Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2023-07-20
Page range: 339-362
Abstract views: 619
PDF downloaded: 20

Resolving taxonomic issues of cicadas (Hemiptera: Cicadidae) including new combinations, new synonymies, and revised status, with updates on the diversity of the Brazilian cicada fauna and new records for four South American countries

10900 County Road 352; Blanket; TX 76432; USA
Hemiptera Taxonomy new combination new synonymy new record alpha diversity

Abstract

The taxonomic position and status of historically confused cicada taxa is determined. Species are reassigned based on analysis of type specimens and the application of currently accepted higher taxonomy. Diceroprocta grossa (Fabricius, 1775) is determined to be Megatibicen grossa (Fabricius, 1775) n. comb. with Cicada auletes Germar, 1834 n. syn., Fidicina literata Walker, 1850 n. syn., and Cicada sonora Walker, 1850 n. syn. becoming junior synonyms. Abroma egae (Distant, 1892) is determined to be Brevialavenosa egae (Distant, 1892) n. comb., Abroma temperata (Walker, 1858) is reassigned to become Brevialavenosa temperata (Walker, 1858) n. comb., and Abroma apicifera (Walker, 1850) is determined to be Brevialavenosa apicifera (Walker, 1850) n. comb. Cicada sahlbergi Stål, 1854 is reassigned to become Herrera sahlbergi (Stål, 1854) n. comb. Zammara eximia Erichson, 1848 is reassigned to become Zammaralna eximia (Erichson, 1848) n. comb. Zammara strepens Amyot & Audinet-Serville, 1843 is reassigned to become Orellana strepens (Amyot & Audinet-Serville, 1843) n. comb. Orellana brunneipennis Goding, 1925 is reassigned to become Zammara brunneipennis (Goding, 1925) n. comb. and Orellana pulla Goding, 1925 is reassigned to become Zammara pulla (Goding, 1925) n. comb. Zammara brunneipennis (Goding, 1925) n. comb., n. syn., Zammara pulla (Goding, 1925) n. comb., n. syn., and Zammara erna Schmidt, 1919 n. syn. are made junior synonyms of Zammara intricata Walker, 1850. Orellana pollyae Sanborn, 2011a n. syn. is made a junior synonym of Orellana strepens (Amyot & Audinet-Serville, 1843) n. comb., and Zammara medialinea Sanborn, 2004 n. syn. is made a junior synonym of Zammara luculenta Distant, 1883. Dorisiana brasilianensis Boulard, 2017 n. syn. is determined to be a junior synonym of Dorisiana noriegai Sanborn & Heath, 2014. Two taxa, Carineta viridicollis var. a Stål, 1862 n. syn. and Cicada proponens var. b Walker, 1858c n. syn. are shown to be unavailable as well as junior synonyms of Carineta viridicollis (Germar, 1830) and Parnisa proponens (Walker, 1858c), respectively. Fidicina aldegondae Kulgatz, 1902 (in Kulgatz & Melichar, 1902) rev. stat. is removed from synonymy and reassigned to become Guyalna aldegondae (Kulgatz, 1902 in Kulgatz & Melichar, 1902) rev. stat., n. comb. Cicada costa Dohrn, 1859 is shown to be a misspelling of Cicada casta Stål, 1854 and does not represent a nomen nudum. The alpha diversity of the known Brazilian cicada fauna is now 181 species with new records of Proarna uruguayensis Berg, 1882 and Herrera dentata Sanborn, 2020 and the removal of Diceroprocta grossa (Fabricius, 1775) and Odopoea degiacomii Distant, 1912 from the fauna. New records of Hemisciera maculipennis (de Laporte, 1832) for Bolivia, Guyalna cuta (Walker, 1850) for Paraguay, and Pachypsaltria phaedima Torres, 1960 for Peru are provided.

 

References

  1. Amyot, C.J.B. & Audinet-Serville, A. (1843) Histoire Naturelle des Insectes. Hemiptères. Deuxime partie. Homoptères. Homoptera Latr. Librairie encyclopédique de Roret, Paris, 675 pp. https://doi.org/10.5962/bhl.title.8471 DOI: https://doi.org/10.5962/bhl.title.8471
  2. Audouin, J.V. & Brullé, G.A. (1835) Histoire naturelle des insectes, traitant de leur organization et de leurs moeurs en general, et comprenant leur classification et la description des espèces. Le tout accompagné de planches gravées sur acier, d’après des peintures executes pour cette edition sur la collection du Muséum de Paris. F.D. Pillot, Paris, 56 pls. https://doi.org/10.5962/bhl.title.34228 DOI: https://doi.org/10.5962/bhl.title.34228
  3. Berg, C. (1882) Contribuciones al estudio, de las Cicadidae de la República Argentina y países limítrofes. Anales de la Sociedad Cientifica Argentina, 14, 38–48.
