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Published: 2023-08-10
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On some commensal anomuran and brachyuran crabs (Crustacea: Decapoda) off Visakhapatnam, Northwestern Bay of Bengal

Fishery Survey of India; Beach Road; Visakhapatnam; Andhra Pradesh; India
Fishery Survey of India; Beach Road; Visakhapatnam; Andhra Pradesh; India
Fishery Survey of India; Beach Road; Visakhapatnam; Andhra Pradesh; India
Fishery Survey of India; Beach Road; Visakhapatnam; Andhra Pradesh; India
Fishery Survey of India; Beach Road; Visakhapatnam; Andhra Pradesh; India
Fishery Survey of India; Beach Road; Visakhapatnam; Andhra Pradesh; India
Fishery Survey of India; New Fishing Jetty; Sasson Dock; Colaba; Mumbai; India
Crustacea Coral reefs commensal Brachyura Anomura Visakhapatnam


Commensal anomuran and brachyuran crabs were collected from extensive exploratory surveys on rocky areas where soft corals, sea anemones and sponges are found off the Visakhapatnam coast. The present study includes nine species of commensal decapod crustaceans, comprising four porcellanid crabs (Petrolisthes lamarckii (Leach, 1820), Porcellanella triloba White, 1852, Polyonyx hendersoni Southwell, 1909 and Pisidia gordoni (Johnson, 1970)), one squat lobster (Allogalathea elegans (Adams, 1847) and four brachyuran crabs (Dorippoides facchino (Herbst, 1785), Quadrella coronata Dana, 1852, Quadrella maculosa Alcock, 1898 and Sphenomerides trapezioides (Wood-Mason & Alcock, 1891). Allogalathea elegans, Polyonyx hendersoni, Pisidia gordoni, Quadrella coronata and Quadrella maculosa are new records from off Visakhapatnam. The porcellanid crabs are associated with the sea pen Pteroeides esperi, desmosponges and black soft corals Antipathes sp. whereas the squat lobster was associated with the soft coral Dendronephthya sp. The brachyuran crabs were observed in association with Antipathes sp. Here, we provide details on the morphology, distribution, habitat and commensalism of decapod crustaceans studied.



  1. Adams, A. & White, A. (1848) Crustacea. In: Adams, A.F. (Ed.), Under the command of captain sir Edward belcher, C.B., F.R.A.S., F.G.S., during the years 1843–1846. Vol. VIII. Benham and Leeve, London, pp. 1–66.
  2. Ahyong, S.T. (2007) Decapod Crustacea collected by the NORFANZ Expedition from the northern Tasman Sea: Galatheidae and Polychelidae. Zootaxa, 1593 (1), 1–54.
  3. Alcock, A. & Anderson, A.R.S. (1899) Natural history notes from H. M. Royal Indian Marine Survey Ship Investigator, Commander T. H. Heming, R. N., commanding. Series III. No. 2. An account of the deep-sea Crustacea dredged during the surveying-season of 1897–98. Annals and Magazine of Natural History, Series 7, (13), 1–27.
  4. Alcock, A. (1898) Materials for the carcinological fauna of India, No. 3, The Brachyura Cyclometopa, part 1. The family Xanthidae. Journal of Asiatic Society of Bengal, 67 (1), 67–233.
  5. Alcock, A. (1899a) An Account of the Deep-Sea Brachyura collected by the Royal Indian Marine Survey Ship Investigator. Trustees of the Indian Museum, Calcutta, iv + 85 pp., index, pls. I–IV.
  6. Alcock, A. (1899b) Crustacea. Part VII. In: Illustrations of the Zoology of the Royal Indian Marine Survey Ship Investigator, under the command of Commander T.H. Heming, R.N. Office of the Superintendent of Government Printing, Calcutta, pp. 36–45.
  7. Alcock, A.W. (1896) Materials for the carcinological fauna of India, No.2, The Brachyura Oxystomata. Journal of Asiatic Society of Bengal, 65, 134–296.
