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Type: Monograph
Published: 2023-03-08
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Review of Indo-West Pacific jawfishes (Opistognathus: Opistognathidae), with descriptions of 18 new species

Florida Museum of Natural History, University of Florida, Gainesville, FL 32611‒7800, USA.
taxonomy zoogeography endemism behavior Opistognathidae


Sixty species of jawfishes (Opistognathus) from the Indo-West Pacific are reported in an updated review, including descriptions of 18 new species: Opistognathus albomaculatus n.sp., O. asper n.sp., O. aurolineatus n.sp., O. bathyphilus n.sp., O. biporus n.sp., O. challenger n.sp., O. erdmanni n.sp., O. flavidus n.sp., O. helvolus n.sp., O. hyalinus n.sp., O. megalops n.sp., O. microspilus n.sp., O. nigripinnis n.sp., O. parvus n.sp., O. pholeter n.sp., O. triops n.sp., O. vigilax n.sp., and O. wassi n.sp.. Species accounts are provided for each species, including illustrations or color photographs, complete synonymies, specimens examined (or appropriate citation if previously published in detail), diagnosis, comparisons, etymology, and distribution maps. Geographic range extensions are reported for a number of species. An identification key is given for all species and frequency tables of important characters are also provided. The taxonomic status of Opistognathus inornatus and O. rosenbergii annulatus are discussed in detail but not completely resolved pending unavailable molecular data. Geographic variation is also described for Opistognathus adelus, O. albomaculatus n.sp., O. castelnaui, O. margaretae, O. variabilis, and O. vigilax n.sp. Many species are known only from holotypes and others from single localities, indicating how much more remains to be known about these jawfishes.


  1. Aizawa, M. (2002) Opistognathidae. In: Nakabo, T. (Ed.), Fishes of Japan with pictorial keys to the species. English Edition. Tokai University Press, Tokyo, pp.741–743.

  2. Al-Abdessalaam, T.Z S. (1995) Marine species of the Sultanate of Oman. Marine Science Fisheries Center, Ministry of Agriculture and Fisheries. Muscat Printing Press, Muscat, 412 pp.

  3. Allen, G.R. (1985) Fishes of Western Australia. In: Burgess, W.A. & Axelrod, H.R. (Eds.), Pacific Marine Fishes Book 9. T.F.H. Publications, Inc., Neptune City, New Jersey, pp. i–v +2205–2534.

  4. Allen, G.R. 2006. Coral Reef Fish Diversity. In: Green, A., Lokani, P., Atu, W., Ramohia, P., Thomas, P. & Almany, J. (Eds.), Solomon Islands Marine Assessment: Technical report of survey conducted May 13 to June 17, 2004. TNC Pacific Island Countries Report, 1/06, 112–156.

  5. Allen, G.R. (2008) Conservation hotspots of biodiversity and endemism for Indo-Pacific coral reef fishes. Aquatic Conservation: Marine and Freshwater Ecosystems, 18, 541–556.

  6. Allen, G.R. & Adrim, M. (2003) Coral reef fishes of Indonesia. Zoological Studies, 42 (1), 1–72.

  7. Allen, G.R. & Erdmann, M.V. (2012) Reef fishes of the East Indies. Vol. 1. Tropical Reef Research, Perth, 424 pp.

  8. Allen, G.R & Smith-Vaniz, W.F. (1994) Fishes of the Cocos (keeling) Islands. Atoll Research Bulletin, 412, 1–21.

  9. Allen, G.R., Steene, R., Humann, P. & DeLoach, N. (2003) Reef fish identification tropical Pacific. New World Publications, Jacksonville, Florida, 457 pp.

  10. Allen, G.R. & Swainston, R. (1988) The Marine Fishes of North-Western Australia A Field Guide for Anglers and Divers. Western Australian Museum, Perth, 201 pp.

  11. Allen, R. 2006. Gorontalo hidden paradise. Snow Publishing, Singapore, 156 pp.

  12. Alleyne, H.G. & Macleay, W. (1877) The ichthyology of the Chevert Expedition. Proceedings of the Linnean Society of New South Wales, 1 (3), 261–281.

  13. Anderson, R. C. (2000) An underwater guide to Indonesia. University of Hawaii Press, Honolulu, 160 pp.

  14. Araga, C. & Tanase, H. (1966) Fish stranding caused by a typhoon in the vicinity of Seto. Publications of the Seto Marine Biological Laboratory, 14 (2), 155–160.

  15. Arai, R. & Abe, T. (1970) Fishes of the Tsushima Islands. Memoir of the National Science Museum, 3, 83–100. [in Japanese]

  16. Baker, J.L. (2013) Status report on rare and endemic species and other marine fauna of conservation concern in the northern rivers CMA region, New South Wales. Part 2. Bony Fishes. Report for Northern Rivers Catchment Management Authority, New South Wales, 232 pp.

  17. Barnard, K.H. (1927) A monograph of the marine fishes of South Africa. Part 2. Annals of the South African Museum, 21, 419–1033.

  18. Berg, L.S. (1940) Classification of fishes, both recent and fossil. Travaux de 1’Institut Zoologique 1’Academie des sciences de l’URSS, 5 (2), 7–517. [in Russian and English]

  19. Betancur-R, R., Wiley, E.O., Arratia, G., Acero, A., Bailly, N., Miya, M., Lecointre, G. & Orti, G. (2017) Phylogenetic classification of bony fishes. BMC Evolutionary Biology, 17, 162.

  20. Biswas, S., Mishra, S.S., Das, N.P.I., Satpathy, K.K., Nayak, L. & Selvanayagam, M. (2013) First record and range-extension of bigscale jawfish, Opistognathus macrolepis (Perciformes: Opistognathidae), from India. Marine Biodiversity Records, 6 (e8), 1–4.

  21. Bleeker, P. (1851) Nieuwe bijdrage tot de kennis der ichthyologische fauna van Celebes. Natuurkundige Tijdschrift voor Nederlandsch-Indië, 2 (2), 209–224.

  22. Bleeker, P. (1853) Bijdrage tot de kennis der ichthyologische fauna van Solor. Natuurkundige Tijdschrift voor Nederlandsch-Indië, 5, 67–96.

  23. Bleeker, P. (1856) Bijdrag tot de kennis der ichthyologische fauna van het eiland Nias. Natuurkundige Tijdschrift voor Nederlandsch-Indië, 12, 211–288.

  24. Bleeker, P. (1859) Visschen van Singapore, verzameld door Graaf Fr. De Castelnau. Natuurkundige Tijdschrift voor Nederlandsch-Indië, 20 (2), 236–239.

  25. Bleeker, P. (1860) Dertiende bijdrage tot de Kennis der vischfauna van Celebes (visschen van Bonthain, Badjoa, Sindjai, Lagoesi en Pompenoea). Acta Societatis Regiae Scientiarum Indo-Neêrlandicae, 8 (7), 1–60.

  26. Bleeker, P. (1868) Description de trois especes inédites de poissons des iles D’Amboine et de Waigiou. Verslagen en Mededeelingen der Koninklijke Akademie van Wetenschappen, 2 (2), 331–335.

  27. Bleeker, P. (1874) Les especes insulindiennes de la famille des Opisthognathoides. Archives Neerlandaises des Sciences exactes et Naturelles, 9, 466–476.

  28. Boeseman, M. (1954) On the rediscovery of the type specimen of Opisthognathus papuensis Bleeker, 1868. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen, Series C, 57 (3), 271–273.

  29. Bogorodsky, S.V. & Randall, J.E. (2019) Endemic fishes of the Red Sea. In: Rasul, N.M.A. & Stewart, I.C.F. (Eds.), Oceanographic and Biological Aspects of the Red Sea. Springer, Cham, pp. 239–265.

  30. Böhlke, J.E. (1953) A catalogue of the type specimens of recent fishes in the natural history museum of Stanford University. Stanford Ichthyological Bulletin, 5, 1–168.

  31. Böhlke, J.E. (1960) Comments on fishes with disjunct lateral lines, with the description of a new one from the Bahamas. Notulae naturae, Academy of Natural Sciences of Philadelphia, 330, 1–11.

  32. Boulenger, G.A. (1888) An account of the fishes obtained by Surgeon-Major A.S.G. Jayakar at Muscat, east coast of Arabia. Proceedings of the Zoological Society of London, 1887 (4), 653–667.

  33. Briggs, J.C. (1961) Emendated generic names in Berg’s classification of fishes. Copeia, 1961 (2), 161–166.

  34. Briggs, J.C. (1987) Antitropical distributions and evolution in the Indo-West Pacific Ocean. Systematic Zoology, 36, 237–247.

  35. Bruton, M. (2018) The Fishy Smiths: A Biography of J.L.B. & Margaret Smith. Penguin Books, Cape Town, 344 pp.

  36. Carpenter, K.E. Harrison, P.L., Hodgon, G., Alsaffar, A.H. and Alhazeem, H. (1997) The corals and coral reef fishes of Kuwait. Kuwait Institute for Scientific Research, Fahad Al-Marzouk Printing Establishment, Kuwait City, 165 pp.

  37. Chan, W.L. (1966) Notes on opisthognathid jawfishes from Hong Kong, with description of a new species. Japanese Journal of Ichthyology, 14 (1/3), 9–11.

  38. Chan, W.L. (1968) Opistognathus hongkongiensis, a replacement name for the jawfish Opisthognathus fasciatus Chan. Copeia, 1968 (1), 198.

  39. Chen, J.-P., Jan, R.-Q. & Shao, K.-T. (1997) Checklist of reef fishes from Taiping Island (Itu Aba Island), Spratly Islands, South China Sea. Pacific Science, 51 (2), 143–166.

  40. Chen, J.-P., Shao, K.-T., Jan, R.-Q., Kuo, J.-W. & Chen, J.-Y. (2010) Marine fishes of Kenting National Park. 1st Revised Edition. Kenting National Park Headquarters, Kenting, 650 pp. [in Chinese]

  41. Choat, J.H., van Herwerden, L., Robbins, W.D., Hobbs, J.P. & Ayling, A.M. (2006) A report on the ecological surveys undertaken at Middleton and Elizabeth Reefs, February 2006. Report by James Cook University to the Department of the Environment and Heritage, Townsville, 65 pp.

  42. Choi, Y., Kim, J.-H. & Park, J.-Y. (2003) Marine fishes of Korea. Kyuo-Hak, Seoul, 747 pp.

  43. Colin, P.L. (1972) Daily activity patterns and effects of environmental conditions on the behavior of the yellowhead jawfish, Opistognathus aurifrons, with notes on its ecology. Zoologica, 57, 137–169.

  44. Colin, P.L. (1973) Burrowing behavior of the Yellowhead Jawfish, Opistognathus aurifrons. Copeia, 1973 (1), 84–90.

  45. Conlu, P.V. (1986) Fishes. Guide to Philippine Flora and Fauna 9. Natural Resources Management Center, Ministry of Natural Resources and University of the Philippines, JMC Press, Inc., Quezon City, 495 pp.

  46. Costello, M.J. (2020) Taxonomy as the key to life. Megataxa, 1 (2), 105–113.

  47. Cuvier, G. (1816) Le regne animal. Vol. 2. 1st Edition. s.n., Paris, 532 pp.

  48. Cuvier, G. & Valenciennes, A. (1836) Histoire naturalle des poissons. Vol. 11. F.G. Levrault, Paris, 506 pp.

  49. Datovo, A. & Vari, R.P. (2013) The jaw adductor muscle complex in Teleostean fishes: Evolution, homologies and revised nomenclature (Osteichthyes: Actinopterygii). PLoS ONE, 8 (4), e60846.

  50. Day, F. (1876) The fishes of India; being a natural history of the fishes known to inhabit the seas and fresh waters of India, Burma, and Ceylon, London, 2, 169–368.

  51. DeLoach, N. (1999) Reef Fish Behavior–Florida Caribbean Bahamas/ with photographer Paul Human. New World Publications, Inc., Jacksonville, Florida, 359 pp.

  52. Dor, M. (1970) Nouveaux poissons pour la faune de la Mer Rouge. Contributions to the knowledge of the Red Sea 44. Sea Fisheries Research Station, State of Israel Bulletin, 54, 7–28.

  53. Dor, M. (1984) CLOFRES: Checklist of the fishes of the Red Sea. Israel Academy of Science and Humanities, Jerusalem, 427 pp.

  54. Eagderi, S., Fricke, R., Esmaeili, H.R. & Jlili, P. (2019) Annotated checklist of the fishes of the Persian Gulf: diversity and conservation status. Iranian Journal of Ichthyology, 6 (Supplement 1), 1–171.

  55. Eibl-Eibesfeldt, I. von & Klausewitz, W. (1961) Gnathypops rosenbergi annulata n. ssp. Von den Nikobaren (Pisces, Percomorphi, Opisthognathidae). Senckenbergiana Biologica, 42 (5/6), 421–426.

  56. Fourmanoir, P. (1965) Liste complementaire des poissons marins de Nha-Trang. Cashiers O.R.S.T.O.M. (Oceanographie), No. Special, 1–114.

  57. Fowler, H.W. (1925) Fishes from Natal, Zululand, and Portuguese East Africa. Proceedings of the Academy of Natural Sciences of Philadelphia, 77, 187–268.

  58. Fowler, H.W. (1934) Fishes obtained by Mr. H. W. Bell-Marley chiefly in Natal and Zululand in 1929 to 1932. Proceedings of the Academy of Natural Sciences of Philadelphia, 86, 405–514.

  59. Fowler, H.W. (1946) Notes on Bahama fishes with a description of a new jawfish (Opisthognathidae). Notulae Naturae, Philadelphia, 181, 1–8.

  60. Fowler, H.W. & Silvester, C.F. (1922) A collection of fishes from Samoa. Carnegie Institution of Washington, 18 (312), 109–126.

  61. Fricke, R., Allen, G.R., Andréfouët, S., Chen, W-J., Hamel, M.A., Laboute, P., Mana, R., Hui, T.H. & Uyeno, D. (2014) Checklist of the marine and estuarine fishes of Madang District, Papua New Guinea, western Pacific Ocean, with 820 new records. Zootaxa monograph, 3832 (1), 1–247.

  62. Fricke, R. & Eschmeyer, W.N. (2022) Eschmeyer’s catalog of fishes: Guide to fish collections. Electronic version. Available from: (accessed 7 February 2022)

  63. Fricke, R., Eschmeyer, W.N. & van der Laan, R. (Eds.) (2022) Eschmeyer’s catalog of fishes: gender, species references. Electronic version. Available from: (accessed 7 February 2022)

  64. Fricke, R., Kulbicki, M. & Wantiez, L. (2011) Checklist of the fishes of New Caledonia, and their distribution in the southwest Pacific Ocean. Stuttgarter Beiträge zur Naturkunde A, Neue Serie 4, 341‒463.