  4. Billberg, G.J. (1820) Enumeratio inectorum in Museo. Typis Gadelianis, Stockholm, 138 pp. https://doi.org/10.5962/bhl.title.49763 DOI: https://doi.org/10.5962/bhl.title.49763
  5. Blanchard, E. (1840) Homoptères. Histoire naturelle des insectes Orthoptères, Névroptères, Hémiptères, Lépidoptères et Diptères, 3, 1–672, 71 pls. https://doi.org/10.5962/bhl.title.124185 DOI: https://doi.org/10.5962/bhl.title.59226
  6. Boer, A.J. de (1995) The phylogeny and taxonomy of the Chlorocystini (sensu stricto) (Homoptera, Tibicinidae). Contributions to Zoology, 65, 201–231. https://doi.org/10.1163/26660644-06504001 DOI: https://doi.org/10.1163/26660644-06504001
  7. Boulard, M. (1976) Sur une deuxieme cigale africaine depourvue d’appareil sonore (Homoptera). Bulletin de l’Institut Fondamental d’Afrique Noire, Series A, 37, 629–636.
  8. Boulard, M. (1986) Nouvelles cigales guyano-amazoniennes du genre Carineta (Homoptera, Tibicinidae). Nouvelle Revue d’Entomologie, New Series, 2, 415–429.
  9. Boulard, M. (2017) Michel Boulard, Entomologiste, éco-éthologiste, photographe. Autobiographie d’un bourlingueur. Ecole Pratique des Hautes Etudes, Laboratoire Biologie et Evolution des Insectes, 21a, i–xii + 1–352.
  10. Boulard, M. (2022) My life with the cicadas, some nearby relations and others. Ecole Pratique des Hautes Etudes, Laboratoire Biologie et Evolution des Insectes, Paris, 512 pp.
  11. Boulard, M. & Martinelli, N.M. (1996) Révision des Fidicini, nouveau statut de la tribu, espèces connues et nouvelles espèces (Cicadomorpha, Cicadidae, Cicadinae). Premiere partie: Sous-tribu nouvelle des Fidicinina. Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea, 9, 11–81.
  12. Boulard, M. & Martinelli, N.M. (2011) Nouvelles cigales néotropicales de la sous-tribu des Guyalinina (Rhynchota, Cicadidae, Cicadinae, Fidicinini). Lambillionea, 61, 219–232.
  13. Buckton, G.B. (1890) s.n. In: Mongraph of the British Cicadae, or Tettigiidae, illustrated by more than four hundred coloured drawings. Vol. I. Part III. Macmillan and Co., London, pp. xxxiv–xlviii + 65–96, pls. xxi–xxx. https://doi.org/10.5962/bhl.title.9473 DOI: https://doi.org/10.5962/bhl.title.9473
  14. Delétang, L.F. (1923) Monografia de los Cicádidos (Cicadidae) argentinos y relacion de estos con la fauna sudamericana. Anales del Museo Nacional de Historia Natural de Buenos Aires, 31, 538–649. https://doi.org/10.22201/iie.18703062e.1938.2.47 DOI: https://doi.org/10.22201/iie.18703062e.1938.2.47
  15. Distant, W.L. (1883) Contributions to a proposed monograph of the homopterous family Cicadidae. Part I. Proceedings of the Zoological Society of London, 1883, 187–194, pl. 25. https://doi.org/10/1111/j/1469-7998.1883.tb06649.x DOI: https://doi.org/10.1111/j.1469-7998.1883.tb06649.x
  16. Distant, W.L. (1892) On some undescribed Cicadidae, with synonymical notes. Annals and Magazine of Natural History, Series 6, 10, 54–67. https://doi.org/10.1080/00222939208677373 DOI: https://doi.org/10.1080/00222939208677373
  17. Distant, W.L. (1904) Rhynchotal notes XXVII. Annals and Magazine of Natural History, Series 7, 14, 329–336. https://doi.org/10.1080/03745480409443017 DOI: https://doi.org/10.1080/03745480409443017
  18. Distant, W.L. (1905a) Rhynchotal notes XXXIII. Annals and Magazine of Natural History, Series 7, 16, 22–35. https://doi.org/10.1080/03745480509443650 DOI: https://doi.org/10.1080/03745480509443650
