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Issue: Vol. 5432 No. 4: 3 Apr. 2024
Type: Article
Published: 2024-04-03
DOI: 10.11646/zootaxa.5432.4.2
Page range: 461-508
Abstract views: 106
PDF downloaded: 10

Two New Taxa of Goniasteridae (Asteroidea, Echinodermata) and Noteworthy Observations of Deep-Sea Asteroidea by the NOAA Ship Okeanos Explorer in the North and Tropical Atlantic

Department of Invertebrate Zoology; National Museum of Natural History; Smithsonian Institution; Washington D.C. 20007
Echinodermata NOAA ship Okeanos Explorer deep-sea feeding observations new species taxonomy Stichasteridae Zoroasteridae

Abstract

Two new species and one new genus, collected by NOAA ship Okeanos Explorer, Bathyceramaster kelliottae n. sp. and Rhianastra isosceles n. gen., n. sp. (Goniasteridae; Asteroidea) are described from deep-sea settings in the North Atlantic. Two potentially undescribed species are observed from imagery and new in situ observations further elucidate life modes and feeding behavior of several species from North Atlantic deep-sea habitats. New occurrences of Atlantic Neomorphaster are presented and Neomorphaster forcipatus is synonymized with Neomorphaster margaritaceus.

References

  1. Acloque, A. (1900) Faune de France contenant la description de toutes les espèces indigènes-avec une préface par E. Perrier. Baillière, Paris, 500 pp. https://doi.org/10.5962/bhl.title.34225
  2. Benfield, M.C. & Kupchik, M.J. (2020) Continuing and Expanding a Deepwater Biological Observation System in the Northern Gulf of Mexico. Cooperative agreement No. M13AC00010, OCS Study BOEM 2020–022. US Department of the Interior, Bureau of Ocean Energy Management, New Orleans, Louisiana, 225 pp.
  3. Bernasconi, I. (1956) Dos nuevos Equinodermos de la costa del Brasil. Neotropica, 2, 33–36, 2 figs.
  4. Bernasconi, I. (1958) Equinoideos y asteroide de la colección del Instituto Oceanográfico de la Univeridad de San Pablo. 2. Boletin Instituto Oceanográfico de Sao Paulo, 7, 119–149. https://doi.org/10.1590/S0373-55241958000100002
  5. Bernasconi, I. (1961) Una nueva especie de asteroideo. Neotropica, 7 (22), 1–2, 1 fig.
  6. Bernasconi, I. (1963) Asteroideos argentinos. 4. Familia Goniasteridae. Revista Museo Argentino de Ciencias Naturales Bernardino Rivadavia. Zoologia, 9 (1), 1–25, pls. 1–5.
  7. Bernasconi, I. (1964) Asteroideos argentinos. Claves para los órdenes, familias, subfamilies y géneros. Physis Buenos Aires, 24, 241–277.
  8. Bernasconi, I. (1970) Equinodermos Antarticos. II. Asteroideos. 3. Asteroideos de la extremidad Norte de la Peninsula Antarctica. Revista del Museo Argentino de Ciencias Naturales “Bernardino Rivadavia” e Instituto Nacional de Investigación de las Ciencias Naturales, 9 (10), 211–281, 20 pls.
  9. Birkeland, C. (1974) Interactions between a sea pen and seven of its predators. Ecological Monographs, 44, 211–232. https://doi.org/10.2307/1942312
  10. Blacker, R.W. (1957) Benthic animals as indicators of hydrographic conditions and climatic change in Svalbard waters. Fisheries Investigations of the Ministry of Agriculture of London, 20, 1–49, 30 figures.
  11. Branch, M.L., Jangoux, M., Alva, V., Massin, C.L. & Stampanato, S. (1993) The Echinodermata of subantarctic Marion and Prince Edward Islands. South African Journal of Antarctic Research, 23, 1(2), 37−70.
  12. Clark, A.H. (1916) A new starfish (Lydiaster americanus) from the Gulf of Mexico. Journal of the Washington Academy of Sciences, 6, 141–144.