  65. Fricke, R., Mahafina, J., Behivoke, F., Jaonalison, H., Léopold, M. & Ponton, D. (2018) Annotated checklist of the fishes of Madagascar, southwestern Indian Ocean, with 158 new records. FishTaxa, 3 (1), 1–432.

  66. Gaither, M.R., Bowen, B.W., Bordenave, T-R., Rocha, L.A., Newman, S.J., Gomez, J.A., Herwerden, L. van, and Graig, M.T. (2011) Phylogeography of the reef fish Cephalopholis argus (Epinephelidae) indicates Pleistocene isolation across the Indo-Pacific barrier with contemporary overlap in the coral triangle. BMC Evolutionary Biology, 11, 189. []

  67. Gaither, M.R. & Rocha, L.A. (2013) Origins of species richness in the Indo-Malay-Philippine biodiversity hotspot: evidence for the centre of overlap hypothesis. Journal of Biogeography, 40, 1638–1648.

  68. Gill, A.C. & Mooi, R.D. (1993) Monophyly of the Grammatidae and of the Notograptidae, with evidence for their phylogenetic positions among perciforms. Bulletin of Marine Science, 52 (1), 327–350.

  69. Gill, A.C., Tea, Y.-K. & Senou, H. (2016) Plectranthias takasei, new species of anthiadine fish from southern Japan (Teleostei: Serranidae). Zootaxa, 4205 (4), 349–356.

  70. Gill, T.N. (1862) Remarks on the relations of the genera and other groups of Cuban fishes. Proceedings of the Academy of Natural Sciences of Philadelphia, 14, 235–242.

  71. Gloerfelt-Tarp, T. & Kailola, P.J. (1985) Trawled fishes of southern Indonesia and northwestern Australia. Australian Development Assistance Bureau, Sydney, 406 pp.

  72. Golani, D. & Bogorodsky, S.V. (2010) The fishes of the Red Sea–Reappraisal and updated checklist. Zootaxa monograph, 2463 (1), 1–135.

  73. Golani, D. & Fricke, R. (2018) Checklist of the Red Sea fishes with delineation of the Gulf of Suez, Gulf of Aqaba, endemism and Lessepsian migrants. Zootaxa, 4509 (1), 1–215.


  75. Gon, O. (1996) Fifty years of marine fish systematics at the J.L.B. Smith Institute of Ichthyology. Transactions of the Royal Society of South Africa, 51, 45–78.

  76. Gosline, W.A. (1968) The suborders of perciforms fishes. Proceedings of the United States National Museum, 124 (3647), 1–78.

  77. Grant, E.M. (1982) Guide to Fishes, 5th edition. Department of Harbours and Marine, Queensland, 896 pp.

  78. Greenfield, D.W. (2001) John E. Randall. Copeia, 2001 (3), 872–877.[0872:JER]2.0.CO;2

  79. Günther, A. (1860) Catalogue of the fishes in the British Museum. Catalogue of the acanthopterygian fishes in the collection of the British Museum. Vol. 2. Squamipinnes, Cirrhitidae, Triglidae, Trachinidae, Sciaenidae, Polynemidae, Sphyraenidae, Trichiuridae, Scombridae, Carangidae, Xiphiidae. British Museum, London, xxi + 548 pp.

  80. Günther, A. (1880) Report on the shore fishes procured during the voyage of H.M.S. Challenger in the years 1873–1876. Report on the scientific results of the exploring voyage of H.M.S. Challenger 1873–76, Zoology, 1 (6), 1–82.

  81. Gushiken, S. (1972) Fishes of the Okinawa Islands, with 330 species colored. Tiger Print Company, Okinawa, 251 pp. [in Japanese]

  82. Hall, R. (2002) Cenozoic geological and plate tectonic evolution of SE Asia and the SW Pacific: computer-based reconstructions, model and animations. Journal of Asian Earth Science, 20, 353–431.

  83. Hayashi, M. & Okuri, S. (2007) Review of the genus Opistognathus (Perciformes: Opistognathidae) from Japan. Science Report Yokosuka City Museum, 54, 27–57. [in Japanese with English summary]

  84. Heemstra, P.C. & Bruton, M.N. (1989) Obituary: Margaret Mary Smith 1916–1987. Copeia, 1988 (2), 523–524.

  85. Herald, E.S. (1972) Fishes of North America. Doubleday & Company, Inc., Garden City, New York, 254 pp.

  86. Herre, A.W.C.T. (1933) Twelve new Philippine fishes. Copeia, 1933 (1), 17–25.

  87. Herre, A.W.C.T. (1934) Notes on fishes in the Zoological Museum of Stanford University. 1. The fishes of the Herre Philippine expedition of 1931. The fishes of the Herre 1931 Philippine expedition with descriptions of 17 new species. Newspaper Enterprise Limited, Hong Kong, Herre Philippine Expedition, 1934, 1–106.

  88. Herre, A.W.C.T. (1953) Check list of Philippine fishes. United States Fish and Wildlife Service Research Report, No. 20, 1–977.

  89. Hess, H.C. (1993) Male mouth brooding in jawfishes (Opistognathidae): constraints on polygyny. Bulletin of Marine Science, 52 (2), 806–818.

  90. Hibino, Y., Sasaki, D. & Kimura, S. (2016) A new record of a jawfish (Perciformes: Opistognathidae) Opistognathus variabilis from Yonaguni Island, Yaeyama Islands, Japan. Japanese Journal of Ichthyology, 63 (1), 33‒38 [in Japanese]

  91. Hiramatsu, W. & Endo, H. (2013) Opistognathus trimaculatus, a new species of jawfish (Teleostei: Opistognathidae) from Tosa Bay, southern Japan. Bulletin of National Museum of Natural History, series A, supplement, 7, 19–23.

  92. Ho, H.-C. & Shao, K-T. (2011) Annotated checklist and type catalog of fish genera and species described from Taiwan. Zootaxa, 2957 (1), 1–74.

  93. Hobbs, J-P.A. & Feary, D.A. (2007) Monitoring the ecological status of Elizabeth and Middleton Reefs. Report to the Department of the Environment and Water Resources, s.n., 37 pp.

  94. Hoeksema, B.W. (2007) Delineation of the Indo-Malayan centre of maximum marine biodiversity: the Coral Triangle. In: Renema, W. (Ed.), Biogeography, time, and place: distributions, barriers, and islands. Springer, Cham, pp. 117178.


  96. Hoese, D.F. & Bray, D.J. (2006) Opistognathidae. In: Zoological Catalogue of Australia. 35 (1). Fishes. CSIRO Publishing, Collingwood, pp. 1073–1077.

  97. Honma, Y. (1957) Further additions to “a list of the fishes collected in the Province of Echigo, including Sado Island.” Japanese Journal of Ichthyology, 6 (4–6), 109112. [in Japanese]

  98. Hughes, L.C., Orti, G., Huang, Y., Sun, Y., Baldwin, C.C., Thompson, A.W., Arcila, D., Betanchur-R, R., Li, C., Becker, L., Bellora, N., Zhao, X., Li, X., Wang, M., Fang, C., Xie, B., Zhou, Z., Huang, H., Chen, S. & Venkatesh, B. (2018) Comprehensive phylogeny of ray-finned fishes (Actinopterygii) based on transcriptomic and genomic data. Proceedings of the National Academy of Sciences, 115 (24), 62496254.

  99. Hussain, S. & Jawad, L.A. (2014) First records of Opistognathus muscatensis Boulenger, 1888 (Opistognathidae), Trachinotus baillonii (Lacepède, 1801) (Carangidae) and Atrobucca nibe (Jordan & Thompson, 1911) (Sciaenidae) off the Iraq Coast Iraq, Arabian Gulf. International Journal of Marine Science, 4 (28), 253–258.

  100. Hutchins, J.B. (1974) Fish attacks man. Western Australian Naturalist, 12, 182183.

  101. Hutchins, J.B. (2001) Checklist of the fishes of Western Australia. Records of the Western Australian, Supplement 63, 9–50.

  102. Hutchins, J.B. & Smith, K.N. (1991) A catalogue of type specimens of fishes in the Western Australian Museum. Records of the Western Australian Museum Supplement No. 38, 156.

  103. Hylleberg, J. & Aungtonya, C. (2013) Biodiversity survey in Chalong Bay and surrounding areas, east coast of Phuket, Thailand. Phuket Marine Biological Center Special Publication, 32, 81112.

  104. Ikeda, H. & Nakabo, T. (2015) Fishes of the Pacific coasts of southern Japan. Tokai University Press, Hadano, 597 pp. [in Japanese]

  105. Jaafar, T.N.A.M., Lee, J.N., Mazlan, A.G., Kadir, S.T.S.A. & Seah, Y.G. (2019) Opistognathus nigromarginatus Rüppell, 1830 (Perciformes, Opistognathidae), Bridled jawfish: a first record from Malaysia. Check List, 15 (5), 883–886.

  106. Jawad, L.A., Faddagh Ziyadi, M.S., Näslund, J., Pohl, T. & Al-Mukhtar, M.A. (2018) Checklist of the fishes of the newly discovered coral reef in Iraq, north-west Arabian Gulf, with 10 new records to the Arabian Gulf. Aqua, Journal of Ichthyology and Aquatic Biology, 24 (3), 89–138.

  107. Johnson, J.W. (1999) Annotated checklist of the fishes of Moreton Bay, Queensland, Australia. Memoirs of the Queensland Museum, 43 (2), 709‒762.

  108. Jordan, D.S. & Richardson, R.E. (1908) Fishes from islands of the Philippine Archipelago. Bulletin of the Bureau of Fisheries, 27, 233–287. [for 1907]

  109. Jordan, D.S. & Snyder, J.O. (1902) A review of the trachinoid fishes and their supposed allies found in the waters of Japan. Proceedings of the United States National Museum, 24 (1263), 461–497.

  110. Jordan, D.S. & Thompson, W.F. (1913) Notes on a collection of fishes from the island of Shikoku in Japan, with a description of a new species, Gnathypops iyonis. Proceedings of the United States National Museum, 46 (2011), 65–72.

  111. Kailola, P.J. (1973) Additions to the fish fauna of Papua New Guinea—I. Papua New Guinea Agricultural Journal, 24 (1), 1–15.

  112. Kailola, P.J. (1975) A catalogue of the fish reference collection at the Kanudi Fisheries Research Laboratory, Port Moresby. Department of Agriculture, Stock and Fisheries, Port Moresby, Research Bulletin, 16, 1–277.

  113. Kailola, P.J. (1987) The fishes of Papua New Guinea: A revised and annotated checklist. Volume two: Scorpaenidae to Callionymidae. Department of Fisheries and Marine Resources, Port Moresby, Papua New Guinea, Research Bulletin, 41, xxii + 195–418.

  114. Kamohara, T. (1952) Revised descriptions of the off-shore bottom-fishes of Prov. Tosa, Shikoku, Japan. Research Reports of the Kochi University, Natural Science, 3, 1–122.

  115. Kamohara, T. (1956) On some rare species of fishes from Prov. Tosa, Japan. Reports of the Usa Marine Biological Station, 2 (2), 1–4.

  116. Katayama, M. & Fujioka, Y. (1958) Fishes of Ooshima-gun, Yamaguti Prefecture. Bulletin Faculty of Agriculture, 9, 1149–1168. [in Japanese]

  117. Kimura, S., Imamura, H., Nguyen, V.Q. & Pham, T.D. (Eds.) (2018) Fishes of Ha Long Bay, the natural world heritage site of northern Vietnam. Fisheries Research Laboratory, Mie University, Shima, ix + 314 pp.

  118. Kimura, S. & Matsuura, K. (2003) Fishes of Bitung northern tip of Sulawesi, Indonesia. Ocean Research Institute, University of Tokyo, 244 pp.

  119. Kimura, S., Satapoomin, U. & Matsuura, K., eds. (2009) Fishes of Andaman Sea, west coast of southern Thailand. National Museum of Nature and Science, Tokyo, 346 pp.

  120. Kimura, S. & Suzuki, K. (1982) Fish fauna of Ago Bay and its adjacent waters, Mie Prefecture, Japan. Supplement I. Report of the Fisheries Research Laboratory, Mie University, 3, 1–20.

  121. Klunzinger, C.B. (1871) Synopsis der Fische des Rothen Meeres. II. Theil. Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien, 21, 441–688.

  122. Krishnan, S. & Mishra, S.S. (1993) On a collection of fish from Kakinada–Gopalpur sector of the east coast of India. Records of the Zoological Survey of India, 93 (1–2), 201–240.

  123. Kulbicki, M., Randall, J.E. & Rivaton, J. (1994) Checklist of the fishes of the Chester Islands (Coral Sea). Micronesia, 27 (1–2), 1–43.

  124. Kulbicki, M. & Williams, J.T. (1997) Checklist of Shorefishes of Ouvéa Atoll, New Caledonia. Atoll Research Bulletin, 444, 1–26.

  125. Kuiter, R.H. (1992) Tropical reef fishes of western Pacific Indonesia and adjacent waters. Gramedia Pustaka Utama, Jakarta, 314 pp.

  126. Kuiter, R.H. (1993) Coastal fishes of south-eastern Australia. University of Hawaii Press, Honolulu, 437 pp.

  127. Kuiter, R.H. & Debelius, H. (1994) Southeast Asia tropical fish guide. IKAN-Unterwasseraarchiv, Frankfurt, 321 pp.

  128. Kuiter, R.H. & Debelius, H. (2006) World Atlas of Marine Fishes. IKAN-Unterwasseraarchiv, Frankfurt, 720 pp.

  129. Kuiter, R.H. & Tonozuka, T. (2004) s.n. In: Pictorial guide to Indonesian reef fishes. Vol. 3. 2nd Edition. PT Dive & Dive’s, Denpasar, Bali, pp. 623–893.

  130. Kyushin, K., Amaoka, K., Nakaya, K., Ida, H., Tanino, Y. & Senta, T. (1982) Fishes of the South China Sea. Japan Marine Fishery Resource Research Center, Tokyo, 333 pp.

  131. Laboute, P. & Grandperrin, R. (2000). Poissons de Nouvelle-Calédonie. Editions Catherine Ledru, Nouméa, New Caledonia, 520 pp.

  132. Larson, H.K. (1999) Report to parks Australia north, on the estuarine fish inventory of Kakadu National Park, Northern Territory, Australia. Magnt Research Report, 5, 3–50.

  133. Larson, H.K. & Williams, R.S. (1997) Darwin Harbour fishes: a survey and annotated checklist. In: Hanley, J.R., Caswell, G., Megirian, D. & Larson, H.K. (Eds.), Proceedings of the Sixth International Marine Biological Workshop. The marine flora and fauna of Darwin Harbour, Northern Territory, Australia. Museums and Art Galleries of the Northern Territory and the Australian Marine Sciences Association: Darwin, pp. 339–380.

  134. Larson, H.K., Williams, R.S. & Hammer, M.P. (2013) An annotated checklist of the fishes of the Northern Territory, Australia. Zootaxa Monograph, 3696 (1), 1–293.

  135. Leis, J.M. & Carson-Ewart, B.M. (Eds.) (2000) The larvae of Indo-Pacific coral reef fishes. An identification guide to marine larvae. Fauna Malesiana Handbook, 2, 1‒850.

  136. Leis, J.M. & Trnski, T. (1989) The larvae of Indo-Pacific Shorefishes. University of Hawaii Press, Honolulu, 371 pp.

  137. Linklater, E. (1972) The Voyage of the Challenger. John Murray (publishers), Ltd, London, 288 pp.

  138. Longley, W.H. (1922) Habits and local distributions of Tortugas fishes. Yearbook, Carnegie Institution of Washington, 20 (1921), 204–205.

  139. Longley, W.H. (1927) Observations upon the ecology of Tortugas fishes with notes upon the taxonomy of species new or little known. (Definition of three new genera and two species). Carnegie Institution of Washington Yearbook, 26, 222–224.

  140. Longley, W.H. & Hildebrand, S.F. (1940) New genera and species of fishes from Tortugas, Florida. Papers Tortugas Laboratory, Carnegie Institution of Washington, 32, 223‒285.

  141. Macleay, W. (1878) The fishes of Port Darwin. Proceedings of the Linnean Society of New South Wales, 2, 344–367.

  142. Macleay, W. (1881) Descriptive catalogue of the fishes of Australia Part II. Proceedings of the Linnean Society of New South Wales, 5 (4), 510–629.

  143. Mariasingarayan, Y., Karuppiah, K., Danaraj, J., Sekar, S. & Ayyappan, S. (2021) Opistognathus evermanni (Jordan & Snyder, 1902), the first distributional record from Indian coastal waters. Journal of Applied Ichthyology, 37, 7861–789.

  144. Marshall, T.C. (1964) Fishes of the Great Barrier Reef and Coastal Waters of Queensland. Angus and Robertson, Sydney, 566 pp.

  145. Masuda, H., Amaoka, K., Araga, C., Uyeno, T. & Yoshino, T. (Eds.) (1984) The fishes of the Japanese Archipelago. Tokai University Press, Tokyo, 438 pp.

  146. Masuda, H., Araga, C. & Yoshino, T. (1975) Coastal fishes of southern Japan. Tokai University Press, Tokyo, 438 pp.

  147. McCulloch, A.R. (1914) Notes on some Western Australian fishes. Records of the Western Australian Museum, 1 (3), 211–227.

  148. McCulloch, A.R. (1929) A checklist of the fishes recorded from Australia. Part III. Memoirs of the Australian Museum, Sydney, 5 (3), 329–436.

  149. McKay, R.J. (1969) The genus Tandya in Western Australia, with a description of a new opistognathid fish, Tandya reticulata sp. nov. Journal of the Royal Society of Western Australia, 52 (1), 1–2.

  150. McKay, R.J. (1970) Additions of the fish fauna of Western Australia 5, Fisheries Bulletin of Western Australia, 9 (5), 3–24.

  151. Meek, S.E. & Hildebrand, S.F. (1928) The marine fishes of Panama. Part III. Field Museum of Natural History, Zoological Series, 15 (publication 249), i–xxxi +709–1045, pls. 72–102.

  152. Mees, G.F. (1959) Additions to the fish fauna of western Australia–I. Fisheries Bulletin, 9 (1), 5–11.

  153. Mok, H.-K., Chang, H.-J. & Lee, C.-Y. (1990) Phylogenetic interrelationships of the perciform Acanthoclinidae, Grammidae, Plesiopidae, Pseudochromidae and Opistognathidae. Bulletin of the Institute of Zoology, Academia Sinica, 29 (1), 29–39.

  154. Mooi, R.D. (1990) Egg surface morphology of pseudochromoids (Perciformes: Percoidei), with comments on its phylogenetic implications. Copeia, 1990 (2), 455–475.

  155. Mooi, R.D. (1993) Phylogeny of the Plesiopidae (Pisces: Perciformes) with evidence for the inclusion of the Acanthoclinidae. Bulletin of Marine Science, 52 (1), 284–326.

  156. Mooi, R.D. & Gill, A.C. (1995) Association of epaxial musculature with dorsal-fin pterygiophores in acanthomorph fishes, and its phylogenetic significance. Bulletin of Natural History Museum of London, Zoology, 61 (2), 121–137.

  157. Moore, G.I., Hutchins, J.B., Smith, K.N. & Morrison, S.M. (2008) Catalogue of type specimens of fishes in the Western Australian Museum (second edition). Records of the western Australian Museum Supplement, 74, 1–69.

  158. Moore, G.I., Morrison, S.M., Hutchins, J.B., Allen, G.R. & Sampey, A. (2004) Kimberley marine biota. Historical data: fishes. Records of the Western Australian Museum Supplement, 84, 161–206.

  159. Motomura, H., Alama, U.B., Muto, N., Babaran, R.P. & Ishikawa, S. (Eds.) (2017) Commercial and bycatch market fishes of Panay Island, Republic of the Philippines. The Kagoshima University Museum, University of the Philippines Visayas, Iloilo, and Research Institute of Humanity and Nature, Kyoto, 246 pp., 911 figs.

  160. Motomura, H. & Harazaki, S. (2017) Annotated checklist of marine and freshwater fishes of Yaku-shima Island in the Osumi Island, Kagoshima, southern Japan, with 129 new records. Bulletin of the Kagoshima University Museum, 9, 1–183.

  161. Motomura, H., Kimura, S., Gait, Y.G., Sheikh Abdul Kadir, S.T. & Ghaffar, M.A. (Eds.) (2021) Reef and shore fishes of Bidong Island, off east coast of Malay Peninsula. The Kagoshima University Museum, Kagoshima, 80 pp.

  162. Motomura, H. & Satapoomin, U. (2009) Opistognathidae. In: Kimura, S., Satapoomin, U, and Matsuura, K. (Eds.), Fishes of Andaman Sea west coast of southern Thailand. National Museum of Nature and Science, Tokyo, pp. i–vi + 1–346.

  163. Munro, I.S.R. (1958) The fishes of the New Guinea Region. A check-list of New Guinea incorporating new records of species collected by the fisheries survey vessel Fairwind during the years 1948 to 1950. The Papua and New Guinea Agricultural Journal, 10 (4), 97–369. [1956]

  164. Munro, I.S.R. (1967) The fishes of New Guinea. Department of Agriculture, Stock and Fisheries, Port Moresby, New Guinea, xxxvii + 651 pp., 78 pls.

  165. Myers, R.F. (1999) Micronesian Reef Fishes. 3rd Edition. Coral Graphics, Guam, 330 pp., 192 color pls.

  166. Myoung, J.-G., Cho, S.-H., Kim, J.M. & Kim, Y.U. (1999) First record of the jawfish, Opistognathus iyonis (Opistognathidae, Perciformes) from Korea. Korean Journal of Ichthyology, 1 (2), 139–142.

