Abstract
Twenty-six genera and five subgenera of the tribe Smicromyrmini are reviewed. The tribe (528 valid species) is Afrotropical (15 genera /126 species), Palaearctic (13/213) and Oriental (14/204) in distribution. The females are known for 80%, and males for 87%, of the 26 genera and five subgenera. The hitherto unknown female of Rasnitsynitilla Lelej in Lelej & van Harten, 2006 is described. New combinations are proposed for Rasnitsynitilla bartolozzii (Lo Cascio et Grita, 2014), comb. nov. (from the genus Smicromyrme Thomson, 1870) and Skorikovia sanguinicollis (Klug, 1829), comb. nov. (from the genus Ephutomma Ashmead, 1899). The genus Indratilla Lelej, 1993 is newly recorded from India and the subgenus Eremotilla Lelej, 1985 from the Oriental Region. Keys to females and males of these genera and subgenera of Smicromyrmini are provided. Habitus photos of males and/or females are provided for each genus and subgenus.
References
André, E. (1896) Notes pour servir à la connaissance de mutilles paléarctiques et description de quelques espèces nouvelles. Deuxieme partie. Mémoires de la Société Zoologique de France, 9, 261–277.
André, E. (1898) Étude sur les mutillides du Muséum de Paris. Annales de la Société Entomologique de France, 67, 1–79.
André, E. (1899–1902) Species de Hyménoptères d’Europe & d’Algérie. Tome Huitième. Les Mutillides. Vve Dubosclard and A. Hermann, Paris, 480 pp., 15 pls. [pp. 1–64, pls. I–VIII (1899b), pp. 65–144 (1900), pp. 145–304 (1901a), pp. 305–384 (1902b), pp. 385–478 (1902a), pls. IX–XV (unknown date)]
André, E. (1902b [“1903”]) Fam. Mutillidae. In: Wytsman, P., Genera Insectorum. Fasc. 11. s.n., Bruxelles, pp. 1–77 + 3 pls.
André, E. (1905) Mutillides nouveaux ou peu connues de l’Ancien Monds et de l’Australie. Zeitschrift für systematische Hymenopterologie und Dipterologie, 5 (1–6), 201–216 + 265–277.
Arnold, G. (1946) New species of African Hymenoptera. No. 6. Occasional Papers of the National Museum of Southern Rhodesia, 12, 49–97.
Arnold, G. (1956) New species of African Hymenoptera. No. 12. Occasional Papers of the National Museum of Southern Rhodesia, 3(21B), 52–77.
Arnold, G. (1962) New species of African Hymenoptera. No. 16. Occasional Papers of the National Museum of Southern Rhodesia, 3 (26B), 844–855.
Ashmead, W.H. (1899) Superfamilies in the Hymenoptera and generic synopses of the families Thynnidae, Myrmosidae, and Mutillidae. Journal of the New York Entomological Society, 7, 45–60.
Ashmead, W.H. (1900–1904) Classification of the fossorial, predaceous and parasitic wasps, or the superfamily Vespoidea. The Canadian Entomologist, 32 (1900), 34 (1902), 35 (1903), 36 (1904), 145–155 + 185–188 + 295–296, 79–88 + 131–137 + 163–166 + 203–210 + 219–231 + 268–273 + 287–291, 3–8 + 39–44 + 95–107 + 155–158 + 199–205 + 303–310 + 323–332, 5–9. https://doi.org/10.4039/Ent32145-5 https://doi.org/10.4039/Ent3479-4 https://doi.org/10.4039/Ent353-1
Bischoff, H. (1920–1921) Monographie der Mutilliden Afrikas. Archiv für Naturgeschichte, 86A (1–3) & 86A (4–5), 1–480 (1920) & 481–830 (1921) .
Brothers, D.J. (1975) Phylogeny and classification of the aculeate Hymenoptera, with special reference to Mutillidae. The University of Kansas Science Bulletin, 50 (11), 483–648.