  19. Distant, W.L. (1905b) Rhynchotal notes XXXIV. Annals and Magazine of Natural History, Series 7, 16, 203–216. https://doi.org/10.1080/03745480509443671 DOI: https://doi.org/10.1080/03745480509443671
  20. Distant, W.L. (1905c) Rhynchotal notes XXXV. Annals and Magazine of Natural History, Series 7, 16, 265–281. https://doi.org/10.1080/03745480509442862 DOI: https://doi.org/10.1080/03745480509442862
  21. Distant, W.L. (1905d) Rhynchotal notes XXXI. Annals and Magazine of Natural History, Series 7, 15, 379–387. https://doi.org/10.1080/03745480509443064 DOI: https://doi.org/10.1080/03745480509443064
  22. Distant, W.L. (1905e) Rhynchotal notes XXX. Annals and Magazine of Natural History, Series 7, 15, 304–319. https://doi.org/10.1080/03745480509443047 DOI: https://doi.org/10.1080/03745480509443047
  23. Distant, W.L. (1905f) Rhynchotal notes XXXII. Annals and Magazine of Natural History, Series 7, 15, 478–486. https://doi.org/10.1080/03745480509442837 DOI: https://doi.org/10.1080/03745480509442837
  24. Distant, W.L. (1906) A synonymic catalogue of Homoptera. Part I. Cicadidae. Trustees of the British Museum, London, 207 pp. https://doi.org/10.5962/bhl.title.8554 DOI: https://doi.org/10.5962/bhl.title.8554
  25. Dohrn, F.A. (1859) Catalogus Hemipterorum. Herausgegeben von dem Entomologischen Verein zu Stettin, Buchdruckerei von Herrecke & Lebeling, Stettin, 112 pp. https://doi.org/10.5962/bhl.title.8515
  26. Dorval, A., Filho, O.P., Costa, R.B. da & de Souza, M.D. (2011) Diversidade de cigarras (Hemiptera-Homoptera) em ambientes florestais no município de Cotriguaçu, estado de Mato Grosso. Diversity of cicadas (Hemiptera-Homoptera) in forest environments in the municipality of Cotriguaçu, state of Mato Grosso. Multitemas, Campo Grande, 40, 7–19.
  27. Duffels, J.P. & van der Laan, P.A. (1985) Catalogue of the Cicadoidea (Homoptera, Auchenorhyncha) 1956–1980. Series Entomologica 34. Dr. W. Junk Publishers, Dordrect, 414 pp.
  28. Erichson, W.F. (1848) Homoptera. In: Schomburgk, R. (Ed.), Reisen in Britisch-Guiana in den Jahren 1840–1844 im auftrag Sr. Mäjestat des Königs von Preussen ausgeführt von Richard Schomburgk. Nebst einer Fauna und Flora Guiana’s nach Vorlagen von Johannes Müller, Ehrenberg, Erichson, Klotzsch, Troschel, Cabanis und andern. Band 3. Weber, Leipzig, pp. 533–1260. https://doi.org/10.5962/bhl.title.109982 DOI: https://doi.org/10.5962/bhl.title.109982
  29. Fabricius, J.C. (1775) Systema entomologiae, sistens insectorum classes, ordines, genera, species, adiectis, synonymis, locis, descriptionibus, observationibus. In Officina Libraria Kortii, Flensbvrgi et Lipsiae, 832 pp. https://doi.org/10.5962/bhl.title.36510 DOI: https://doi.org/10.5962/bhl.title.36510
  30. Fabricius J. C. (1803) Systema Rhyngotorum secundum ordines, genera, species, adiectis synonymis, locis, observationibus, descriptionibus. Reichard, Brunsvigae, 314 pp. https://doi.org/10.5962/bhl.title.11644 DOI: https://doi.org/10.5962/bhl.title.11644
  31. Germar, E.F. (1830) Species Cicadarium enumeratae et sub genera distributae. Thon’s Entomologisches Archiv, 2, 37–57.
  32. Germar, E.F. (1834) Observations sur plusieurs especes du genre Cicada, Latr. Revue Entomologique Publiée par Gustav Silbermann, 2, 49–82.