  167. Nakabo, T. (Ed.) (2000) Fishes of Japan with pictorial keys to the species. 2nd Edition. Tokai University Press, Tokyo, lvi + 866 pp. [English edition]

  168. Nakae, M., Motomura, H., Hagiwara, K., Senou, H., Koeda, K., Yoshida, T., Tashiro, S., Jeong, B., Hata, H., Fukuki, Fujiwara, K., Yamakawa, T., Aizawa, M., Shinohara, K. & Matsuura, K. (2018) An annotated checklist of fishes of Amami-oshima Island, the Ryukyu Islands, Japan. Memoirs of the National Museum of Natural Science, Tokyo, 52, 205–361.

  169. Nakayama, N., Hiramatsu, W., Kai, Y. & Endo, H. (2016) First record of the jawfish Opistognathus trimaculatus Hiramatsu and Endo 2013 (Actinopterygii: Perciformes: Opistognathidae) from the East China Sea, with comments on its diagnosis. Marine Biology, 1–4. [published online]

  170. Norman, J.R. (1939) Fishes. The John Murray Expedition 1933-34, Scientific Reports, 7 (1), 1–116.

  171. Ochiai, A. & Asano, H. (1964) Two rare fishes, Gnathypops hopkinsi Jordan & Snyder, and Chauliodus sloani sloani Bloch & Schneider, obtained from Japan. Bulletin of the Misaki Marine Biological Institute, Kyoto University, 4, 75–81. [1963]

  172. Ogilby, J.D. (1908) On new genera and species of fishes. Proceedings of the Royal Society of Queensland 21, 1–26.

  173. Ogilby, J.D. (1920) Edible fishes of Queensland. Part XVI. Opisthognathidae (No. 1). Memoirs of the Queensland Museum, 7 (1), 21–30.

  174. Okamura, O. & Amaoka, K. (1997) Sea fishes of Japan. Yama-Kei Publishers Co., Ltd., Tokyo, 783 pp. [in Japanese]

  175. Okuri, S., Iwashita, M. & Hagiwara, K. (2004) Opistognathus iyonis (Jordan et Thompson, 1913). I.O.P. Diving News, 15 (6), 1. [in Japanese]

  176. Park, J-H., Kim, J.K., Choi, J.H. & Choi, Y.M. (2008) First record of a jawfish, Opistognathus hongkongiensis (Opistognathidae: Perciformes) from Korea. Korean Journal of Ichthyology, 20 (1), 74–77.

  177. Peters, W. (1866) Mittheilung über Fische (Protopterus, Auliscops, Labrax, Labracoglossa, Nematocentris, Serranus, Scorpis, Opisthognathus, Scombresox, Acharnes, Anguilla, Gymnomuraena, Chilorhinus, Ophichthys, Helmichthys). Monatsbericht der Königlich Preussischen Akademie der Wissenschaften zu Berlin, 1866, 509–526.

  178. Playfair, R.L. & Günther, A. (1867) The Fishes of Zanzibar, with a list of the fishes of the whole east coast of Africa. s.n., London, xiv + 153 pp., 21 pls. [reprinted in 1971, with a new introduction by G. S. Myers and a new forward by A. E. Gunther; Newton K. Gregg, publisher, Kentfield, California]

  179. Popta, C.M.L. (1922) Vierte und letzte fortsetzung der Beschreibung von neuen Fischarten der Sunda-Expedition. Zoologische Mededeelingen, Leiden, 7, 27–39.

  180. Rainboth, W.J., Vidthayanan, C. & Dinhyen, M. (2012) Fishes of the greater Mekong ecosystem with species list and photographic atlas. Miscellaneous Publications, Museum of Zoology, University of Michigan, No. 21, 1–172, 121 pls.

  181. Ramsay, E.P. (1883) Descriptions of some new Australian fishes. Proceedings of the Linnean Society of New South Wales, 8 (1), 177–179.

  182. Ramsay, E.P. & Ogilby, J.D. (1887) Descriptions of new Australian fishes. Proceedings of the Linnean Society of New South Wales, Series 2, 2 (3), 561–564.

  183. Randall, J.E. (1982) Examples of antitropical and antiequatorial distribution of Indo-West Pacific fishes. Pacific Science, 35, 197–209. [1981]

  184. Randall, J.E. (1995) Coastal fishes of Oman. Crawford House Publishing Pty Ltd, Bathurst, 439 pp.

  185. Randall, J.E. (2001) Reminiscing. Atoll Research Bulletin, 494, 23–50.

  186. Randall, J.E. (2005) Reef and shore fishes of the South Pacific, New Caledonia to Tahiti and the Pitcairn Islands. University of Hawai’i Press, Honolulu, 707 pp.

  187. Randall, J.E., Allen, G.R. & Steene, R.C. (1990) Fishes of the Great Barrier Reef and Coral Sea. University of Hawaii Press, Honolulu, xx + 507 pp.

  188. Randall, J.E. & Anderson, R.C. (1993) Annotated checklist of the epipelagic and shore fishes of the Maldive Islands. J.L.B. Smith Institute of Ichthyology Ichthyological Bulletin, 59, 1–49.

  189. Randall, J.E., Downing, N., McCarthy, L.J. & Stanaland, B.E. (1994) Fifty-one new records of fishes from the Arabian Gulf. Fauna of Saudi Arabia, 14, 220–258.

  190. Randall, J.E. & Lim, K.K.P., eds. (2000) A checklist of the fishes of the South China Sea. Raffles Bulletin of Zoology Supplement, 8, 569–667.

  191. Ray, D. & Mohapatra, A. (2015) First report of two jawfishes (Perciformes: Opistognathidae) from northern east coast of India. Records of the Zoological Society of India, 115 (1), 109–112.

  192. Regan, C.T. (1913) The classification of the percoid fishes. Annuals of Magazine and Natural History, London, Series 8, 12, 111–145.

  193. Richer de Forges, B., Bouchet, P., Dayrat, B., Warén & Philippe, J.-S. (2000) La campagne BORDAU I sur la ride de Lau (iles Fidji). Compte rendu et liste des stations. In: Crosnier, A. (ed.). Résultats des Campagnes MUSORSTOM. Vol. 21. Mémoires du Muséum national d’Histoire naturelle, 184, 25–38.

  194. Robertson, D.R. & Smith-Vaniz, W.F. (2008) Rotenone: an essential but demonized tool for assessing marine fish biodiversity. Bioscience, 58 (2), 165–170.


  196. Robertson, D.R. & Smith-Vaniz, W.F. (2010) Use of clove oil in collecting coral reef fishes for research. Marine Ecology Progress Series, 401, 295–302.

  197. Roughley, T.C. (1937) Wonders of the Great Barrier Reef. Angus & Robertson Limited, Sydney, 282 pp.

  198. Rüppell, E. (1830) Atlas zu Reise im nordlichen Afrika. Fische des Rothen Meeres, Frankfurt-am-Main, 16, 95–118,

  199. Russell, B.C. (1983) Checklist of fishes of the Great Barrier Reef Marine Park Capricorn Section. Great Barrier Reef Marine Park Authority Special Publication, Series 1, 1–184.

  200. Russell, B.C. & Houston, W. (1989) Offshore fishes of the Arafura Sea. The Beagle, records of the Northern Territory Museum of Arts and Sciences, 6 (1), 69–84.

  201. Russell, B.C., Fraser, T.H. & Larson, H.K. (2010) Castelnau’s collection of Singapore fishes described by Pieter Bleeker. Raffles Bulletin of Zoology, 58 (1), 93–102.