  13. Clark, A.H. (1954) Echinoderms (other than Holothurians) of the Gulf of Mexico. Fisheries Bulletin, U.S. Fisheries & Wildlife Service, 55 (89), 373–379.
  14. Clark, A.M. (1993) An index of names of recent Asteroidea. Part 2: Valvatida. Echinoderm Studies, 4, 187–366. https://doi.org/10.1201/9781003072553-4
  15. Clark, A.M. & Courtman-Stock, J. (1976) The Echinoderms of Southern Africa. British Museum of Natural History, London, 277 pp.
  16. Clark, A.M. & Downey, M.E. (1992) Starfishes of the Atlantic. Chapman and Hall, London, 794 pp.
  17. Clark, H.E.S. & McKnight, D.G. (2001) The Marine Fauna of New Zealand: Echinodermata: Asteroidea (sea-stars), Order Valvatida. NIWA Biodiversity Memoir, 117, 1–270.
  18. Clark, H.L. (1941) Reports on the scientific results of the Atlantis expeditions to the West Indies, under the joint auspices of the University of Havana and Harvard University. The Echinoderms (other than holothurians). Memorias de la Sociedad Cubana de Historia Natural, 15 (1), 1–154, 10 pls.
  19. Clark, M.R. & Bowden, D.A. (2015) Seamount biodiversity: high variability both within and between seamounts in the Ross Sea region of Antarctica. Hydrobiologia, 761, 161–180. https://doi.org/10.1007/s10750-015-2327-9
  20. Danielssen, D.C. & Koren, J. (1882) Fra den norske Nordhavsexpedition, Echinodermer. Nyt Magazin for Naturvidenskaberne, 27, 267–299.
  21. Dilman, A.B. (2013) Changes in deep-sea asteroid fauna along the northern Mid-Atlantic Ridge. Cahiers Biologie Marine, 54, 505–509. https://doi.org/10.1080/17451000.2012.749993
  22. Djakonov, A.M. (1950) n.k. Morskie Zvezdy Morei SSSR T U, 34, 1–203. [translated as Dyakonov, A.M. (1968) In: Strelkov, A.A. (Ed.), Sea stars (Asteroids) of the USSR Seas. Keys to the Fauna of the USSR. Vol. 34. Zoological Institute of the Academy of Sciences of the USSR, Israel Program for scientific translations Ltd., Jerusalem, pp. 1–183.]
  23. Downey, M.E. (1971) A new species of the genus Solaster (Echinodermata: Asteroidea) from Martinique. Proceedings of the Biological Society of Washington, 84, 421–426.
  24. Downey, M.E. (1973) Starfishes from the Caribbean and the Gulf of Mexico. Smithsonian Contributions to Zoology, 126, 1–158. https://doi.org/10.5479/si.00810282.126
  25. Downey, M.E. (1980) Floriaster maya, new genus and species of the family Goniasteridae (Echinodermata: Asteroidea). Proceedings of the Biological Society of Washington, 93, 346–349.
  26. Downey, M.E. (1982) Evoplosoma virgo, a new goniasterid starfish (Echinodermata: Asteroidea) from the Gulf of Mexico. Proceedings of the Biological Society of Washington, 95 (4), 772–773.
  27. Elliott, K.P., Kennedy, B.R.C., Lobecker, M. & Potter, J. (2016) Okeanos Explorer 2015 Field Season Overview, New Frontiers in Ocean Exploration: The E/V Nautilus and NOAA Ship Okeanos Explorer 2015 Field Season. Oceanography (supplement), 29 (1), 58–61. https://doi.org/10.5670/oceanog.2016.supplement.01
  28. Fisher, W.K. (1906) The starfishes of the Hawaiian islands. Bulletin of the United States Fish Commission, 23, 987–1130.
  29. Fisher, W.K. (1908) Necessary changes in the nomenclature of starfishes. Smithsonian Miscellaneous collections (Quarterly), 52, 87–93.