  33. Gmelin, J.F. (1789) Caroli a Linné, Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tom. 1, Pars IV. Impensis Georg. Emanuel. Beer, Lipsiae, pp. 1517–2224. https://doi.org/10.5962/bhl.title.36932 DOI: https://doi.org/10.5962/bhl.title.36932
  34. Goding, F.W. (1925) Synopsis of the Cicadidae of Ecuador. Revista del Colegio Nacional Rocafuerte, 7, 1–34.
  35. Goemans, G. (2016) The classification and phylogeny of the Neotropical cicada tribe Zammarini (Hemiptera, Cicadidae) and a revision of its type genus Zammara Amyot & Audinet-Serville, 1843 and its sister genus Zammaralna Boulard & Sueur, 1996. Ph.D. dissertation, University of Connecticut, Storrs, 206 pp.
  36. Gogala, M., Šporar, K., Sanborn, A.F. & Maccagnan, D.H.B. (2015) New cicada species of the genus Guyalna (Hemiptera: Cicadidae) from Brazil. Acta Entomologica Slovenica, 23, 105–116. https://doi.org/10.1603/ice.2016.112976 DOI: https://doi.org/10.1603/ICE.2016.112976
  37. International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature, fourth edition. International Trust for Zoological Nomenclature, London, 306 pp. https://doi.org/10.5962/bhl.title.50608 DOI: https://doi.org/10.5962/bhl.title.50608
  38. Kuhlgatz, T. & Melichar, L. (1902) VI. Rhynchoten. In: Von Ihrer Königlichen Hoheit der Prinzessin Therese von Bayern auf einer Reise in Sudamerika gesammelte Insekten. Berliner Entomologischer Zeitschrift, 47, pp. 246–278, pls. IV–V. https://doi.org/10.1002/mmnd.47919000312 DOI: https://doi.org/10.1002/mmnd.47919000312
  39. de Laporte, F.L. (1832) Mémoire sur cinquante espèces nouvelles ou peu connues d’insectes. Annales de la Société Entomologique de France, 1, 386–415.
  40. Latreille, P.A. (1802) s.n. In: Histoire naturelle, générale et particulière des crustacés et des insectes. Ouvrage faisant suite à l’histoire naturelle générale et particulière, composée par Leclerc de Buffon, et rédigée par C.S. Sonnini, membre de plusieurs sociétés savantes. Familles naturelles des genres. Tome Troisième. F. Dufast, Paris, pp. i–xii + 13–467. https://doi.org/10.5962/bhl.title.60678 DOI: https://doi.org/10.5962/bhl.title.15764
  41. Linnaeus, C. (1758) Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tomus I. Edito duodecima, reformata. Laurentius Salvius, Holmiae, 824 pp. https://doi.org/10.5962/bhl.title.157601 DOI: https://doi.org/10.5962/bhl.title.542
  42. MacArthur, R.H. & Wilson, E.O. (1967) The theory of island biogeography. Princeton University Press, Princeton, New Jersey, 203 pp.
  43. Maccagnan, D.H.B., Martinelli, N.M., Goemans, G. & Guilherme, E. (2011) Zammara smaragdula Walker, 1858 (Hemiptera: Cicadidae): first record for Brazil. Check List, 7, 563–564. https://doi.org/10.15560/7.4.563 DOI: https://doi.org/10.15560/7.4.563
  44. Maccagnan, D.H.B. & Sanborn, A.F. (2015) Orialella aerizulae (Hemiptera: Cicadidae): first record in Brazil. Florida Entomologist, 98, 984–986. https://doi.org/10.1653/024.098.0331 DOI: https://doi.org/10.1653/024.098.0331
  45. Marshall, D.C., Moulds, M.S., Hill, K.B.R., Price, B.W., Wade, E.J., Owen, C.J., Goemans, G., Marathe, K., Sarkar, V., Cooley, J.R., Sanborn, A.F., Kunte, K., Villet, M.H. & Simon, C. (2018) A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily level classification. Zootaxa, 4424 (1), 1–64. https://doi.org/10.11646/zootaxa.4424.1.1 DOI: https://doi.org/10.11646/zootaxa.4424.1.1
  46. Metcalf, Z.P. (1952) New names in the Homoptera. Journal of the Washington Academy of Sciences, 42, 226–231.
  47. Metcalf, Z.P. (1963a) General catalogue of the Homoptera, Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section I. Tibiceninae. North Carolina State College Contribution, 1502, i–vii + 1–585.
  48. Metcalf, Z.P. (1963b) General catalogue of the Homoptera, Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section II. Gaeninae and Cicadinae. North Carolina State College Contribution, 1502, 587–919.
  49. Metcalf, Z.P. (1963c) General catalogue of the Homoptera, Fascicle VIII. Cicadoidea. Part 2. Tibicinidae. North Carolina State College Contribution, 1564, i–vi + 1–492.