Brothers, D.J. (1983) Identity and classification of Physetopoda, Chaetotilla and Paramyrme. Journal of the Kansas Entomological Society, 56 (3), 441–445.
Brothers, D.J. (1999) Phylogeny and evolution of wasps, ants and bees (Hymenoptera, Chrysidoidea, Vespoidea and Apoidea). Zoologica Scripta, 28, 233–249. https://doi.org/10.1046/j.1463-6409.1999.00003.x
Brothers, D.J. (2022) Critical analysis of the type material of Mutillidae described from the Australasian Region (Hymenoptera). Zootaxa, 5140 (1), 1–215. https://doi.org/10.11646/zootaxa.5140.1.1
Brothers, D.J. & Lelej, A.S. (2017) Phylogeny and higher classification of Mutillidae (Hymenoptera) based on morphological reanalyses. Journal of Hymenoptera Research, 60, 1–97.
https://doi.org/10.3897/jhr.60.20091
Cameron, P. (1902) Descriptions of new genera and species of Hymenoptera collected by Major C. S. Nurse at Deesa, Simla and Ferozepore. Part I. Journal of the Bombay Natural History Society, 14 (2), 267–293, 1 pl.
Chen, C.-W. (1957) A revision of the velvety ants or Mutillidae of China (Hymenoptera). Quarterly Journal of the Taiwan Museum, 10 (3–4), 135–224, 6 pls.
Dalla Torre, C.G. de (1897) Catalogus Hymenopterorum hucusque Descriptorum Systematicus et Synonimicus. Vol. VIII. Fossores (Sphegidae). Guilelmi Engelmann, Lipsiae [Leipzig], [4] + viii + 750 pp. https://doi.org/10.5962/bhl.title.10348
De Stefani, T. (1887) Un nuovo genere di crabronidi ed altri imenotteri nuovi o poco cogniti raccolti in Sicilia. Naturalista Siciliano, 6, 59–62 + 85–90 + 110–114 + 143–147, tab. 2.
Fabricius, I.C. (1787) Mantissa Insectorum sistens eorum Species nuper detecta, adjectis Characteribus Genericis, Differentiis Specificis, Emendationibus, Observationibus. Tom. 1. Christian Gottlob Proft, Hafnia [Copenhagen], 348 pp.
https://doi.org/10.5962/bhl.title.36471
Fabricius, J.C. (1804) s.n. In: Systema Piezatorum secundum Ordines, Genera, Species adjectis Synonymis, Locis, Observationibus, Descriptionibus. Carolum Reichard, Brunsvigae [Braunschweig], pp. i–xiv + 15–440 + 1–30. https://doi.org/10.5962/bhl.title.10490
Food and Agriculture Organisation of the United Nations [FAO] (1975) Final Rapport. Rapport préparé pour le Gouvernement de la République Unie du Cameroun par l’Organisation pour l’Alimentation et l’Agriculture des Nations Unies agissant en Qualité d’Agence d’Exécution pour le Programme des Nations Unies pour le Developpement basé sur les Activités du Prof. Dr. Guido Nonveiller Expert en Entomologie Agricole. FAO, Rome and Université de Yaounde, École Nationale Superieure Agronomique, Laboratoire d’Entomologie, Nkolbisson, 2 + xii + 184 pp.
Hoffer, A. (1936) Nové kudulky (Mutillidae, Нуmеnорteга, Vespoidea) z Československé republiky (Diagnoses novae mulillidarum Čechosloveniae). Časopis České Společnosti Entomologické, 33, 157–163. [in Czech]
Hymenoptera Anatomy Ontology (2013) Hymenoptera Glossary. Available from: http://glossary.hymao.org (accessed 19 July 2022)
Integrated insect types database of Taiwanese species Available from: http://twinsecttype.nmns.edu.tw/specimen/?id=NMNS-SDEI-01389 (accessed 15 July 2022)
Invrea, F. (1950) Mutillidi nuovi o notevoli del Bacino Mediterraneo (Hymenoptera, Mutillidae). Memorie della Società Entomologica Italiana, 29, 19–27.
Invrea, F. (1964) Mutillidae—Myrmosidae. Fauna d’Italia. Vol. 5. Edizioni Calderini, Bologna, i–xii + 302 + [1] pp.
Kimsey, L.S. & Brothers, D.J. (2016) The life, publications and new taxa of Qabir Argaman (Carol Nagy). Journal of Hymenoptera Research, 50, 141–178. https://doi.org/10.3897/JHR.50.7973
Klug, J.C.F. (1829) Symbolae Physicae seu Icones et Descriptiones Insectorum quae ex Itinere per Africam Borealem et Asiam Occidentalem Friderici Guilelmi Hemprich et Cristiani Godofredi Ehrenberg Studio Novae aut Illustratae redierunt. III, Insectorum. Hymenoptera, Decas Prima. Berolini, [17] pp. [and Symbolae Physicae seu Icones et Descriptiones Insectorum quae ex Itineribus per Libyam, Aegyptum, Nubiam, Dongalam, Syriam Arabiam et Habbesiniam Publico Institutis sumptu Friderici Guilelmi Hemprich et Cristiani Godofredi Ehrenberg Medicinae et Chirurgiae Doctorum, Studio Annis MDCCXX–MDCCCXXV redierunt. III, Insectorum. Hymenoptera, Decas Prima. Berolini, 10 pls.]
Latreille, P.A. (1802) Histoire naturelle, générale et particulière des Crustacés et des Insectes. 3. F. Dufart, Paris, xii + 467 pp. https://doi.org/10.5962/bhl.title.15764
Latreille, P.A. (1825) Famillies Naturelles du Règne Animal, exposées succinctement et dans un Ordre Analytique, avec l’Indication de leurs Genres. Baillière, Paris, 570 pp. https://doi.org/10.5962/bhl.title.16094
Latreille, P.A. (1829) Suite et fin des Insectes. In: Cuvier, G., Le Règne Animal distribué d’après son Organisation, pour servir de Base a l’Histoire Naturelle des Animaux et d’Introduction a l’Anatomie Comparée. Nouvelle Édition, revue et augmentée. Tome V. Déterville, Paris, pp. i–xxiv + 1–556.
Lelej, A.S. (1977) New data on the velvet ant fauna of Mongolia (Hymenoptera, Mutillidae). Insects of Mongolia, 5, 526–536. [in Russian]
Lelej, A.S. (1979) A new mutillid species of the genus Smicromyrme Thomson (Hymenoptera, Mutillidae). Trudy vsesoyuznogo entomologicheskogo obshchestva, 61, 169–171. [in Russian]
Lelej, A.S. (1984). Velvet ants of the genus Раrаmуrmе (Hymenoptera, Mutillidae). In: Lehr, P.A. (Ed.), Systematica Nasekomykh Dalnego Vostoka. DVNC AN SSSR, Vladivostok, pp. 101–115. [in Russian]
Lelej, A.S. (1985) Osy-nemki (Hymenoptera, Mutillidae) Fauny SSSR i Sopredelnykh Stran [The Velvet Ants (Hymenoptera, Mutillidae) of the USSR and Neighbouring Countries]. Nauka, Leningrad, 268 pp. [in Russian]
Lelej, A.S. (1993) A new genus of velvet ants from Sri Lanka (Insecta, Hymenoptera, Mutillidae). Spixiana, 16 (3), 233–236.
Lelej, A.S. (1995a) A review of Palaearctic and Oriental species of genus Nemka Lelej with description of Oriental genus Mickelomyrme gen. n. (Hymenoptera, Mutillidae). Far Eastern Entomologist, 6, 1–20.
Lelej, A.S. (1995b) To the knowledge of East Asian species of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera, Mutillidae) with description of four new genera and eight new species. Far Eastern Entomologist, 13, 1–28.
Lelej, A.S. (1995c) To the knowledge of the velvet ants (Hymenoptera, Mutillidae) of Rajasthan, Western India. Far Eastern entomologist, 20, 1–11.
Lelej, A.S. (1996a) Mutillid wasps collected in Malaysia and Indonesia by Dr. Sk. Yamane (Hymenoptera, Mutillidae). Tropics, 6 (1–2), 91–104.
https://doi.org/10.3759/tropics.6.91
Lelej, A.S. (1996b) A review of the East Asian species of Mickelomyrme Lelej, 1995 (Hymenoptera, Mutillidae). Entomofauna, 17 (15), 277–292.
Lelej, A.S. (2002) Catalogue of the Mutillidae (Hymenoptera) of the Palaearctic Region. Dalnauka, Vladivostok, 172 pp.
Lelej, A.S. (2005) Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region. Dalnauka, Vladivostok, 252 pp.
Lelej, A.S. (2007) Biogeography of mutillid wasps (Hymenoptera, Mutillidae). In: Rasnitsyn, A.P. & Gokhman, V.E. (Eds.), Studies on Hymenopterous Insects. Collection of Scientific Papers. KMK Scientific Press Ltd., Moscow, pp. 82–111. [in Russian]
Lelej, A.S. & Brothers, D.J. (2008) The genus-group names of Mutillidae (Hymenoptera) and their type species, with a new genus, new name, new synonymies, new combinations and lectotypifications. Zootaxa, 1889 (1), 1–79.
https://doi.org/10.11646/zootaxa.1889.1.1
Lelej, A.S. & Kabakov, O.N. (1980) On Bradynobaenidae and Mutillidae (Hymenoptera) of Afghanistan. Entomologicheskoe Obozrenie, 59 (1), 181–196. [in Russian, English translation: (1981) Entomological Review, 59 (1), 141–156]
Lelej, A.S. & Nemkov, P.G. (1997) Phylogeny, evolution and classification of Mutillidae (Hymenoptera). Far Eastern Entomologist, 46, 1–24.
Lelej, A.S. & Osten, T. (2004) To the knowledge of the mutillid and bradynobaenid wasps of Iran (Hymenoptera: Mutililidae, Bradynobaenidae). Proceeding of the Russian Entomological Society, St. Petersburg, 75 (1), 253–262.
Lelej, A.S., Ullah, M. & Mahmood, K. (2007) Additions to the knowledge of the Mutillidae (Hymenoptera) of Pakistan. Zootaxa, 1444 (1), 53–60.
https://doi.org/10.11646/zootaxa.1444.1.4
Lelej, A.S. & van Harten, A. (2006) A review of the Mutillidae (Hymenoptera) of Yemen. Zootaxa, 1226 (1), 1–50. https://doi.org/10.11646/zootaxa.1226.1.1
Lelej, A.S. & van Harten, A. (2011) Mutillidae (Hymenoptera) of the United Arab Emirates. Part 2. Genus Dentilla Lelej. Russian Entomological Journal, 20 (3), 299–302.
Lelej, A.S. & Yildirim, E. (2009) A review of the Mutillidae (Hymenoptera) of Turkey. Zootaxa, 2160, 1–28. https://doi.org/10.11646/zootaxa.2160.1.1
Lo Cascio, P. (2015) Worldwide checklist of the island mutillid wasps (Hymenoptera Mutillidae). Biodiversity Journal, 6 (2), 529–592.
Lo Cascio, P. & Grita, F. (2014) A new species of Smicromyrme from Israel (Hymenoptera: Mutillidae). Fragmenta entomologica, Roma, 46 (1–2), 125–128. https://doi.org/10.4081/fe.2014.76
Lucas, H. (1848) Exploration Scientifique de l’Algérie pendant des Années 1840, 1841, 1842. Sciences Physiques, Zoologie, III. Histoire Naturelle des Animaux Articulés, Troisieme Partie, Insectes. Imprimerie Nationale, Paris, [6] + 527 pp. [and Zoologie, IV. Histoire Naturelle des Animaux Articulés, Atlas. Imprimerie Nationale, Paris, 122 pls.]
Magretti, P. (1884) Risultati di raccolte imenotterologiche nell’Africa Orientale. Annali del Museo Civico di Storia Naturale di Genova, Series 2°, 1 (21), 523–636, pl. 1.
Magretti, P. (1892 [“1890–1891”]) Di alcune specie d’Imenotteri raccolte dall’Ing. L. Bricchetti-Robecchi nel paese dei Somali. Museo Civico di Storia Naturale di Genova. Series 2°, 10 (30), 950–960.
Magretti, P. (1906 [“1905”]) Materiali per la conoscenza della Fauna Eritrea. Insetti. Ord. Imenotteri. Parte I. Fam. Mutillidi. Bolletino della Società Entomologica Italiana, 37 (1), 3–96.
Mickel, C.E. (1933) The Mutillidae of Formosa. Annals of the Entomological Society of America, 26 (2), 381–423. https://doi.org/10.1093/aesa/26.2.381
Morawitz, F. (1893) Catalog der von D. Glasunov in Turkestan gesammelten Hymenoptera Fossoria. Horae Societatis Entomologicae Rossicae, 27 (3–4), 391–428.
Muskovits, J. & György, Z. (2011) Velvet ants of Hungary / Magyarooszág hangyadarazsai. Budapest: Kaposvár, 198 pp. (Natura Somogyiensis, 18).
Nagy, C.G. (1966) Ein neues Mutilliden-Subgenus aus Rumänien (Hymenoptera). Reichenbachia, 6 (13–34), 113–117.
Nagy, C.G. (1972) Two new species of Mutillidae (Hymenoptera) from Asia Minor and description of female Mutilla erronea. Bulletin of the Iraq Natural History Museum, 5 (2), 4–9.
Nonveiller, G. (1979 [“1978”]) Recherches sur les Mutillides de l’Afrique (Hymenoptera, Mutillidae). VIII. Révision des genres Ctenotilla, Cephalotilla et Pseudocephalotilla sensu Bischoff. Memoires publies par l’Institut pour la Protection des Plantes, Beograd, 13, 1–184.
Nonveiller, G. (1980) Recherches sur les Mutillides de l’Afrique (Hymenoptera, Mutillidae). X. Bref aperçu des résultats des recherches sur la faune des Mutillides du Cameroun, effectuées au cours de la période de 1962–1975. Memoires publies par l’Institut pour la Protection des Plantes, Beograd, 14, 11–68.
Nonveiller, G. (1993) Studies on African Mutillidae. XIV. Ctenoceraea, a new genus with pectinate antennae and description of the type-species C. pectinella sp. n. (Hymenoptera). Revue Française d’Entomologie, New Series, 15 (1), 15–18.
Nurse, C.G. (1903) New species of Indian Aculeate Hymenoptera. The Annals and Magazine of Natural History, Series 7, 11, 393–403 + 511–526 + 528–549. https://doi.org/10.1080/00222930308678789
Okayasu, J. (2017) Two new species of the genus Sinotilla Lelej, 1995 (Hymenoptera: Mutillidae), with notes on taxonomic characters. Zootaxa, 4294 (2), 151–169. https://doi.org/10.11646/zootaxa.4294.2.1
Okayasu, J. (2018) Taxonomic review of Chin-wen Chen’s species described in the genus Smicromyrme (Hymenoptera: Mutillidae: Smicromyrmini). Acta Entomologica Musei Nationalis Pragae, 58 (2), 479–494. https://doi.org/10.2478/aemnp-2018-0036
Okayasu, J. (2020) Velvet ants of the tribe Smicromyrmini Bischoff (Hymenoptera: Mutillidae) of Japan. Zootaxa, 4723 (1), 1–110.