  30. Fisher, W.K. (1911) Asteroidea of the North Pacific and adjacent waters. 1. Phanerozonia and Spinulosida. Bulletin of the US National Museum, 76, 1–420. https://doi.org/10.5479/si.03629236.76.i
  31. Fisher, W.K. (1919) Starfishes of the Philippine seas and adjacent waters. Bulletin of the United States National Museum, 3 (100), 1–547. 156 pls.
  32. Fisher, W.K. (1930) Asteroidea of the North Pacific and adjacent waters. 3. Forcipulata (concluded). Bulletin of the US National Museum, 78, 1–356, 93 pls. https://doi.org/10.5479/si.03629236.76.3
  33. Florez, C. & Martinez de Rodriguez, A. (1971) Status de la familia Goniasteridae (Asteroidea: Phanerozonia) en la region oriental de Venezuela. Boletim Instituto Oceanografico, 9, 3–8, 6 figs.
  34. Forbes, E. (1839) On the Asteriadae of the Irish Sea. Memoirs of the Wernerian Natural History Society of Edinburgh, 8, 114–129.
  35. France, S.C., Amon, D.J., Messing, C., Skarke, A., Wagner, D., White, M.P., Kennedy, B.R.C. & Pawlenko, N. (2019) Deep-Sea Exploration of the US Gulf of Mexico with NOAA Ship Okeanos Explorer, Supplement: New Frontiers in Ocean Exploration: The E/V Nautilus, NOAA Ship Okeanos Explorer, and R/V Falkor 2018 Field Season. Oceanography, 32 (1), 78–81. https://doi.org/10.5670/oceanog.2019.supplement.01
  36. Gale, K.S.P., Hamel, J.-F. & Mercier, A. (2013) Trophic ecology of deep-sea Asteroidea (Echinodermata) from eastern Canada. Deep-Sea Research, Part I, 80, 25–36. https://doi.org/10.1016/j.dsr.2013.05.016
  37. García-Guillén, L.M. , Macías-Ramírez, A., Ríos, P., Manjón-Cabeza, M.E. (2022) Deep sea starfishes (Echinodermata: Asteroidea) from the Avilés Canyon System (Bay of Biscay), including two new records. Estuarine, Coastal and Shelf Science, 277, 1–14. https://doi.org/10.1016/j.ecss.2022.107993
  38. Gebruk, A., Budaeva, N. & King, N. (2010) Bathyal benthic fauna of the Mid-Atlantic Ridge between the Azores and the Reykjanes Ridge. Journal of the Marine Biological Association of the United Kingdom, 90 (1), 1–14. https://doi.org/10.1017/S0025315409991111
  39. Gray, J.E. (1840) XXXII. A synopsis of the genera and species of the class Hypostoma (Asterias, Linnaeus). Annals of the Magazine of Natural History, Series 1, 6, 175–184 + 275–290. https://doi.org/10.1080/03745484009443296
  40. Halpern, J.A. (1970a) A monographic revision of the goniasterid sea stars of the North Atlantic. Unpublished PhD dissertation, University of Miami, Coral Gables, Florida, 253 pp.
  41. Halpern, J.A. (1970b) Goniasteridae (Echinodermata: Asteroidea) of the Straits of Florida. Bulletin of Marine Science, 20 (1), 193–286.
  42. Harvey, R., Gage, J.D., Billett, D.S.M., Clark, A.M. & Paterson, L.J. (1988) Echinoderms of the Rockall Trough and adjacent areas. 3. Additional records. Bulletin of the British Museum of Natural History, Zoology, 54 (4), 153–198, 5 figs.
  43. Hayashi, R. (1939) Solasterids in Japanese waters. Journal of the Faculty of Science of Hokkaido Imperial University, Zoology, 6, 297–311.
  44. Hayashi, R. (1940) Contributions to the Classification of the sea-stars of Japan. I. Spinulosa. Journal of the Faculty of Imperial Science of Hokkaido University, 6 (7), 107–204.
  45. Howell, K.L., Rogers, A.D., Tyler, P.A. & Billett, D.S.M. (2004) Reproductive isolation among morphotypes of the Atlantic sea star species Zoroaster fulgens (Asteroidea: Echinodermata). Marine Biology, 144, 977–984. https://doi.org/10.1007/s00227-003-1248-8
  46. Jangoux, M. (1982) Food and feeding mechanisms: Asteroidea. In: Jangoux, M. & Lawrence, J. (Eds.), Echinoderm Nutrition. Balkema, Rotterdam, pp. 117–159. https://doi.org/10.1201/9781003078920-6
  47. Jangoux, M. (1982b) On Tremaster Verrill 1878, an odd genus of Recent starfish (Echinodermata: Asteroidea). In: Lawrence, J.M. (Ed.), Echinoderms, Proceedings of the International Echinoderm Conference, Tampa Bay 1981. AA Balkema, Rotterdam, pp. 155–163.
  48. John, D.D., Clark, A.M. (1954) The Rosaura Expedition. 3. The Echinodermata. Bulletin of the British Museum of Natural History, Zoology, 2, 139–162, 1 pl., 12 figs. https://doi.org/10.5962/p.314149
  49. Kennedy, B.R.C., Cantwell, K., Sowers, D., Quattrini, A.M., Cheadle, M.J. & McKenna, L. (2015) EX1502L3 Expedition Report- Océano Profundo 2015: Exploring Puerto Rico’s Seamounts, Trenches, and Troughs. OER Expedition Rep. 2015-02-03. Office of Ocean Exploration and Research, Office of Oceanic &Atmospheric Research, NOAA, Silver Spring, Maryland, 93 pp. https://doi.org/10.7289/V5NG4NM8
  50. Koehler, R. (1909) Astéries recueillies par l’Investigator dans l’océan Indien. I. Les Asteries de Mer Profonde. Echinoderma of the Indian Museum, 5, 1–143, 13 pls. https://doi.org/10.5038/bin.books.1017
  51. Leeling, B. (1984) Zur Morphologie und Systematik der Gattung Tremaster Verrill, 1879 (Echinodermata, Asteroidea) [Translation: The morphology and systematics of the genus Tremaster Verrill, 1879]. Mitteilungen aus dem Naturhistorischen Museum in Hamburg, 81, 261–276.
  52. Linchangco, G.V., Foltz, D.W., Reid, R., Williams, J., Nodzak, C., Kerr, A.M., Miller, A.K., Hunter, R., Wilson, N.G., Nielsen, W.J., Mah, C.L., Rouse, G.W., Wray, G.A. & Janies, D.A. (2017) The phylogeny of extant starfish (Asteroidea: Echinodermta) including Xyloplax based on comparative transcriptomics. Molecular Phylogenetics and Evolution, 115, 161–170. https://doi.org/10.1016/j.ympev.2017.07.022
  53. Madeira, P., Kroh, A., Cordeiro, R., De Frias Martins, A. & Avila, S.P. (2019) The Echinoderm Fauna of the Azores (NE Atlantic Ocean). Zootaxa, 4639 (1), 1–231. https://doi.org/10.11646/zootaxa.4639.1.1
  54. Mah, C.L. (2006) Phylogeny and biogeography of the deep-sea goniasterid, Circeaster (Echinodermata: Asteroidea) including descriptions of six new species. Zoosystema, 28 (4), 917–954.
  55. Mah, C.L. (2007) Phylogeny of the Zoroasteridae (Zorocallina; Forcipulatida): Evolutionary Events in Deep-Sea Asteroidea displaying Paleozoic Features. Zoological Journal of the Linnean Society, 150, 177–210. https://doi.org/10.1111/j.1096-3642.2007.00291.x
  56. Mah, C.L. (2011) Revision and Key to High-Latitude (Antarctic and subAntarctic) Goniasteridae. Zootaxa, 2759 (1), 1–48. https://doi.org/10.11646/zootaxa.2759.1.1
  57. Mah, C.L. (2015) A new Atlantic species of Evoplosoma with taxonomic summary and in situ observations of Atlantic deep-sea corallivorous Goniasteridae (Valvatida; Asteroidea). Marine Biodiversity Records, 8 (e5), 1–8. https://doi.org/10.1017/S1755267214001407
  58. Mah, C.L. (2016) Deep-sea (>1000 m) Goniasteridae (Valvatida; Asteroidea) from the North Pacific, including an overview of Sibogaster, Bathyceramaster n. gen. and three new species. Zootaxa, 4175 (2), 101–141. https://doi.org/10.11646/zootaxa.4175.2.1
  59. Mah, C.L. (2018) New genera, species and occurrence records of Goniasteridae (Asteroidea; Echinodermata) from the Indian Ocean. Zootaxa, 4539 (1), 1–116. https://doi.org/10.11646/zootaxa.4539.1.1
  60. Mah, C.L. (2020) New Species, Occurrence Records and Observations of Predation by Deep-sea Asteroidea (Echinodermata) from the North Atlantic by NOAA Ship Okeanos nExplorer. Zootaxa, 4766 (2), 201–260. https://doi.org/10.11646/zootaxa.4766.2.1
  61. Mah, C.L. (2021) The East Pacific/South Pacific Boundary: New Taxa and Occurrences from Rapa Nui (Easter Island), New Caledonia and adjacent regions. Zootaxa, 4980 (3), 401–450. https://doi.org/10.11646/zootaxa.4980.3.1
  62. Mah, C.L. (2022) New Genera, Species and Occurrence of Deep-Sea Asteroidea (Valvatacea, Forcipulatacea, Echinodermata) collected from the Tropical Pacific Ocean by the CAPSTONE Expedition. Zootaxa, 5164 (1), 1–75. https://doi.org/10.11646/zootaxa.5164.1.1
  63. Mah, C.L. (2023a) World Asteroidea Database. Anthenoides peircei Perrier, 1881. Available from: https://marinespecies.org/asteroidea./aphia.php?p=taxdetails&id=178061 (accessed 26 October 2023)
  64. Mah, C.L. (2023b) New Genera, Species, and observations on the biology of Antarctic Valvatida (Asteroidea). Zootaxa, 5310 (1), 1–88. https://doi.org/10.11646/zootaxa.5310.1.1
  65. Mah, C. & Blake, D.B. (2012) Global Diversity and Phylogeny of the Asteroidea (Echinodermata). PLoS ONE, 7 (4), e35644. https://doi.org/10.1371/journal.pone.0035644
  66. Mah, C.L. & Foltz, D.W. (2011a) Molecular Phylogeny of the Valvatacea (Asteroidea, Echinodermata). Zoological Journal of the Linnean Society, 161, 769–788. https://doi.org/10.1111/j.1096-3642.2010.00659.x
  67. Mah, C. & Foltz, D.W. (2011b) Molecular Phylogeny of the Forcipulatacea (Asteroidea: Echinodermata): Systematics & Biogeography. Zoological Journal of the Linnean Society, 162, 646–660. https://doi.org/10.1111/j.1096-3642.2010.00659.x
  68. Mah, C.L. & Foltz, D.W. (2014) New Taxa and Taxonomic Revisions to the Poraniidae (Valvatacea; Asteroidea) with Comments on Feeding Biology. Zootaxa, 3795 (3), 327–372. https://doi.org/10.11646/zootaxa.3795.3.7
  69. Mah, C.L., Nizinski, M. & Lundsten, L. (2010) Phylogenetic Revision of the Hippasterinae (Goniasteridae; Asteroidea): Systematics of Deep Sea Corallivores, including one new genus and three new species. Zoological Journal of the Linnean Society, 160, 266–301. https://doi.org/10.1111/j.1096-3642.2010.00638.x
  70. Mauzey, K.P., Birkeland, C. & Dayton, P.K. (1968) Feeding behavior of asteroids and escape responses of their prey in the Puget Sound region. Ecology, 49, 603–619. https://doi.org/10.2307/1935526
  71. Meadows, P.S., Meadows, A. & Murray, J.M.H. (2012) Biological modifiers of marine benthic seascapes: Their role as ecosystem engineers. Geomorphology, 157–158, 31–48. https://doi.org/10.1016/j.geomorph.2011.07.007
  72. Mecho, A., Easton, E.E., Sellanes, J., Gorny, M. & Mah, C. (2019) Unexplored diversity of the mesophotic echinoderm fauna of the Easter Island ecoregion. Marine Biology, 166, 91. https://doi.org/10.1007/s00227-019-3537-x
  73. Milne-Edwards, A. (1882) Rapport sur les Travaux de la Commission chargée par M. le Ministre de l’Instruction Publique d’étudier la faune sous-marine dans les grandes profondeurs de la Méditerranée et de l’Océan Atlantique. Archives des missions scientifiques et littéraires, 9 (3), 1–59.
  74. Mortensen, T. (1927) Handbook of the Echinoderms of the British Isles. Oxford University Press, London, ix + 471 pp., 269 figs. https://doi.org/10.5962/bhl.title.6841
  75. Nance, J.M. & Braithwaite, L.F. (1979) The function of mucous secretions in the cushion star Pteraster tesselatus Ives. Journal of Experimental Marine Biology and Ecology, 40 (3), 259–266. https://doi.org/10.1016/0022-0981(79)90055-8
  76. Nesis, K.N. (1960) On the distribution of the starfish Tremaster mirabilis Verrill. Zoologiechesky Zhurnal, 39, 1886–1888, 1 fig.
  77. O’Loughlin, P.M. & Waters, J.W. (2004) A molecular and morphological revision of genera of Asterinidae (Echinodermata: Asteroidea). Memoirs of Museum Victoria, 61 (1), 1–40. https://doi.org/10.24199/j.mmv.2004.61.1
  78. Pawson, D.L. (1976) Some aspects of the biology of deep-sea echinoderms. Thalassia Jugoslavica, 12 (1), 287–293.
  79. Perrier, E. (1881) Report on the Results of dredging in the Gulf of Mexico and in the Caribbean Sea, 1877–79, by the United States Coastal Survey Steamer Blake. 14. Description sommaire des espèces nouvelles d’Astéries. Bulletin of the Museum of Comparative Zoology, 9, 1–31.
  80. Perrier, E. (1884) Mémoire sur les étoiles de mer recueillies dans la mer des Antilles et le golfe du Mexique : durant les expéditions de dragage faites sous la direction de M. Alexandre Agassiz. Nouvelles archives du Muséum national d’histoire naturelle de Paris, 6 (2), 127–276. https://doi.org/10.5962/bhl.title.82184
  81. Perrier, E. (1885) Sur les stellérides recueillis durant la mission du Talisman. Comptes-rendus Hebdomadaires des Séances de l’Académie des Sciences, Paris, 101, 884–887.
  82. Perrier, E. (1891) Stellerides nouveaux provenant des campagnes du yacht l’Hirondelle. Mémoires de la Société zoologique de France, 4, 258–271.
  83. Perrier, E. (1894) Stellérides. Expéditions Scientifiques du Travailleur et du Talisman. Masson, Paris, 431 pp., 26 pls.
  84. Ringvold, H. (1999) Starfishes (Echinodermata, Asteroidea) new for the Faroe Islands, collected by “BIOFAR.” Sarsia, 84 (5–6), 469–472. https://doi.org/10.1080/00364827.1999.10807353
  85. Ringvold, H. & Andersen, T. (2016) Starfish (Asteroidea, Echinodermata) from the Faroe Islands; spatial distribution and abundance. Deep Sea Research Part I: Oceanographic Research Papers, 107, 22–30. https://doi.org/10.1016/j.dsr.2015.09.004
  86. Sladen, W.P. (1889) Report on the Asteroidea. Report on the Scientific Results of the Voyage of H.M.S. Challenger during the years 1873–1876. Zoology, 30 (51), 1–893, 118 pls.
  87. Spencer, W.K. & Wright, C.R. (1966) Asterozoans, Part U: Echinodermata In: Moore, R.C. (Ed.), Treatise on Invertebrate Paleontology. 3 (1). University of Kansas Press, Lawrence, Kansas, U4–U107.
  88. Stampanato, S. & Jangoux, M. (2004) The asteroid fauna (Echinodermata) of Marion and Prince Edward Islands. Annals of the South African Museum, 112 (1), 1–16.