  202. Sadovy, Y. & Cornish, A.S. (2000) Reef fishes of Hong Kong. Hong Kong University Press, Aberdeen, xi + 321 pp.

  203. Sainsbury, K.J., Kailola, P.J. & Leyland, G.G. (1984) Continental Shelf Fishes of Northern and North-Western Australia. Cloustan & Hall and Peter Pownall Fisheries Information Service Canberra, 375 pp.

  204. Saruwatari, T., Lopez, J.A. & Pietsch, T.W. (1997) Cyanine blue: a versatile and harmless stain for specimen observation. Copeia, 1997 (4), 840–841.

  205. Satapoomin, U. (1999) A survey of fish fauna at the Cape Panwa Reef, southeastern Phuket. Phuket Marine Biological Center Research Bulletin, 62, 9–32.

  206. Satapoomin, U. (2011) The fishes of southwestern Thailand, the Andaman Sea - A review of research and provisional checklist of species. Phuket Marine Biological Center Research Bulletin, 70, 29–77.

  207. Saville-Kent, W.S. (1889) Preliminary observations on a natural history collection made in connection with the surveying cruise of H.M.S. Myrmidon, at Port Darwin and Cambridge Gulf–September to November 1888. Proceedings of the Royal Society of Queensland, 6, 219–242.

  208. Schroeder, R.E. (1980) Philippine shore fishes of the western Sulu Sea. National Media Production Center, Manila, 266 pp.

  209. Senou, H., Matsuura, K. & Shinohara, G. (2006) Checklist of fishes in the Sagami Sea with Zoogeographical comments on shallow water fishes occurring along the coastlines under the influence of the Kuroshio Current. Memoirs of the National Science Museum, Tokyo, 41, 389–542.

  210. Shao, K.-T., Ho, H.-C., Lin, P.-L., Lee, P.-F., Lee, M.-Y., Tsai, C.-Y., Liao, Y.-C. & Lin, Y.-C. (2008) A checklist of the fishes of southern Taiwan, northern South China Sea. The Raffles Bulletin of Zoology, Supplement 19, 233–271.

  211. Shen, S-C. (Ed.) (1995) Fishes of Taiwan. National Taiwan University, Taipei, 956 pp., 208 pls. [in Chinese]

  212. Shen, S.-C., Shao, K.-T., Chen, C.-T., Chen, C.-H., Lee, S.-C. & Mok, H.-K. (1993) Fishes of Taiwan. Department of Zoology, National Taiwan University, Taipei, 960 pp. [in Chinese]

  213. Shen, S.-C., Yu, L.-C. & Yeh, H.-S. (1986) Additions to the fish-fauna from the adjacent waters around Taiwan (I). Journal of the Taiwan Museum, 39 (1), 65–74.

  214. Shiino, S.M. (1972) List of English names of Japanese fishes proposition of new names. Science Report of Shima Marineland, 1, 1–210.

  215. Shinohara, G. (1999) A new jawfish, Stalix toyoshio, from Kyushu, Japan (Perciformes: Opistognathidae). Ichthyological Research, 46 (3), 267–270.

  216. Shinohara, G. (2021) A new jawfish of the genus Opistognathus (Perciformes: Opistognathidae) from Japan. Zootaxa, 4964 (1), 157–168.


  218. Smale, M.J., Watson, G. & Hecht, T. (1995) Otolith atlas of southern African marine fishes. Ichthyological Monographs of the J.L.B. Smith Institute of Ichthyology, 1, i–xiv + 1–253, 149 pls.

  219. Smith, J.L.B. (1935) New and little known fishes from South Africa. Records of the Albany Museum, Grahamstown, 4, 169–235.

  220. Smith, J.L.B. (1949) The Sea Fishes of South Africa. Central News Agency, Ltd., Johannesburg, 580 pp.

  221. Smith, J.L.B. & Smith, M.M. (1963) The Fishes of Seychelles. Department of Ichthyology, Rhodes University, Grahamstown, 77 pp.

  222. Smith, M.M. & Heemstra, P.C. (Eds.) (1986) Smith’s Sea Fishes. Macmillan South Africa, Johannesburg, XX + 1047 pp., 144 pls.

  223. Smith-Vaniz, W.F. (1983) Opistognathus margaretae, a new species of jawfish (Perciformes: Opistognathidae) from the Indian Ocean, with notes on O. nigromarginatus Rüppell and O. muscatensis Boulenger. J.L.B. Smith Institute of Ichthyology, Special Publication, 30, 1–10.

  224. Smith-Vaniz, W.F. (1986) Family No. 225: Opistognathidae. In: Smith, M.M. & Heemstra, P.C. (Eds.), Smith’s Sea Fishes. Macmillan South Africa, Johannesburg, pp. 726–727.

  225. Smith-Vaniz, W.F. (1989) Revision of the jawfish genus Stalix (Pisces: Opistognathidae), with descriptions of four new species. Proceeding of the Academy of Natural Sciences of Philadelphia, 141, 375–407.

  226. Smith-Vaniz, W.F. (1997) Five new species of jawfishes (Opistognathus: Opistognathidae) from the western Atlantic Ocean. Bulletin of Marine Science, 60 (3), 1074–1128.

  227. Smith-Vaniz, W.F. (2000) Family Opistognathidae. In: Randall, J.E. & Lim, K.K.P. (Eds.), A checklist of the fishes of the South China Sea. Raffles Bulletin of Zoology Supplement, 8, pp. 612.

  228. Smith-Vaniz, W.F. (2004) Descriptions of six new species of jawfishes (Opistognathus; Opistognathidae) from Australia. Records of the Australian Museum, 56 (2), 209–224.

  229. Smith-Vaniz, W.F. (2009) Three new species of jawfishes (Opistognathus: Opistognathidae), with the posterior end of the upper jaw produced as a thin flexible lamina. Aqua, International Journal of Ichthyology, 15 (2), 69–108.

  230. Smith-Vaniz, W.F. (2010) New species of Indo-Pacific jawfishes (Opistognathus: Opistognathidae) from the Western Indian Ocean and Red Sea. Smithiana, Publications in Aquatic Biodiversity, 12, 39–54.

  231. Smith-Vaniz, W.F. (2011) Opistognathus albicaudatus, a new species of jawfish (Teleostei: Opistognathidae) from the Andaman Islands. Zootaxa, 3085 (1), 34–40.

  232. Smith-Vaniz, W.F. (2016) Opistognathus ensiferus, a new species of jawfish (Opistognathidae) from the Gulf of Mannar, India, with redescription of O. solorensis Bleeker. Zootaxa, 4196 (2), 278‒288.

  233. Smith-Vaniz, W.F. (2017) Descriptions of a new genus and two new species of Caribbean deep-water jawfishes (Teleostei: Opistognathidae). Journal of the Ocean Science Foundation, 26, 46‒58.

  234. Smith-Vaniz, W.F. (2022) Family Opistognathidae. In: Heemstra, P.C., Heemstra, E., Ebert, D.R., Holleman, W. & Randall, J.E. (Eds.), Coastal fishes of the western Indian Ocean, 4, pp. 309–315

  235. Smith-Vaniz, W.F. & Allen, G.R. (2007) Opistognathus rufilineatus, a new species of jawfish (Opistognathidae) from the Bird’s Head Peninsula, western New Guinea. Aqua, International Journal of Ichthyology, 13 (1), 35–42.

  236. Smith-Vaniz, W.F., Bineesh, K.K. & Akhile, K.V. (2012) Opistognathus pardus, a new species of jawfish (Teleostei: Opistognathidae) from the western Indian Ocean. Zootaxa, 3523 (1), 20–24.