  50. Moulds, M.S. (2005) An appraisal of the higher classification of cicadas (Hemiptera: Cicadoidea) with special reference to the Australian fauna. Records of the Australian Museum, 57, 375–446. https://doi.org/10.3853/j.0067-1975.57.2005.1447 DOI: https://doi.org/10.3853/j.0067-1975.57.2005.1447
  51. Nunes, V.L., Ruschel, T.P., Maccagnan, D.H.B., Simões, P.C. & Acosta, R.C. (2023) Cigarras do Brasil, Guia Fotográphica. 1a edição. ebook. 46 pp. [https://zenodo.org/record/7712483]
  52. Olivier, G.A. (1790) Cigale, Cicada. Encyclopedie Méthodique Histoire Naturelles Insectes, 5, 735–760.
  53. Ruschel, T.P. (2017) Seven new species of the cicada genus Guyalna Boulard & Martinelli, 1996 (Hemiptera: Cicadidae: Fidicinini) with a re-description of the type species. Zootaxa, 4281 (1), 246–279. https://doi.org/10.11646/zootaxa.4281.1.25 DOI: https://doi.org/10.11646/zootaxa.4281.1.25
  54. Ruschel, T.P. (2018) Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae). Acta Entomologica Musei Nationalis Pragae, 58, 599–566. https://doi.org/10.2478/aemnp-2018-0047 DOI: https://doi.org/10.2478/aemnp-2018-0047
  55. Ruschel, T.P., Bianchi, F.M., Campos, L.A. & G.S. Carvalho, G.S. (2023) Total evidence analysis elucidates the tangled systematic scenario within Fidicinini (Hemiptera: Auchenorrhyncha, Cicadidae). Arthropod Systematics & Phylogeny, 81, 35–77. https://doi.org/10.3897/asp.81.e85755 DOI: https://doi.org/10.3897/asp.81.e85755.figure10
  56. Ruschel, T.P. & Sanborn, A.F. (2021) Two new genera and new species of Fidicinini Distant, 1905 with a re-description of Nosola Stål, 1866 (Hemiptera, Auchenorrhyncha, Cicadidae). Zootaxa, 4920 (4), 509–527. https://doi.org/10.11646/zootaxa.4920.4.3 DOI: https://doi.org/10.11646/zootaxa.4920.4.3
  57. Sanborn, A.F. (2004) Two new Zammara species from South America (Hemiptera: Cicadoidea: Cicadidae). Florida Entomologist, 87, 365–371. https://doi.org/10.1653/0015-4040(2004)087[0365:tnzsfs]2.0.co;2 DOI: https://doi.org/10.1653/0015-4040(2004)087[0365:TNZSFS]2.0.CO;2
  58. Sanborn, A.F. (2008a) The identity of Cicada tibicen Linné, (=Tibicen chloromerus (Walker, 1850)) (Hemiptera: Cicadoidea: Cicadidae). Entomological News, 119, 227–231. https://doi.org/10.3157/0013-872x(2008)119[227:tioctl]2.0.co;2 DOI: https://doi.org/10.3157/0013-872X(2008)119[227:TIOCTL]2.0.CO;2
  59. Sanborn, A.F. (2008b) New records of Brazilian cicadas including the description of a new species (Hemiptera: Cicadoidea: Cicadidae). Neotropical Entomology, 37, 685–690. https://doi.org/10.1590/s1519-566x2008000600010 DOI: https://doi.org/10.1590/S1519-566X2008000600010
  60. Sanborn, A.F. (2011a) A new species of Orellana Distant from Brazil (Hemiptera: Cicadoidea: Cicadidae). Proceedings of the Entomological Society of Washington, 113, 377–384. https://doi.org/10.4289/0013-8797.113.3.377 DOI: https://doi.org/10.4289/0013-8797.113.3.377
  61. Sanborn, A.F. (2011b) Checklist of the cicadas of French Guiana including new records and the description of nine new species (Insecta, Hemiptera, Cicadoidea, Cicadidae). Zoosystema, 33, 377–418. https://doi.org/10.5252/z2011n3a6 DOI: https://doi.org/10.5252/z2011n3a6
  62. Sanborn, A.F. (2011c) Checklist of the cicadas (Insecta: Hemiptera: Cicadidae) of Paraguay including new records for six species. Check List, 7, 465–467. https://doi.org/10.15560/7.4.465 DOI: https://doi.org/10.15560/7.4.465
  63. Sanborn, A.F. (2013) Catalogue of the Cicadoidea (Hemiptera: Cicadoidea). With contributions to the bibliography by Martin H. Villet. Elsevier/Academic Press, San Diego, 1001 pp. https://doi.org/10.1016/b978-0-12-416647-9.00001-2 DOI: https://doi.org/10.1016/B978-0-12-416647-9.00001-2