  8. Baba, K. (1969) Four new genera with their representatives and six new species of the Galatheidae in the collection of the Zoological Laboratory, Kyushu University, with redefinition of the genus Galathea. Ohmu Japan, 2, 1–32.
  9. Baba, K., Macpherson, E., Lin, C.W. & Chan, T.-Y. (2009) Crustacean fauna of Taiwan: Squat lobsters (Chirostylidae and Galatheidae). National Taiwan Ocean University, Keelung, Taiwan, 311 pp.
  10. Becker, K. (1996) Epibionts on carapaces of some malacostracans from the Gulf of Thailand. Journal of Crustacean Biology, 16 (1), 92–104.
  11. Beleem, I., Poriya, P. & Gohil, B. (2017) Diversity and distribution of porcelain crabs of Gujarat, India. Iranian Journal of Animal Biosystematics, 13 (1), 41–51.
  12. Borradaile, L.A. (1902b) Marine Crustaceans. III. The Xanthidae and other crabs. In: Gardiner, J.S. (Ed.), The Fauna and Geography of the Maldive and Laccadive Archipelagoes: Being the Account of the Work carried on and of the Collections made by an Expedition during the years 1899 and 1900, 1 (III), 237–271, figs. 41–60.
  13. Cabezas, P., Macpherson, E. & Machordom, A. (2011) Allogalathea (Decapoda: Galatheidae): a monospecific genus of squat lobster? Zoological Journal of the Linnaean Society, 162, 245–270.
  14. Castro, P. (1997) Trapeziid crabs (Brachyura: Xanthoidea: Trapeziidae) of French Polynesia. In: Richer De Forges, B. (Ed.), Les fonds meubles des lagons de Nouvelle-Calédonie (Sédimentologie, Benthos). Études & Thèses. Vol. 3. ORSTOM, Paris, pp. 109–139.
  15. Castro, P. (2013) Brachyuran crabs (Crustacea, Brachyura: Crossotonotidae, Ethusidae, Euryplacidae, Goneplacidae, Latreilliidae, Palicidae, Tetraliidae, Trapeziidae) of the Mainbaza, Miriki, and Atimo Vatae expeditions to the Mozambique Channel and Madagascar. In: Ahyong, S.T., Chan, T.-Y., Corbari, L. & Ng, P.K.L. (Eds.), Tropical deep sea benthos. Vol. 27. Memories of Museum Natural History, Paris, 204, pp. 437–466.
  16. Castro, P., Ng, P.K.L. & Ahyong, S.T. (2004) Phylogeny and systematics of the Trapeziidae Miers, 1886 (Crustacea: Brachyura), with the description of a new family. Zootaxa, 643 (1), 1–70.
  17. Chia, D.G.B. & Ng, P.K.L. (2000) A revision of Eumedonus H. Milne Edwards, 1834 and Gonatonotus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with sea urchins. Journal of Natural History, 34, 15–56.
  18. Chopra, B.N. (1933) Further notes on Crustacea Decapoda in the Indian Museum. III. On the Decapod Crustacea collected by the Bengal Pilot Service off the mouth of the River Hughli. Dromiacea and Oxystomata. Records of Indian Museum, 35 (1), 25–52.
  19. Dana, J.D. (1852) Conspectus Crustaceorum, quae in orbisterrarumcircumnavigatione, Carolo Wilkese Classe Reipublicae Foederatae Duce, lexit et descripsit. Proceedings of the Academy of Natural Science of Philadelphia, 6, 73–86.
  20. Daniel, A. & Krishnan, S. (1979) A parthenopid crab, Zebrida adamsii White, 1847 inhabiting interspaces of spines of the sea urchin, Salmacis virgulata L. Agassiz, 1846. Bulletin of Zoological Survey of India, 1 (2), 171–175.
  21. Dai, A. & Yang, S. (1991) Crabs of the China Seas. China Ocean Press, Beijing and Springer-Verlag, Berlin, iv + 608 pp., 295 figs., 74 pls.