  237. Smith-Vaniz, W.F. & Richards, W.J. (2006) Opistognathidae. In: Richards, W.J. (Ed.), Early stages of Atlantic fishes: An identification guide for the Western Central North Atlantic. Vol. 2. CRC Press, Miami, Florida, pp. 1827–1832.

  238. Smith-Vaniz, W.F. & Walsh, S.J. (2017) Revision of the jawfish genus Lonchopisthus with description of a new Atlantic species (Teleostei: Opistognathidae). Journal of Ocean Science Foundation, 28, 52–89.

  239. Smith-Vaniz, W.F. & Yoshino, T. (1985) Review of Japanese jawfishes of the genus Opistognathus (Opistognathidae) with description of two new species. Japanese Journal of Ichthyology, 32 (1), 18–27.

  240. Soniyama, T., Ogimoto, K., Hori, S., Uchida, Y. & Kawano, M. (2020) An annotated checklist of marine fishes of the Sea of Japan off Yamaguichi Prefecture, Japan, with 74 new records. Bulletin of the Kagoshima University Museum, 11, 1–152.

  241. Springer, V.G. (1982) Pacific Plate biogeography, with special reference to shorefishes. Smithsonian Contributions to Zoology, 367, 1–182.

  242. Springer, V.G. (1993) Definition of the suborder Blennioidei and its included families (Pisces: Perciformes) Bulletin of Marine Science, 52 (1), 472–495.

  243. Springer, V.G. & Johnson, G.D. (2004) Study of the gill-arch musculature of teleostome fishes, with special reference to the Actinopterygii. Bulletin of the Biological Society of Washington, 11, 1–260. [with Appendix by Springer, V.G. and Orrell, T.M.]

  244. Springer, V.G., Smith, C.L. & Fraser, T.H. (1977) Anisochromis straussi, new species of protogynous hermaphroditic fish, and synonymy of Anisochromidae, Pseudoplesiopidae, and Pseudochromidae. Smithsonian Contributions to Zoology, 252, 1–15.

  245. Springer, V.G. & Williams, J.T. (1990) Widely distributed Pacific plate endemics and lowered sea-level. Bulletin of Marine Science, 47 (3), 631–640.

  246. Stanbury, P.J. (1969) Type specimens in the Macleay Museum, University of Sydney. I. Fishes. Proceedings of the Linnean Society of New South Wales, 93 (2), 203–210.

  247. Steene, R. (1998) Coral Seas. Firefly Books (U.S.), Inc., Buffalo, New York, 271 pp.

  248. Tan, W. (2007) Fight club. Australian Scuba Diver, 3 (4), 29–31.

  249. Taquet, M. & Diringer, M.T. (2012) Fishes of the Indian Ocean and Red Sea. Éditions Quae, France, 703 pp.

  250. Tashiro, S. (2017) Family Opistognathidae. In: Motomura, H., Alama, U.B., Muto, N., Babaran, R.P. & Ishikawa, S. (Eds.), Commercial and bycatch market fishes of Panay Island, Republic of the Philippines. The Kagoshima University Museum, Kagoshima, University of the Philippines Visayas, Iloilo, and Research Institute for Humanity and Nature, Kyoto, pp. 1–246.

  251. Tashiro, S., Uyeno, D. & Motomura, H. (2018) First Japanese records of the jawfish Opistognathus solorensis (Actinopterygii: Perciformes: Opistognathidae) from the Osumi Islands. Species diversity, 23, 233–237.

  252. Taylor, W.R. (1964) Fishes of Arnhem Land. Records of the American-Australian Scientific Expedition to Arnhem Land, 4, 45–307.

  253. Valenciennes, A. (1840) Poissons. Livraison 18. In: Cuvier’s Regne animal. Disciples Edition. 3rd Edition. s.n., Paris, pp. 121–128, pls. 69 + 71 + 78 + 79

  254. Voris, H.K. (2000) Maps of Pleistocene sea levels in southeast Asia: shorelines, river systems and time durations. Journal of Biogeography, 27, 1153–1167.

  255. Wainwright, P.C., Smith, W.L., Price, S.A., Tang, K.L., Sparks, J.S., Ferry, L.A., Kuhn, K.L., Eytan, R.I. & Near, T.J. (2012) The evolution of pharyngognathy: A phylogenetic and functional appraisal of the pharyngeal jaw key innovation in labroid fishes and beyond. Systematic Biology, 61, 1001–1027.

  256. Waite, E.R. (1904) New records or recurrences of rare fishes from eastern Australia. No. 3. Records of the Australian Museum, 5 (4), 231–244.

  257. Wass, R.C. (1984) An annotated checklist of the fishes of Samoa. NOAA Technical Report, SSRF-781, 1–43.

  258. Watson, H.G. (1996) Opistognathidae. In: The early stages of fishes in the California Current Region, Moser, H.G. (Ed.), CalCOFI Atlas, 33, pp. 1068–1071

  259. Weber, W. (1913) Die Fische der Siboga-Expedition. Siboga Expedition Report, 57, 1–710.

  260. White, W.T., Last, P.R., Dharmadi, Faizah, R., Chodrijah, U., Prisantoso, B.I., Pogonoski, J.J., Puckridge, M. & Blaber, S.J.M. (2013) Market fishes of Indonesia. Australian Centre for International Agricultural Research, Monograph 155, 1–438 pp.

  261. Whitley, G.P. (1932) Fishes. Great Barrier Reef Expedition, 1928–29: Scientific reports, 4 (9), 267–316.

  262. Whitley, G.P. (1937) Studies in Ichthyology No. 10. Records of the Australian Museum, 20 (1), 3–24.

  263. Whitley, G.P. (1948) A list of the fishes of Western Australia. Western Australia Fisheries Bulletin, 2, 1–35.

  264. Whitley, G.P. (1964) Presidential address. A survey of Australian Ichthyology. Proceedings of the Linnean Society of New South Wales, 89 (1), 11–127.

  265. Whitley, G.P. (1966) Notes on some Queensland fishes. Australian Zoologist, 13 (3), 235–243.

  266. Winterbottom, R. (1974) A descriptive synonymy of the striated muscles of the Teleostei. Proceedings of the Academy of Natural Sciences of Philadelphia, 125 (12), 225–317.

  267. Wongratana, T. (1975) Description of a new jaw-fish, Opisthognathus rex from Thailand (Pisces: Opisthognathidae). Natural History Bulletin of the Siam Society, 26 (1–2), 98–104.

  268. Xu, C., Denig, S., Xiong, G. & Zhan, H. (1980) Two new fishes from east China Sea. Oceanologia et Limnologia Sinica, 11 (2), 179–184.

  269. Yamada, U., Horikawa, H., Yamashita, H. & Tagawa, M. (1997) Opistognathus sp. Seikaiku Suisan Kenkyusho News, 90, 1. [in Japanese]

  270. Yoshida, T., Motomura, H., Musikasinthorn, P. & Matsuura, K. (Eds.) (2013) Fishes of northern Gulf of Thailand. National Museum of Nature and Science, Tsukuba, Research Institute for Humanity and Nature, Kyoto and Kagoshima University Museum, Kagoshima, 239 pp.

  271. Zajonz, U., Lavergne, E., Bogorodsky, S.V., Saeed, F.N., Aideed, M.S. & Krupp, F. (2019) Coastal fish diversity of the Socotra Archipelago, Yemen. Zootaxa, 4636 (1), 1–108.