  64. Sanborn, A.F. (2014) A new genus and new tribe of cicada from South America (Hemiptera: Cicadoidea: Cicadidae) with a note on the taxonomic position of Ahomana Distant, 1905. Proceedings of the Entomological Society of Washington, 116, 339–348. https://doi.org/10.4289/0013-8797.116.3.339 DOI: https://doi.org/10.4289/0013-8797.116.3.339
  65. Sanborn, A.F. (2016a) Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa, 4137 (4), 501–519. https://doi.org/10.11646/zootaxa.4137.4.3 DOI: https://doi.org/10.11646/zootaxa.4137.4.3
  66. Sanborn, A.F. (2016b) Five new species, a new genus and a new record of cicadas from French Guiana with four new combinations and three new synonymies (Insecta, Hemiptera, Cicadoidea, Cicadidae). Zoosystema, 38, 177–199. https://doi.org/10.5252/z2016n2a2 DOI: https://doi.org/10.5252/z2016n2a2
  67. Sanborn, A.F. (2016c) The cicada genus Guyalna Boulard & Martinelli, 1996 (Hemiptera: Cicadidae: Cicadinae: Fidicinini): generic description, twelve new combinations, and a key to species. Zootaxa, 4105 (5), 430–454. https://doi.org/10.11646/zootaxa.4105.5.2 DOI: https://doi.org/10.11646/zootaxa.4105.5.2
  68. Sanborn, A.F. (2017) Generic redescription, seven new species and a key to the Taphura Stål, 1862 (Hemiptera: Cicadidae: Cicadettinae: Taphurini). Zootaxa, 4324 (3), 451–481. https://doi.org/10.11646/zootaxa.4324.3.3 DOI: https://doi.org/10.11646/zootaxa.4324.3.3
  69. Sanborn, A.F. (2018) The cicadas (Hemiptera: Cicadidae) of Panama including the description of six new species, three new combinations, one new synonymy, and nine new records. Zootaxa, 4493 (1), 1–69. https://doi.org/10.11646/zootaxa.4493.1.1 DOI: https://doi.org/10.11646/zootaxa.4493.1.1
  70. Sanborn, A.F. (2019a) The cicadas (Hemiptera: Cicadidae) of Bolivia including the descriptions of fifteen new species, the resurrection of one genus and two species, seven new combinations, six new synonymies, and twenty-eight new records. Zootaxa, 4655 (1), 1–104. https://doi.org/10.11646/zootaxa.4655.1.1 DOI: https://doi.org/10.11646/zootaxa.4655.1.1
  71. Sanborn, A.F. (2019b) The cicada genus Selymbria Stå1, 1861 (Hemiptera: Cicadidae: Tibicininae: Selymbrini): redescription including ten new species and a key to the genus. Zootaxa, 4614 (3), 401–448. https://doi.org/10.11646/zootaxa.4614.3.1 DOI: https://doi.org/10.11646/zootaxa.4614.3.1
  72. Sanborn, A.F. (2019c) Two new species of Neotropical cicadas (Hemiptera: Cicadidae: Cicadettinae: Carinetini) from Bolivia. Journal of Insect Biodiversity, 13, 27–35. https://doi.org/10.12976/jib/2019.13.2.2 DOI: https://doi.org/10.12976/jib/2019.13.2.2
  73. Sanborn, A.F. (2020a) The cicadas (Hemiptera: Cicadidae) of Peru including the descriptions of twenty-four new species, three new synonymies, and thirty-seven new records. Zootaxa, 4785 (1), 1–129. https://doi.org/10.11646/zootaxa.4785.1.1 DOI: https://doi.org/10.11646/zootaxa.4785.1.1
  74. Sanborn, A.F. (2020b) Two new species of Neotropical cicadas (Hemiptera: Cicadidae: Cicadettinae) from southeastern Brazil. Journal of Insect Biodiversity, 19, 14–23. https://doi.org/10.12976/jib/2020.19.1.3 DOI: https://doi.org/10.12976/jib/2020.19.1.3
  75. Sanborn, A.F. (2020c) Dyticodopoea and Pygmaeodopoea, new genera for the Central American cicada species previously assigned to Odopoea Stål, 1861 (Hemiptera: Cicadoidea: Cicadidae: Cicadinae: Zammarini). Proceedings of the Entomological Society of Washington, 122, 117–126. https://doi.org/10.4289/0013-8797.122.1.117 DOI: https://doi.org/10.4289/0013-8797.122.1.117
  76. Sanborn, A.F. (2020d) The cicadas (Hemiptera: Cicadidae) of Ecuador including the descriptions of five new species, a new subtribe, four new synonymies, and fifteen new records. Zootaxa, 4880 (1), 1–80. https://doi.org/10.11646/zootaxa.4880.1.1 DOI: https://doi.org/10.11646/zootaxa.4880.1.
