Skip to main content Skip to main navigation menu Skip to site footer
Responsive image
Issue: Vol. 5529 No. 3: 29 Oct. 2024
Type: Article
Published: 2024-10-29
DOI: 10.11646/zootaxa.5529.3.7
Page range: 551-569
Abstract views: 101
PDF downloaded: 8

New records of the solitary Vespidae (Hymenoptera: Vespidae: Masarinae and Eumeninae s. l.) from Central Asia, with new synonyms and description of a new species of Eumenes Latreille

T.I. Vyazemsky Karadag Scientific Station—Nature Reserve of RAS—Branch of A.O. Kovalevsky Institute of Biology of the Southern Seas of RAS; Nauki Str. 24; Kurortnoye; 298188 Feodosiya; Russia.
Federal Scientific Center of the East Asia Terrestrial Biodiversity; Far Eastern Branch of the Russian Academy of Sciences; 100-let Vladivostoku Ave. 159; 690022 Vladivostok; Russia
Hymenoptera pollen wasps eumenine wasps fauna taxonomy new synonymy trophic relationships Palaearctic region

Abstract

A new species, Eumenes selisi Fateryga, sp. nov. is described from Uzbekistan. Three new synonymies are proposed: Masaris carli von Schulthess, 1922 = M. elegans Gusenleitner, 2002, syn. nov.; Parodontodynerus ephippium (Klug, 1817) = P. ephippium dalanicus Kurzenko, 1976, syn. nov.; Psiliglossa odyneroides (S.S. Saunders, 1850) = P. odyneroides kozhantshikovi Kostylev, 1940, syn. nov. Twenty-nine new country records are presented: two from Turkmenistan, four from Tajikistan, seven from Uzbekistan, seven from Kyrgyzstan, and nine from Kazakhstan. Masaris carli is excluded from the fauna of Iran and Psiliglossa odyneroides is excluded from the fauna of Russia. Observations on flower-visiting behavior of Chlorodynerus arenicola and Euodynerus setosus are reported.

 