https://doi.org/10.11646/zootaxa.4723.1.1
Okayasu, J. (2022) Remarkable range extension of the genus Nordeniella Lelej, 2005 (Hymenoptera: Mutillidae: Smicromyrmini): Description of a new species from Sumbawa, Indonesia. Journal of Insect Biodiversity, 32 (2), 56–63. https://doi.org/10.12976/jib/2022.32.2.3
Okayasu, J., Williams, K.A. & Lelej, A.S. (2018) A remarkable new species of the genus Sinotilla Lelej (Hymenoptera: Mutillidae: Smicromyrmini) from Taiwan and an overview of color diversity in East Asian mutillid females. Zootaxa, 4446 (3), 301–324.
https://doi.org/10.11646/zootaxa.4446.3.1
Okayasu, J., Williams, K.A., Lelej, A.S. & Pham, T.H. (2021) Review of female Andreimyrme Lelej (Hymenoptera: Mutillidae: Smicromyrmini). Zootaxa, 5061 (1), 1–38.
https://doi.org/10.11646/zootaxa.5061.1.1
Ovtchinnikov, S.V. (2002) Materials to the fauna of velvet ants (Hymenoptera, Mutillidae) of Kyrghizia. In: Zonstein, S.L. (Ed.), Entomological Investigations in Kyrgystan. Vol. 22. Kyrghyz Society of Entomology, Bishkek, pp. 79–104. [in Russian]
Pagden, H.T. (1949) Description and records of Austro-Malaysian Methocidae and Mutillidae (Hymenoptera). Transactions of the Royal Entomological Society of London, 100 (8), 191–231. https://doi.org/10.1111/j.1365-2311.1949.tb01431.x
Pagliano, G., Brothers, D.J., Cambra, R., Lelej, A.S., Lo Cascio, P., Matteini Palmerini, M., Scaramozzino, P.L., Williams, K.A. & Romano, M. (2020 [“2018”]) Checklist of names in Mutillidae (Hymenoptera), with illustrations of selected species. Bollettino del Museo Regionale di Scienze Naturali di Torino, 36 (1–2), 5–425.
Pagliano, G. & Strumia, F. (2007) Contributo alla conoscenza dei Mutillidae italiani (Hymenoptera, Scolioidea). Bollettino del Museo Regionale di Scienze Naturali di Torino, 24 (1), 25–110.
Pallas, P.S. (1773) Reisen durch verschiedene Provinzen des Russischen Reichs. Zweiter Theil. Zweites Buch vom Jahr 1771. Kayserlichen Academie der Wissenschaften, St. Petersburg, [12] + 744 pp.
Panzer, G.W.F. (1792–1844) s.n. In: Faunae Insectorum Germaniae initia oder Deutschlands Insecten. Hf. 39–48 (1797), Hf. 49–60 (1798), Hf. 61–72 (1799), Hf. 73–79 (1800), Hf. 80–85 (1801), Hf. 97–99 (1805), Hf. 101–107 (1806–1809). s.n., Nürnberg, pls. 18–20 (Mutillidae: Hf. 46), pl. 24 (Mutillidae: Hf. 55), pls. 19–20 (Mutillidae: Hf. 62), pl. 20 (Mutillidae: Hf. 76), pl. 22 (Mutillidae: Hf. 80) + pl. 20 (Hf. 83) + pl. 14 (Hf. 85), pl. 20 (Mutillidae: Hf. 97), pl. 21 (Mutillidae: Hf. 106). [dates of publication after Sherborn, 1923]
Panzer, G.W.F. (1806) Entomologischer Versuch die Jürineschen Gattungen der Linnéschen Hymenopteren nach dem Fabriziusschen System zu prufen: im Bezug auf die in der Deutschen Insektenfaune bekannt gemachten Gattungen und Arten dieser Klasse. Kritische Revision der Insektenfaune Deutschlands nach dem System bearbeitet. I–C. Heft. II. Bändchen mit zwey illuminirten Kupfertafeln. Felssekerschen Buchhandlung, Nürnberg, [14] + 270 pp., pls. 1–2. https://doi.org/10.5962/bhl.title.6539
Péringuey, L. (1899) A contribution to the knowledge of South African Mutillidae (Order Hymenoptera). Annals of the South African Museum, 1 (2 & 3), 352–378, pl. 8 & 451.