  89. Thomson, C.W. (1873) The Depth of the Sea. An Account of the General Results of the Dredging Cruises of the H.M.S.S. ‘Porcupine’ and ‘Lightning’ During the Summers of 1868, 1869, and 1870, Under the Scientific Direction of Dr. Carpenter, F.R.S., J. Gwyn Jeffreys, F.R.S. and Dr. Wyville Thomson, F.R.S. Macmillan and Co., London, 527 pp., pls. I–VIII. https://doi.org/10.5962/bhl.title.19314
  90. Thomson, C.W. & Murray, J. (1885) Report on the scientific results of the voyage of H.M.S. Challenger during the years 1873–76 under the command of Captain George S. Nares and the late Captain Frank Tourle Thomson, R.N. Vol. 1. Part 2. Neill, Edinburgh, pp. i–viii + 511–1110. https://doi.org/10.5962/bhl.title.109677
  91. Tortonese, E. & Clark, A.M. (1956) On the generic position of the asteroid Goniodiscus placenta Müller and Troschel. Annals of the Magazine of Natural History, Series 12, 9 (101), 347–352. https://doi.org/10.1080/00222935608655821
  92. Tortonese, E. (1984) Notes on the Mediterranean sea-star Peltaster placenta (M& T.). Bollettino Museo civico di Storia Naturale di Verona, 11, 99–112.
  93. VanVeldhuizen, H.D. & Oakes, V.J. (1981) Behavioral responses of seven species of asteroids to the asteroid predator, Solaster dawsoni (responses of asteroids to the predator Solaster dawsoni). Oecologia, 48 (2), 214–220. https://doi.org/10.1007/BF00347967
  94. Verrill, A.E. (1879) Notice of recent additions to the marine invertebrata of the northeastern coast of America, with descriptions of new genera and species and critical remarks on others. Part I. Annelida, Gephyraea, Nemertina, Nematoda, Polyzoa, Tunicata, Mollusca, Anthozoa, Echinodermata, Porifera. Proceedings of the United States National Museum, 2, 165–205. https://doi.org/10.5962/bhl.title.14749
  95. Verrill, A.E. (1884) Notice of the remarkable marine fauna occupying the outer banks off the southern coast of New England, 9. Brief Contributions to Zoology from the Museum of Yale College. American Journal of Science, 28, 213–220. https://doi.org/10.2475/ajs.s3-28.167.377
  96. Verrill, A.E. (1885) Results of the explorations made by the steamer Albatross off the northern coast of the United States in 1883. Reports of the US Fisheries Commission, 1883, 503–645, pls. 10–21. https://doi.org/10.5962/bhl.title.12059
  97. Verrill, A.E. (1894) Descriptions of new species of starfishes and ophiurans, with a revision of certain species formerly described; mostly from the collections made by the United States Commission of Fish and Fisheries. Proceedings of the United States National Museum, 17 (1000), 245–297. https://doi.org/10.5479/si.00963801.1000.245
  98. Verrill, A.E. (1895) Distribution of the echinoderms of north–eastern America. American Journal of Science, 3 (49), 127–141. https://doi.org/10.2475/ajs.s3-49.290.127
  99. Verrill, A.E. (1899) Revision of certain genera and species of starfishes, with descriptions of new forms. Transactions of the Connecticut Academy of Arts and Sciences, 10 (1), 145–234. https://doi.org/10.5962/bhl.part.7031
  100. Verrill, A.E. (1915) Report on the starfishes of the West Indies, Florida, and Brazil, including those obtained by the Bahama expedition from the University of Iowa in 1893. Bulletins from the Laboratories of Natural History, 7 (1), 1–232, 29 pls. https://doi.org/10.5962/bhl.title.12035
  101. Walenkamp, J.H.C. (1976) The asteroids of the coastal waters of Surinam. Zoologische Verhandelingen, 147, 1–91.
  102. Walenkamp, J.H.C. (1979) Asteroidea (Echinodermata) from the Guyana Shelf. Zoologische Verhandelingen, 170, 1–97.