  22. Daniel, A. & Prem Kumar, V.K. (1969) Notes on animal relationships: Dromiid crabs, Cryptodromia tuberculata pileifera Alcock, 1899 sheltering beneath commensal sponges. Journal of Bombay Natural History and Society, 65, 240–242.
  23. Dev Roy, M.K. & Nandi, N.C. (2007) Brachyuran bioresources of coastal Andhra Pradesh. National Symposium on Conservation and Valuation of Marine Biodiversity. Zoological Survey of India, pp. 53–66.
  24. Dev Roy, M.K. & Nandi, N.C. (2012) Brachyuran crabs (Crustacea). In: Fauna of Andaman and Nicobar Islands, State Fauna Series, 19 (1). Zoological Survey of India, Kolkata, pp. 185–236.
  25. Fabricius, J.C. (1798) Supplementatum Entomologiae systematicae. Proft & Storch, Hafniae, 572 pp.
  26. Galil, B. & Clark, P.F. (1990) Crustacea Decapoda: notes on trapeziid crabs from New Caledonia including descriptions of two new species. In: Crosnier, A. (Ed.), Résultats des Campagnes MUSORSTOM, volume 6. Memories of Museum Natural History, Paris, Serie A, 145, pp. 369–388, figs. 1–6.
  27. Galil, B. (1986) Quadrella (Brachyura: Xanthoidea: Trapeziidae). Review and revision. Journal of Crustacean Biology, 6 (2), 275–293.
  28. Gravely, F.H. (1927) Crustacea. In: The littoral fauna of Krusadai Islands in the Gulf of Mannar. Bulletin of Madras Museum, 1, pp. 141–155.
  29. Guinot, D. (1967) La faune carcinologique (Crustacea Brachyura) de l’Océan Indien occidental et de la Mer Rouge. Catalogue, remarques biogéographiques et bibliographie. Mémoires de l’Institut Fondamental d’Afrique Noire, 77, 237–352.
  30. Haig, J. (1966c) The Porcellanidae (Crustacea Anomura) of the Iranian Gulf and Gulf of Oman. Videnskabelige Meddelelser Dansk Naturhistrisk Forening i Kjrbenhavn, 129, 49–65.
  31. Haig, J. (1973) Galatheidea (Crustacea, Decapoda, Anomura) collected by the F.I.S. Endeavour. Records of the Australian Museum, 28, 269–289.
  32. Haig, J. (1974) The anomuran crabs of western Australia: Their distribution in the Indian Ocean and adjacent seas. Journal of Marine Biological Association of India, 14 (2), 443–451.
  33. Haig, J. (1978) Contribution toward a revision of the procellanid genus Porcellana (Crustacea: Decapoda: Anomura). Proceedings of the Biological Society of Washington, 91 (3), 706–714.
  34. Haig, J. (1981) Porcellanid crabs from the Indo-West Pacific, Part II. Steenstrupia, 7 (12), 269–291.
  35. Haig, J. (1992) Hong Kong’s porcellanid crabs. In: Morton, B. (ed.), The Marine Flora and Fauna of Hong Kong and Southern China III. Proceedings of the Fourth International Marine Biological Workshop. Hong Kong University Press, Hong Kong, pp. 303–327.
  36. Henderson, J.R. (1888) Report on the Anomura collected by H.M.S. Challenger during the years 1873–76. Report on the scientific results of the voyage of H.M.S. Challenger during the years 1873–76, 27 (69), i–xi + 1–221, pls. 1–21.
  37. Henderson, J.R. (1893) A contribution to Indian carcinology. Transaction of the Linnaean Society of London, Series 2, 5, 325–458, pis. 36–40.
  38. Herbst, J.F.W. (1785) Versucheiner Naturgeschichte der Krabben und Krebsenebs teinersystematischen Beschreibun gihrerverschiedenen Arten, 1 (6), 183–206, pls. 10–13.
  39. Hiller, A., Harkantra, S. & Werding, B. (2010) Porcellanid crabs from Goa, Eastern Arabian Sea (Crustacea: Decapoda: Porcellanidae). Journal of the Bombay Natural History Society, 107 (3), 201–212.