Péringuey, L. (1914) Notes on South African Mutillidae (Hymenoptera), with descriptions of new or little known species. Annals of the South African Museum, 10 (10), 323–358.
Peters, W.C.H. (1858 [„1857“]) Übersicht der von ihm in Mossambique aufgefundenen und von Hrn. Dr. Gerstäcker bearbeiteten Hymenopteren aus der Familien der Crabronites, Sphegidae, Pompilidae und Heterogyna. Monatsberichte der Königlichen Preussischen Akademie der Wissenschaften zu Berlin, 1857, 509–513.
Petersen, B. (1988) The Palaearctic Mutillidae of I.C. Fabricius and some related material (Insecta, Hymenoptera, Aculeata). Steenstrupia, 14 (6), 129–224.
Radoszkowski, O. (1877) Chrysidiformes, Mutillidae et Sphegidae. Puteshestvie v Turkestan A.P. Fedtshenko [Voyage au Turkestan d‘Alexis Fedtschenko], Sankt-Petersburg, Series 14, 2 (5), 1–87, 3 pls. [in Russian]
Radoszkowski, O. (1885) Revision des armures copulatrices des mâles de la famille de Mutillides. Horae Societatis Entomologicae Rossicae, 19 (1–2), 3–49, 9 pls.
Schuster, R.M. (1949) Contribution towards a monograph of the Mutillidae of the Neotropical region. III. A key to the subfamilies represented and descriptions of several new genera (Hymenoptera). Entomologica Americana, New Series, 29 (3–4), 59–140.
Schmid-Egger, C. & Schmidt, S. (2021) Unexpected diversity in Central European Vespoidea (Hymenoptera, Mutillidae, Myrmosidae, Sapygidae, Scoliidae, Tiphiidae, Thynnidae, Vespidae), with description of two species of Smicromyrme Thomson, 1870. ZooKeys, 1062, 49–72. https://doi.org/10.3897/zookeys.1062.70763
Schwartz, F. (1984) Description d’un genre nouveau, disjoint de Smicromyrme, et d’une espèce nouvelle du sud de la France (Hymenoptera, Mutillidae). L’Entomologiste, 40, 21–25.
Sichel, J. & Radoszkowski, O. (1869–1870) Essai d’une monographie des Mutilles de l’ancien continent. Horae Societatis Entomologicae Rossicae, 6 (3 & 4), 139–172 (1869) & 173–309, tab. 6–11 (1870) .
Skorikov, A.S. (1935) Zur Mutilliden-Fauna Zentralasiens. Trudy Tadzhikskoi Basy Akademii Nauk SSSR, 5, 257–349, 7 pls. [in Russian]
Smith, F. (1855) Catalogue of Hymenopterous Insects in the Collection of the British Museum. Part III. Mutillidae and Pompilidae. Trustees of the British Museum, London, [4] + 206 pp., pl. A, pls. 1–5.
Smith, F. (1860 [“1861”]) Catalogue of hymenopterous insects collected by Mr. A.R. Wallace in the islands of Bachian, Kaisaa, Amboyna, Gilolo, and at Dory in New Guinea. Journal of the Proceedings of the Linnean Society, Zoology, London, 5, 93–143, pl. I. [published as supplement to Vol. 4] https://doi.org/10.1111/j.1096-3642.1860.tb01022.x
Smith, F. (1865) Descriptions of New Species of Hymenopterous Insects from the Islands of Sumatra, Sula, Gilolo, Salwatty, and New Guinea, collected by Mr. A.R. Wallace. Journal of the Proceedings of the Linnean Society, Zoology, 8, 61–94, pl. IV. https://doi.org/10.1111/j.1096-3642.1865.tb02422.x
Spinola, M. (1841) Hyménoptères recueillis à Cayenne en 1839 par M. Leprieur, pharmacien de la Marine Royale. Seconde Partie. Porte-Aiguillons. Annales de la Société Entomologique de France, 10, 85–149.