  77. Sanborn, A.F. (2020e) The cicadas (Hemiptera: Cicadidae) of Suriname including the description of two new species, five new combinations, and three new records. Zootaxa, 4881 (3), 453–481. https://doi.org/10.11646/zootaxa.4881.3.2 DOI: https://doi.org/10.11646/zootaxa.4881.3.2
  78. Sanborn, A.F. (2021a) The cicadas (Hemiptera: Cicadoidea: Cicadidae) of Madagascar including a new tribe, five new genera, twelve new species, four new species synonymies, five revised species status, ten new combinations, new tribal assignments for four genera, one new subtribe synonymy, a checklist and key to the species. Zootaxa, 4937 (1), 1–79. https://doi.org/10.11646/zootaxa.4937.1.1 DOI: https://doi.org/10.11646/zootaxa.4937.1.1
  79. Sanborn, A.F. (2021b) A new genus and species of Neotropical Taphurini Distant, 1905 (Hemiptera: Cicadoidea: Cicadidae: Cicadettinae) from Brazil with a note on the taxonomic position of Prosotettix Jacobi, 1907. Proceedings of the Entomological Society of Washington, 123, 190–198. https://doi.org/10.4289/0013-8797.123.1.190 DOI: https://doi.org/10.4289/0013-8797.123.1.190
  80. Sanborn, A.F. & Heath, M.S. (2014) The cicadas of Argentina with new records, a new genus and fifteen new species (Hemiptera: Cicadoidea: Cicadidae). Zootaxa, 3883 (1), 1–94. https://doi.org/10.11646/zootaxa.3883.1.1 DOI: https://doi.org/10.11646/zootaxa.3883.1.1
  81. Sanborn, A.F. & Heath, M.S. (2016) Megatibicen n. gen., a new North American cicada genus for some former members of the genus Tibicen Latreille, 1825 (Hemiptera: Cicadidae: Cicadinae: Cryptotympanini). Zootaxa, 4168 (3), 577–582. https://doi.org/10/11646/zootaxa.4168.3.10 DOI: https://doi.org/10.11646/zootaxa.4168.3.10
  82. Sanborn, A.F. & Heath, M.S. (2017) Catalogue of the cicadas (Hemiptera: Cicadoidea: Cicadidae) of continental North America North of Mexico. 2nd Edition. Thomas Say Publications in Entomology: Monographs. Entomological Society of America, Lanham, Maryland, 302 pp. https://doi.org/10.3390/d5020166 DOI: https://doi.org/10.4182/ZGJB1076
  83. Sanborn, A.F., Heath, J.E., Philllips, P.K., Heath, M.S. & Noriega, F.G. (2011) Thermal adaptation and diversity in tropical ecosystems: evidence from cicadas (Hemiptera, Cicadidae). PLoS ONE, 6 (e29368), 1–12. https://doi.org/10.1371/journal.pone.0029368 DOI: https://doi.org/10.1371/journal.pone.0029368
  84. Sanborn, A.F., Marshall, D.C., Moulds, M.S. & Simon, C. (2020) Redefinition of the cicada tribe Hemidictyini Distant, 1905, status of the tribe Iruanini Boulard, 1993 rev. stat., and the establishment of Hovanini n. tribe and Sapantangini n. tribe (Hemiptera: Cicadidae). Zootaxa, 4747 (1), 133–155. https://doi.org/10.11646/zootaxa.4747.1.5 DOI: https://doi.org/10.11646/zootaxa.4747.1.5
  85. Sanborn, A.F. & Phillips, P.K. (2013) Biogeography of the cicadas (Hemiptera: Cicadidae) of North America, North of Mexico. Diversity, 5, 166–239. https://doi.org/10.3390/d5020166 DOI: https://doi.org/10.3390/d5020166
  86. Santos, R.S. & Martinelli, N.M. (2011) Morphological characterization of five Brazilian species of Fidicinoides (Hemiptera: Cicadidae). Revista Colombiana de Entomologia, 37, 341–345. https://doi.org/10.25100/socolen.v37i2.9098 DOI: https://doi.org/10.25100/socolen.v37i2.9098
  87. Schellenberg, J.R. (1802) [Cercopis lutea and Cicada gracilis]. Entomologische Beyträge, 1, 1–24.
  88. Schmidt, E. (1919) Beitrag zur kenntnie der genera Zammara Amyot et Serville und Orellana Distant. (Rhynchota-Homoptera.). Stettin Entomologische Zeitung. Herausgegeben von dem entomologischen Vereine zu Stettin, 80, 383–394. https://doi.org/10.5962/bhl.title.8515 DOI: https://doi.org/10.5962/bhl.title.8515
  89. Shydlovskyy, I.V. & Holovachov, O.V. (2005) Homopteran insects from the collection of E.-F. Germar in the Zoological Museum of LNU (Catalogue). Ivan Franko National University of Lviv, Zoological Museum, Lviv, 92 pp., 8 pls. [in Russian, English forward]
  90. Simon, C., Gordon, E.R., Moulds, M.S., Cole, J., Haji, D., Lemmon, A.R., Lemmon, E.M., Kortyna, M., Nazario, K., Wade, E.J., Meister, R., Goemans, G., Chiswell, S.M., Pessacq, P., Veloso, C., McCutchen, J.P. & Łukasik, P. (2019) Off-target capture data, endosymbiont genes and morphology reveal a relict lineage sister to all other singing cicadas. Biological Journal of the Linnean Society, 128, 865–886. https://doi.org/10.1093/biolinnean/blz120 DOI: https://doi.org/10.1093/biolinnean/blz120
  91. Stål, C. (1854) Nya Hemiptera. Öfversigt af Svenska Vetenskaps Akademien Förhandlingar, 11, 231–255.
  92. Stål, C. (1861) Genera nonnulla nova Cicadinorum. Annales de la Société Entomologique de France, Series 4, 1, 613–622.
  93. Stål, C. (1862) Bidrag till Rio Janeiro - traktens Hemipter-fauna. II. Handlingar. Kongliga Svenska Vetenskaps Akademien, Stockholm, 3, 1–75.
  94. Stål, C. (1870) Hemiptera insularum Phillippinarum. Bidrag till Philippinska öarnes Hemipter-fauna. Öfversigt af Svenska Vetenskaps Akademien Förhandlingar, 27, 607–776. https://doi.org/10.5962/bhl.title.61898 DOI: https://doi.org/10.5962/bhl.title.61898
  95. Torres, B.A. (1958) Nuevos géneros Acyroneura y Acuticephala, Guaranisaria bicolor nueva especie (Homoptera-Cicadidae). Neotropica, 4, 17–26.
  96. Torres, B.A. (1960) Estudio del género Pachypsaltria Stål (Homoptera-Cicadidae). Actas y Trabajos del Primer Congresso Sudamericano de Zoologia, La Plata, 12–24 Octubre 1959, 3, 223–243.
  97. Walker, F. (1850) List of the Specimens of Homopterous Insects in the Collection of the British Museum. Part I. British Museum Trustees, London, 260 pp. https://doi.org/10.5962/bhl.title.9063
  98. Walker, F. (1852) List of the Specimens of Homopterous Insects in the Collection of the British Museum. Part IV. British Museum Trustees, London, ix + 280 pp., VIII pls. [pp. i–ix + 909–1188, pls. I–VIII] https://doi.org/10.5962/bhl.title.1584 DOI: https://doi.org/10.5962/bhl.title.1584
  99. Walker, F. (1858a) Insecta Saundersiana: or characters of undescribed insects in the collection of William Wilson Saunders, Esq. Homoptera. John van Voorts, London, 117 pp. https://doi.org/10.5962/bhl.title.5112 DOI: https://doi.org/10.5962/bhl.title.5112
  100. Walker, F. (1858b) List of the specimens of Homopterous insects in the collection of the British Museum. Supplement. British Museum Trustees, London, 307 pp. https://doi.org/10.5962/bhl.title.9063
  101. Walker, F. (1858c) List of the specimens of Homopterous insects in the collection of the British Museum. Addenda. British Museum Trustees, London, pp. 308–369. https://doi.org/10.5962/bhl.title.9063 DOI: https://doi.org/10.5962/bhl.title.9063