  40. Holthuis, L.B. & Manning, R.B. (1990) Crabs of the subfamily Dorippinae MacLeay, 1838, from the Indo-West Pacific Region (Crustacea: Decapoda: Dorippidae). Research of Crustacea, Special Number 3, 1–151.
  41. Hornell, J. (1922) The Madras Aquarium. Madras Fisheries Investigations 1921. Ser., 2, Madras Fisheries Dept of Bulletin, 14, 57–96.
  42. Hornell, J. (1923) Some Commensals of Indian Alcyonarians and Crabs. Journal of Bombay Natural History Society, 28 (4), 926–936.
  43. Huys, R., Low, M.Y., De Grave, S., Ng, P.K.L. & Clark, P.F. (2014) On two reports associated with James Wood-Mason and Alfred William Alcock published by the Indian Museum and the Indian Marine Survey between 1890 and 1891: implications for malacostracan nomenclature. Zootaxa, 3757 (1), 1–78.
  44. Johnson, D.S. (1958) The Indo-West Pacific species of the genus Polyonyx (Crustacea, Decapoda, Porcellanidae). Annals of Zoology, 2, 95–118.
  45. Johnson, D.S. (1970) The Galatheidea (Crustacea Decapoda) of Singapore and adjacent waters. Bulletin of the National Museum Singapore, 35, 1–44.
  46. Jones, S. (1959) Notes on animal associations 1. A porcellanid crab on the sea pen, Pteroeides esperi Herklots. Journal of Marine Biological Association of India, 1 (2), 178–179.
  47. Jones, S. & Sankarankutty, C. (1960) Notes on animal associations. 3. A parthenopid crab, Harovia albolineata Adams & White on a mariametrid crinoid, Lamprometra sp. Journal of Marine Biological Association of India, 2 (2), 194–195.
  48. Kathirvel, M. (2008) Diversity in Indian marine brachyuran crabs. Fisheries Technocrats Forum Technical Bulletin, 5, 1–310.
  49. Kathirvel, M. & Gokul, A. (2010) Checklist of brachyuran crabs from the Gulf of Mannar Marine Biosphere Reserve (Revised). Fisheries Techno craft Forum Technical Bulletin, No. 4, 1–45.
  50. Komai, T. (2000) A checklist of Thalassinidae and Anomura (Crustacea: Decapoda) from the South China Sea. Raffles Bulletin of Zoology. Supplement 8, 343–376.
  51. Krishnamoorthy, P. (2009) Brachyuran crabs from the collections of Marine Biological Centre. Records of Zoological Survey of India, Occasional Paper, 304, 1–46.
  52. Kumar, Biju, A., Sushil Kumar, M., Raffi, S.M. & Ajmalkhan, S. (2007) Diversity of brachyuran crabs associated with trawl by-catch in Kerala coast, India. Indian Journal of Fisheries, 54 (3), 283–290.
  53. Kumaralingam, S. & Raghunathan, C. (2017) First report on Neopetrolisthes maculatus (H. Milne Edwards, 1837) (associated with Stichodactyla haddoni (Saville-Kent, 1893) from Grub Island, Andaman Islands. Proceedings of the International Academy of Ecology and Environmental Sciences, 7 (1), 20–24.
  54. Kumaralingam, S., Raghunathan, C. & Venkataraman, K. (2015) An account of new records of reef associated crabs of Andaman and Nicobar Islands. Records of Zoological Survey of India, 115 (3), 255–271.
  55. Kumaralingam, S., Raghunathan, C., Ajithkumar, T.T. & Chandra, K. (2017) Reef associated common crustaceans of Andaman and Nicobar Islands, Lakshadweep and Gulf of Mannar. Occasional Paper of Zoological Survey of India, 380, 1–249.
  56. Laxmilatha, P., Jasmine, S., Sreeram, M.P. & Rengaiyan, P. (2019) Benthic communities of mesophotic coral ecosystem off Puducherry, east coast of India. Current Science, 116 (6), 982–987.