Suárez, F.J. (1965) Nota sinonímica preliminar y propuesta del nombre genérico Eremomyrme nom. nov. (Hymenoptera, Mutillidae). Memorie della Società Entomologica Italiana, 44, 51–52.
Suárez, F.J. (1975) Especies nuevas o poco conocidas de la fauna Mediterránea. IV. Tres nuevas Smicromyrme de la zona oriental (Hymenoptera, Mutillidae). Archivos Instituto de Aclimatación, Almeria, 20, 155–171.
Suárez, F.J. (1977) Dos nuevos Mutílidos Etiópicos y propuesta de un género nuevo para uno de ellos (Hymenoptera, Mutillidae). Nouvelle Revue d’Entomologie, 7 (2), 213–223.
Terine, J.B., Lelej, A.S. & Girish Kumar, P. (2021) Discovery of the genus Andreimyrme Lelej, 1995 (Hymenoptera: Mutillidae) in India and description of a new species from the Western Ghats. Zootaxa, 5020 (1), 184–190.
https://doi.org/10.11646/zootaxa.5020.1.10
Thaochan, N., Williams, K.A., Thoawan, K., Jeenthong, T. & Sittichaya, W. (2022) Three new species and one new country record of velvet ants (Hymenoptera, Mutillidae) from Thailand. Journal of Hymenoptera Research, 93, 151–165. https://doi.org/10.3897/jhr.93.94727
Tonnoir, A.L. (1930) Notes on Indian Blepharocerid larvae and pupae with remarks on the morphology of Blepharocerid larvae and pupae in general. Records of the Indian Museum, 32, 161–214. https://doi.org/10.26515/rzsi/v32/i2/1930/162531
Thomson, C.G. (1870) Öfversigt af Sveriges rofsteklar. Opuscula Entomologica, 1 (2), 202–251.
Tsuneki, K. (1993) On some Taiwanese Mutillidae, collected in 1976 by Mr. T. Murota, with descriptions of new taxa (Hymenoptera). Special Publications of the Japan Hymenopterists Association, 41, 39–50.
Turner, R.E. (1911) New Hymenoptera from Ceylon. Mutillidae and Scoliidae. Spolia Zeylanica, 7 (27), 141–154.
Waldren, G.C., Roberts, J.D. & Pitts, J.P. (2020) Phoretic copulation in the velvet ant Sphaeropthalma pensylvanica (Lepeletier) (Hymenoptera, Mutillidae): A novel behavior for Sphaeropthalminae with a synthesis of mating strategies in Mutillidae. Journal of Hymenoptera Research, 78, 69–89. https://doi.org/10.3897/jhr.78.55762
Waldren, G.C., Sadler, E.A., Murray, E.A., Bossert, S., Danforth, B.N. & Pitts, J.P. (2023) Phylogenomic inference of the higher classification of velvet ants (Hymenoptera: Mutillidae). Systematic Entomology. [in press]
Williams, K.A., Lelej, A.S., Okayasu, J., Borkent, C.J., Malee, R., Thoawan, K. & Thaochan, N. (2019) The female velvet ants (aka modkhong) of southern Thailand (Hymenoptera: Mutillidae), with a key to the genera of southeast Asia. Zootaxa, 4602 (1), 1–69.
https://doi.org/10.11646/zootaxa.4602.1.1
Zavattari, E. (1913) H. Sauter’s Formosa-Ausbeute. Mutillidae (Hym.). Archiv für Naturgeschichte, Series A,79 (3), 19–42. https://doi.org/10.5962/bhl.part.7709