Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2023-05-26
Page range: 265-274
Abstract views: 1118
PDF downloaded: 23

Description of a geographically variable elongate rock-dwelling cichlid (Cichliformes: Cichlidae) from Lake Malaŵi, Africa

Penn State University; 432 Forest Resources Building; University Park; Pennsylvania 16802; USA. Honorary Research Associate; South African Institute for Aquatic Biodiversity; Makhanda; RSA
Cichlid Press; PO Box 13608; El Paso; TX 79913
Pisces mbuna Lake Malawi Metriaclima


The rock-dwelling cichlids from Lake Malaŵi, known as mbuna, comprise a diverse group of haplochromine fishes that are placed among 14 genera. Within the mbuna, Pseudotropheus is a polyphyletic genus, which has served as a catch-all for many of these fishes. Recently, many of the species-groups within Pseudotropheus have been elevated to separate genera. Herein, we describe an elongate form that was originally placed in the Pseudotropheus elongatus species group but is now described as a member of Metriaclima.



  1. Barel, C.D.N., Van Oijen, M.J.P., Witte, F. & Witte-Maas, E.L.M. (1977) An introduction to the taxonomy and morphology of the haplochromine Cichlidae from Lake Victoria. Netherlands Journal of Zoology, 27 (4), 333–389. DOI:
  2. Bowers, N.J. & Stauffer Jr., J.R. (1993) A new species of rock-dwelling cichlid (Pisces: Cichlidae) from Lake Malaŵi, Africa, with comments on Melanochromis vermivorus Trewavas. Copeia, 1993, 715–722. DOI:
  3. Condé, B. & Géry, J. (1999) Maylandia Meyer et Foerster, 1984, un nom générique disponible (Teleostei, Perciformes, Cichlidae). Revue française d’Aquariologie, 26, 21–22.
  4. Geerts, M. (2002) Stripes do not a zebra make Maylandia versus Metriaclima. Cichlid News, 11, 21–26.
  5. Humphries, J.M., Bookstein, F.L., Chernoff, B., Smith, G.R., Elder, R.L. & Poss, S.G. (1981) Multivariate discrimination by shape in relation to size. Systematic Zoology, 30 (3), 291–308. DOI:
  6. Konings, A. (1995) Malaŵi cichlids in their natural habitat. 2nd Edition. Cichlid Press, Lauenau. [unknown pagination]
  7. Konings, A. (2016) Malaŵi cichlids in their natural habitat. 5th Edition. Cichlid Press, El Paso, Texas. [unknow pagination]
  8. Konings, A.F. & Geerts, M. (1999) Comments on the revision of Pseudotropheus zebra. Cichlid News, 8, 18–26.
  9. Konings, A.F. & Stauffer Jr., J.R. (2006) Revised diagnosis of Metriaclima (Teleostei: Cichlidae) with description of a new species from Lake Malaŵi National Park, Africa. Ichthyological Exploration of Freshwaters, 17 (3), 233–246.
  10. Li, S., Konings, A.F. & Stauffer, J.R. Jr. (2016) A revision of the Pseudotropheus elongatus species group (Teleostei: Cichlidae) with a description of a new genus and seven new species. Zootaxa, 4168 (2), 353–381. DOI:
  11. Meyer, M.K. & Foerster, W. (1984) Un nouveau Pseudotropheus du lac Malawi avec des remarques sur le complexe Pseudotropheus-Melanochromis (Pisces, Perciformes, Cichlidae). Revue française d’Aquariologie, 10, 107–112.
  12. Oliver, M.K. & Arnegard, M.E. (2010) A new genus for Melanochromis labrosus, a problematic Lake Malaŵi cichlid with hypertrophied lips (Teleostei: Cichlidae). Ichthyological Exploration of Freshwaters, 21 (3), 209–232.
  13. Oliver, M.L. & Loiselle, P.V. (1972) A new genus and species of cichlid of the mbuna group (Pisces: Cichlidae) from Lake Malaŵi. Revue de Zoologie et de Botanique Africaines, 855, 309–320.
  14. Ribbink, A.J., Marsh, B.A., Marsh, A.C., Ribbink, A.C. & Sharp, B.J. (1983) A preliminary survey of the cichlid fishes of rocky habitats in Lake Malaŵi. South African Journal of Zoology, 18 (3), 149–310. DOI:
  15. Stauffer, J.R. Jr., Black, K. & Konings, A.F. (2013) Descriptions of five new species of Metriaclima (Teleostei: Cichlidae) from Lake Malaŵi, Africa. Zootaxa, 3647 (1), 101–136. DOI:
  16. Stauffer, J.R. Jr., Bowers, N.J., Kellogg, K.A. & McKaye, K.R. (1997) A revision of the blue-black Pseudotropheus zebra (Teleostei: Cichlidae) complex from Lake Malaŵi, Africa, with a description of a new genus and ten new species. Proceedings of the Academy of Natural Sciences of Philadelphia, 148, 189–230.
  17. Stauffer Jr., J.R. & Kellogg, K.A. (2002) Description of a new species in the Pseudotropheus williamsi complex (Teleostei: Cichlidae), from Lake Malaŵi, Africa. Copeia, 2002, 146–151. DOI:[0146:DOANSI]2.0.CO;2
  18. Trewavas, E. (1935) A synopsis of the cichlid fishes of Lake Nyasa. Annals and Magazine of Natural History, Series 10, 16 (91), 65–118. DOI: