Skip to main content Skip to main navigation menu Skip to site footer
Published: 2022-12-01

Revisional notes on Iolaus aequatorialis Stempffer Bennett, 1958 and related species in the subgenus Philiolaus Stempffer Bennett, 1958 (Lepidoptera, Lycaenidae, Theclinae), with description of two new species

African Butterfly Research Institute, P.O. Box 14308, 0800 Nairobi, Kenya.
African Butterfly Research Institute, P.O. Box 14308, 0800 Nairobi, Kenya.
Lepidoptera taxonomic revision Iolaus aequatorialis species group Iolaus mane Iolaus theodori Iolaus ivani sp. n. Iolaus brazza sp. n. male genitalia


Similarities in male and female genitalic structure isolate a small group of taxa in the genus Iolaus Hübner, [1819], subgenus Philiolaus Stempffer & Bennett, 1958, related to Iolaus aequatorialis Stempffer & Bennett, 1958. They are assigned to the newly proposed I. aequatorialis species group with two new species recognised and described. This action also clarifies previous taxonomic misunderstandings involving a few other species with similar facies because of a lack of information on male and female genital morphology.



  1. Collins, S.C., Congdon, T.C.E., Henning, G.A., Larsen, T.B. & Williams, M.C. (2014) A review of d’Abrera’s Butterflies of the Afrotropical Region—Part III (second edition), 2009. Part 3 (Theclinae, Polyommatinae and Riodinidae). Metamorphosis, 25, 109–122.
    Collins, S.C. & Larsen, T.B. & Warren-Gash, H. (2003) Notes on Afrotropical butterflies with the description of eleven new species and four new subspecies (Lepidoptera: Rhopalocera)—ABRI Research Paper No. 3. Metamorphosis, 14 (3), 63–110.
    d’Abrera, B. (2009) s.n. In: Butterflies of the Afrotropical Region. Part III. Lycaenidae, Riodinidae. New and Revised Edition. Hill House Publishers, Melbourne and London, pp. i–xxvi + 531–880.
    Heath, A., Newport, M.A. & Hancock, D. (2002) The butterflies of Zambia. African Butterfly Research Institute and The Lepidopterists’ Society of Africa, Nairobi, 137 pp., [CD-Rom]
    Larsen, T.B. (1991) The Butterflies of Kenya and their Natural History. Oxford University Press, Oxford, 490 pp., 22 colour pls.
    Larsen, T.B. (2005) Butterflies of West Africa. Apollo Books, Svendborg, 595 pp., 135 colour pls.
    Miller, L.D. (1970) Nomenclature of wing veins and cells. Journal of Research on the Lepidoptera, 8, 37–48.
    Sáfián, Sz. (2021) The butterfly fauna (Lepidoptera: Papilionoidea) of Liberia. PhD thesis. University of Sopron, Sopron, 260 pp., 101 colour pls.
    Sáfián, Sz. , Bayliss, J. & Congdon, T.C.E. (2022) Description of four Iolaus Hübner, 1819 species in the subgenus Philiolaus Stempffer & Bennett, 1958 from East Africa, assigned to the proposed I. maritimus species group (Lepidoptera, Lycaenidae, Theclinae). Zootaxa, 5099 (1), 46–64.
    Sáfián, Sz. , Belcastro, C., Boireau, P. & Collins, S.C. (2020) New taxa of skipper butterflies (Lepidoptera, Hesperiidae) from tropical Africa. Metamorphosis, 31 (1), 56–71.
    Sáfián, Sz. , Collins, S.C., Warren-Gash, H. & Belcastro, C. (2021) Description of five new species of Epitola sensu lato (Lepidoptera: Lycaenidae: Poritiinae) from West and Central Africa. Zootaxa, 4981 (3), 554–576.
    Stempffer, H. (1970) Contribution à l’étude des Lycaenidae de la faune éthiopiènne. Bulletin de la l’Institut Fondamental d’Afrique Noire, Series A, 32, 767–772.
    Stempffer, H. & Bennett, N.H. (1958) Revision des genres appartenant au groupe des Iolaus (Lep. Lycaenidae). (Premiere Partie). Bulletin de la l’Institut Francais d’Afrique Noire, Series A, 20, 1243–1347.