  57. Leach, W.E. (1820) Galatéadées. Dictionaire des Sciences Naturelles, 18, 49–56.
  58. Miyake, S. (1943) Studies on the crab-shaped Anomura of Nippon and adjacent waters. Journal of Department Agriculture Kyushyu University, 7 (3), 49–158, figs. 1–62.
  59. Miyake, S. (1942) Studies on the decapod crustaceans of Micronesia III. Porcellanidae. Palau Tropical Biological Station Studies, 2, 329–379, text-figs. 1–35, pl. 1.
  60. Morgan, G.J. (1990) A collection of Thalassinidea, Anomura and Brachyura (Crustacea: Decapoda) from the Kimberley region of northwestern Australia. Zoologische Verhandelingen, 265, 1–90.
  61. Morton, B. & Morton, J. (1983) The seashore ecology of Hong Kong. Hong Kong University Press, Hong Kong, 350 pp.
  62. Ng, P.K.L., Guinot, D. & Davie, P.J.F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1–286.
  63. Osawa, M. & Chan, T.Y. (2010) Part III. Porcellanidae (porcelain crabs). In: Chan, T.-Y. (Ed.), Crustacean fauna of Taiwan: crab-like anomurans (Hippoidea, Lithoidoidea and Porcellanidae). National Taiwan University, Keelung, pp. 1–197.
  64. Osawa, M. & Mclaughlin, P.A. (2010) Annotated checklist of anomuran decapod crustaceans of the world (exclusive of the Kiwaoidea and families Chirostylidae and Galatheidae of the Galatheoidea). Part II—Porcellanidae. Raffles Bulletin of Zoology, Supplement 23, 109–129.
  65. Panikkar, N.K. & Prasad, R. (1952) Interesting association of ophiuroids, fish and crab with the jellyfish Rhopilema hispidum. Journal of Bombay Natural History Society, 51, 295–296.
  66. Patel, K., Padate, V., Osawa, M., Tiwari, S., Vachhrajani, K. & Trivedi, J. (2022) An annotated checklist of anomuran species (Crustacea: Decapoda) of India. Zootaxa, 5157 (1), 1–27.
  67. Poore, G.C.B., Ahyong, S.T. & Taylor, J. (2011) The Biology of Squat Lobsters. CSIRO, Melbourne, and CRC Press, Boca Raton, Florida, xv + 363 pp.
  68. Poore, G.C.B., McCallum, A.W. & Taylor, J. (2008) Decapod Crustacea of the continental margin of southwestern and central Western Australia: preliminary identifications of 524 species from FRV Southern Surveyor voyage SS10-2005. Museum Victoria Science Report, 11, 1–106.
  69. Poupin, J., Cleva, R., Bouchard, J.M., Dinhut, V. & Dumas, J. (2018) The crabs from Mayotte Island (Crustacea: Decapoda: Brachyura). Atoll Research Bulletin, 617, 1–109.
  70. Prakash, S. & Marimuthu, S. (2020) Notes on some crinoids associated decapods crustaceans (Crustacea: Decapods) of Lakshadweep Archipelago, Central Indian Ocean. Zootaxa, 4766 (1), 86–100.
  71. Prakash, S., Ajith Kumar, T.T. & Ajmal Khan, S. (2013) Checklist of the Porcellanidae (Crustacea: Decapoda: Anomura) of India. Checklist, 9 (6), 1514–1518.
  72. Rao, D.V. (2010) Field Guide to Coral and Coral Associates of Andaman and Nicobar Islands. Zoological Survey of India, Kolkata, 283 pp.
  73. Rathbun, M.J. (1911) The Percy Sladen Trust Expedition to the Indian Ocean in 1905. Under the leadership of Mr. J. Stanley Gardiner. Marine Brachyura. Transactions of the Linnean Society of London, 3 (11), 191–261.
  74. Sakai, T. (1965) The Crabs of Sagami Bay, collected by His Majesty the Emperor of Japan. Maruzen Co., Tokyo, 206 pp. (English text), 27 figs., 100 pls.
  75. Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. In three volumes. Kodansha Ltd., Tokyo, xxix + 773 pp. (English text), 461 pp. (Japanese text), 16 pp., 251 pls. (plates volume).
  76. Sakai, T. (1980) On new or rare crabs taken from Japanese and central Pacific waters. Research Crustacea, 10, 73–84.
  77. Sankarankutty, C. (1961) On the porcellanid crab, Porcellanella triloba (Crustacea-Anomura), a commensal on sea pen, with remarks on allied species. Journal of Marine Biological Association of India, 3 (1–2), 96–100.
  78. Sankoli, K.N. (1963) On a new species of porcellanid crab (Decapoda, Anomura) from India. Journal of the Marine Biological Association India, 5 (2), 280–283.
  79. Sankoli, K.N. (1966) On the Porcellanidae (Crustacea, Anomura) of Ratnagiri along the west coast of India. Proceedings of the Symposium of Crustacea, Cochin, 1, 295–313.
  80. Saravanan, R., Ramamoorthy, N., Sadiq, S., Shanmuganathan, K. & Gopakumar, G. (2015) A note on the obligate symbiotic association between crab Zebrida adamsii White, 1847 (Decapoda: Pilumnidae) and flower urchin Toxopneustes pileolus (Lamarck, 1816) (Camarodonta: Toxopneustidae) from the Gulf of Mannar, India. Journal of Threatened Taxa, 7 (10), 7726–7728.
  81. Sarojini, R. & Nagabhushanam, R. (1968) Notes on porcellanid crabs (Crustacea, Anomura) from Waltair coast. Journal of Zoological Society of India, 20 (1–2), 149–163.
  82. Sastry, D.R.K. (1977) On some crustacean associates of sea-urchins of the Andaman and Nicobar Islands: News Letter. Zoological Survey of India, 3, 119–120.
  83. Sastry, D.R.K. (1981) On some crustacean associates of Echinodermata from the Bay of Bengal. Records of Zoological Survey of India, 79, 19–30.
  84. Schnabel, K.E., Cabezas, P., McCallum, A., Macpherson, E., Ahyong, S.T. & Baba, K. (2011) World-wide distribution patterns of squat lobsters. In: Poore, G.C.B., Ahyong, S.T. & Taylor, J. (Eds.), The Biology of Squat Lobsters. Crustacean Issues. Vol. 19. CSIRO Publishing, Melbourne and CRC Press, Boca Raton, Florida, pp. 149–182.
  85. Serène, R. & Romimohtarto, K. (1969) Observations on the species of Dorippe from the Indo-Malayan Region. Marine Research Indonesia, 9, 1–35.
  86. Serène, R. (1973) Observations sur les espèces des genres Quadrella Dana 1851 et Spheronomides Rathbun 1898 (Decapoda—Brachyura). Bulletin de la Societé zoologique de France, 98 (1), 191–209, figs. 1–28, pls. 1–5.
  87. Shih, H.T. & Mok, H.K. (1996) Quadrella maculosa Alcock, 1898, a genus and species of shallow-water xanthid crab (Brachyura: Xanthoidea: Trapeziidae) new to Taiwan. Zoological Studies, 35, 146–148.
  88. Siddiqui, F.A. & Kazmi, Q.B. (2003) A checklist of marine anomurans (Crustacea: Decapoda) of Pakistan, northern Arabian Sea. Memoirs of Museum Victoria, 60 (1), 87–89.
  89. Sivasubramanian, K., Ravichandran, K. & Anbuchezhian, R. (2014) First discovery of porcellanid crab, Porcellanella picta (Crustacea: Decapods: Porcellanidae), from South-east coast of India. Journal of Asia-Pacific Biodiversity, 7, 248–251.
  90. Southwell, T. (1909) Report on the Anomura collected by Mr. James Hornell at Okha Mandal in Kattiawar in 1905–6, In: Hornell, J. (Ed.), Report to the Government of Baroda on the marine ecology of Okha Mandal in Kattiawar. Part I. Williams and Norgate, London, pp. 105–123.
  91. Sruthi, S., Nansimole, A., Gayathri Devi, T.V. & Tresa, R. (2014) Diversity of brachyuran crabs of Kanyakumari area, South West Coast of India. International Journal of Science and Research, 5, 126–128.
  92. Stimpson, W. (1858) Prodromus descriptions animalium evertebratorum, quae in expeditionead Oceanum Pacificum Septentrionalem, a Republica Federatamissa, Cadwaladaro Ringgold et Johanne Rodgers dueibus, observavit et descripsit. Pars VIL Crustacea Anomura. Proceedings of Academy Natural Science Philadelphia, 10, 225–252.
  93. Thurston, E. (1890) Notes on the Pearl and Chank Fisheries and Marine Fauna of the Gulf of Manaar. Govt. Press, Madras, 124 pp.
  94. Tirmizi, N.M., Yaqoob, M. & Siddiqui, F.A. (1982) An illustrated key to the identification of anomurans (Porcellanidae, Albuneidae and Hippidae) of the northern Arabian Sea. Centre of Excellence, Marine Biology, University of Karachi, Pakistan, 2, 1–29.
  95. Tirmizi, N.M., Yaqoob, M. & Siddiqui, F.A. (1989) Marine fauna of Pakistan: 3 Porcellanid crabs (Crustacea, Anomura). Centre of Excellence, Marine Biology, University of Karachi, Pakistan, 6, 1–46.
  96. Trautwein, S.E. (2007) Four new species of coral crabs belonging to the genus Tetralia Dana, 1851 (Crustacea, Decapoda, Brachyura, Tetraliidae). Zootaxa, 1450 (1), 1–20.
  97. Trivedi, J.N. & Vachhrajani, K.D. (2013) First record of two porcellanid crabs from Gujarat state, India (Crustacea: Decapoda: Porcellanidae). Journal of Marine Biological Association of India, 55 (1), 55–58.
  98. Trivedi, J.N., Trivedi, D.J., Vachhrajani, K.D. & Ng, P.K.L. (2018) An annotated checklist of the marine brachyuran crabs (Crustacea: Decapoda: Brachyura) of India. Zootaxa, 4502 (1), 1–83.
  99. Varadharajan, D. & Soundarapandian, P. (2014) Crab biodiversity from Arukkattuthurai to Pasipattinam, South East coast of India. Indian Journal of Geo-Marine Sciences, 43 (4), 676–698.
  100. Ward, M. (1942) A new genus and eight new species of Brachyura from Mauritius and the Chagos Archipelago. Mauritius Institute of Bulletin, 2 (2), 39–48.
  101. White, A. (1851) Descriptions of some apparently new species of Annulosa, (collected by Mr. Macgillivray during the Voyage of H.M.S. Rattlesnake). Appendix VI. In: MacGillivray, J., Narrative of the Voyage of H.M.S. Rattlesnake, commanded by the late Captain Owen Stanley, R.N., F.R.S. &c. During the years 1846–1850. Including discoveries and surveys in New Guinea, the Louisiade Archipelago, etc. To which is added the account of Mr. E. B. Kennedy’s Expedition for the exploration of the Cape York Peninsula. Vol. II. T. & W. Boone, London, pp. 387–395. [dated 1852, published 1851]
  102. Wood-Mason, J. & Alcock, A. (1891) Natural history notes from H.M. Indian Marine Survey Steamer Investigator, Commander R.F. Hoskyn, R.N., commanding. No. 21. On the Results of the last Season’s Deep-sea Dredging. Annals and Magazine of Natural History, Series 6, 7 (37), 1–19.
  103. Wood-Mason, J. (1891) Report of the Superintendent for the year 1890–91. In: Indian Museum. Annual Report. April 1890 to March 1891. Vol. 22. Printed by of the Trustees, Calcutta, pp. 1–33.
  104. Yang, S.L. & Sun, Z.X. (1990) On the porcellanid crabs from the coast of Fujian Province, China. Bejing Natural History Museum, 45, 1–15.