 26. Staphylinidae (Insecta) aus Afghanistan. Videnskabelige Meddelelser fra Dansk naturhistorisk Forening i Københaven, 123, 33–50.
Schülke, M. & Smetana, A. (2015) Staphylinidae. In: Löbl, I. & Löbl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea—Staphylinoidea. Revised and Updated Edition. Brill, Leiden, pp. 304–1134.
Sharp, D. (1889) The Staphylinidae of Japan. The Annals and Magazine of Natural History, Series 6, 3 (18), 463–476.
Shavrin, A.V. (2000) A new species of the staphylinid genus Geodromicus Redt. (Coleoptera, Staphylinidae) from Southern Siberia. Entomologicheskoe Obozrenie, 79 (1), 87–89. [in Russian, with Abstract in English]
Shavrin, A.V. (2009) A new species and new records of Geodromicus Redtenbacher, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from Pakistan. Caucasian Entomological Bulletin, 5 (2), 183–184.
Shavrin, A.V. (2015) New data on Himalayan Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae) with description of a new species from Nepal. Zootaxa, 3973 (3), 528–538. http://doi.org/10.11646/zootaxa.3973.3.7
Shavrin, A.V. (2017) Five new species of the genus Philydrodes Bernhauer, 1929 from China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa, 4231 (2), 169–186. https://doi.org/10.11646/zootaxa.4231.2.2
Shavrin, A.V. (2018) The lestevoides species group of the genus Geodromicus Redtenbacher, 1857 (Insecta: Coleoptera: Staphylinidae: Omaliinae). Zootaxa, 4378 (2), 151–190. https://doi.org/10.11646/zootaxa.4378.2.1
Shavrin, A.V. (2019) The crassipalpis species group of the genus Geodromicus Redtenbacher, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa, 4686 (4), 571–580. https://doi.org/10.11646/zootaxa.4686.4.6
Shavrin, A.V. (2021) The rousi species group of the genus Geodromicus Redtenbacher, 1857 (Coleoptera: Staphylinidae: Omaliinae). Caucasian Entomological Bulletin, 17 (1), 191–202. https://doi.org/10.23885/181433262021171-191202
Shavrin, A.V. (2022) The plagiatus species group of the genus Geodromicus Redtenbacher, 1857 (Insecta: Coleoptera: Staphylinidae: Omaliinae), and additional data on some species from Japan. Zootaxa, 5188 (5), 401–429. https://doi.org/10.11646/zootaxa.5188.5.1
Tichomirova, A.L. (1973) Opisanie novykh vidov [Descriptions of new species]. In: Gilyarov, M. (Ed.), Ekologiya pochvennykh bespozvonochnykh [Ecology of the soil invertebrates]. Nauka, Moscow, pp. 144–173. [in Russian]
Tronquet, M. (1981) Staphylinidae d’Afghanistan (Coleoptera). Revue Française d’Entomologie, 3 (3), 69–83.
Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus-Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105–142.