References

  1. André, E. (1881–1886) Species des Hyménoptères d’Europe et d’Algérie. Tome 2. André, Beaune, 919 + 48 pp. https://doi.org/10.5962/bhl.title.10281
  2. André, E. (1883) Description de quelques espèces nouvelles d’Odynères de la Sicile. Il Naturalista Siciliano, 2 (10), 229–234.
  3. Antropov, A.V. & Fateryga, A.V. (2017) Family Vespidae. In: Lelej, A.S., Proshchalykin, M.Yu. & Loktionov, V.M. (Eds.), Annotated catalogue of the Hymenoptera of Russia. Vol. 1. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute RAS, Supplement 6, pp. 175–196. https://doi.org/10.31610/trudyzin/2017.supl.6.5
  4. Bank, S., Sann, M., Mayer, C., Meusemann, K., Donath, A., Podsiadlowski, L., Kozlov, A., Petersen, M., Krogmann, L., Meier, R., Rosa, P., Schmitt, T., Wurdack, M., Liu, S., Zhou, X., Misof, B., Peters, R.S. & Niehuis, O. (2017) Transcriptome and target DNA enrichment sequence data provide new insights into the phylogeny of vespid wasps (Hymenoptera: Aculeata: Vespidae). Molecular Phylogenetics and Evolution, 116, 213–226. https://doi.org/10.1016/j.ympev.2017.08.020
  5. Barlow, S.E. & Pavlik, B.M. (2017) Estimating the spatial dimensions of Astragalus holmgreniorum mutualisms. I. Pollinator visitation and behaviors along a seed set gradient. Report to the State of Utah Department of Natural Resources, Endangered Species Recovery Program, and the Nature Conservancy. Red Butte Garden and Arboretum & Natural History Museum of Utah, Salt Lake City, 25 pp.
  6. Blüthgen, P. (1938) Beiträge zur Kenntnis der paläarktischen Eumeniden (Hym. Vespidae). Deutsche Entomologische Zeitschrift, 1938 (2), 434–495. https://doi.org/10.1002/mmnd.193819380208
  7. Blüthgen, P. (1942) Neue paläarktische Faltenwespen (Hym. Vespidae Eumenidinae). Mitteilungen aus dem Zoologischen Museum in Berlin, 25, 300–322.
  8. Blüthgen, P. (1951) Neue oder bemerkenswerte paläarktische Faltenwespen aus der Zoologischen Staatssammlung in München (Hym. Vespidae, Eumenidinae). Mitteilungen der Münchner Entomologischen Gesellschaft, 41, 166–201.
  9. Blüthgen, P. (1952) Weitere neue oder bemerkenswerte paläarktische Faltenwespen aus der Zoologischen Staatssammlung in München (Hym., Eumenidae, Masaridae). Mitteilungen der Münchner Entomologischen Gesellschaft, 42, 1–19.
  10. Blüthgen, P. (1953) Die Eumeniden-Gattung Allodynerus Blüthg. 1938 (Hym., Vespidae). Zoologischer Anzeiger, 150, 50–59.
  11. Blüthgen, P. (1956) Portuguese and Spanish wasps (Hymenoptera, Vespoidea). 1st Supplement. Memórias e Estudos do Museu Zoológico da Universidade de Coimbra, 240, 1–22.
  12. Blüthgen, P. & Gusenleitner, J. (1970) Über Faltenwespen aus dem Iran (Hym., Diploptera). Stuttgarter Beiträge zur Naturkunde, 223, 1–13.
  13. Brauns, H. (1905) Masaridae von Südafrika. Annales Historico-Naturales Musei Nationalis Hungarici, 3, 219–234.
  14. Carl, J. (1921) Une espèce nouvelle de Masaris (Vespidae). Revue Suisse de Zoologie, 28 (19), 449–451. https://doi.org/10.5962/bhl.part.84690
  15. Carpenter, J.M. (2001) Checklist of species of the subfamily Masarinae (Hymenoptera: Vespidae). American Museum Novitates, 3325, 1–39. https://doi.org/10.1206/0003-0082(2001)325<0001:COSOTS>2.0.CO;2
  16. Castro, L. (1992) Sobre los Euménidos (Hym., Vespoidea) del valle medio del Ebro. ZAPATERI. Revista Aragonesa de Entomología, 1 (2), 21–34.
  17. Castro, L. (1997) Familia Vespidae: Subfamilia Eumeninae. Catalogus de la Entomofauna Aragonesa, 16, 3–8.
  18. Christ, J.L. (1791) Naturgeschichte, Klassification und Nomenclatur der Insekten vom Bienen, Wespen und Ameisengeschlecht; als der fünften Klasse fünfte Ordnung des Linneischen Natursystems von den Insekten: Hymenoptera. Mit häutigen Flügeln. Hermannischen Buchhandlung, Frankfurt am Main, 535 pp. https://doi.org/10.5962/bhl.title.87724
  19. Curtis, J. (1823–1840) British entomology: being illustrations and descriptions of the genera of insects found in Great Britain and Ireland: containing coloured figures from nature of the most rare and beautiful species, and in many instances of the plants upon which they are found. Vol. 4. Curtis, London, 70 pls., unnumbered text pages. https://doi.org/10.5962/t.171559
  20. Curtis, J., Haliday, A.H. & Walker, F. (1836) Descriptions, &c. of the insects collected by captain P.P. King, R.N., F.R.S., in the survey of the Straits of Magellan. Transactions of the Linnean Society of London, 17 (3), 315–359. https://doi.org/10.1111/j.1095-8339.1834.tb00026.x
  21. Dalla Torre, K.W. von (1889) Hymenopterologische Notizen. Wiener Entomologische Zeitung, 8, 124–125.
  22. Dalla Torre, K.W. von (1894) Catalogus hymenopterorum hucusque descriptorum et systematicus et synonymicus. Vol. 9. Vespidae (Diploptera). Engelmann, Leipzig, viii + 181 pp. https://doi.org/10.5962/bhl.title.10348
  23. Dalla Torre, K.W. von (1904) Hymenoptera. Fam. Vespidae. In: Witsman, P. (Ed.), Genera Insectorum. Vol. 19. Desmet-Verteneuil, Bruxelles, pp. 1–108, 6 pls.
  24. de Saussure, H.F. (1852–1853) Etudes sur la famille des Vespides. 1. Monographie des guepes solitaires ou de la tribu des Eumeniens, comprenant la classification et la description de toutes les especes connues jusqu’a ce jour, et servant de complement au manuel de Lepeletier de Saint-Fargeau. Cherbuliez & Masson, Genève and Paris, lii + 286 pp. + 22 pls. https://doi.org/10.5962/bhl.title.39973
  25. de Saussure, H.F. (1854–1856) Études sur la famille des Vespides. 3. Monographie des fausses guêpes, ou de la tribu des Masariens. Troisième partie comprenant la monographie des Masariens et un supplément à la monographie des Eumeniens. Masson, Kessmann & Cherbuliez, Paris and Genève, 352 pp., 16 pls. https://doi.org/10.5962/bhl.title.39973
  26. de Saussure, H.F. (1862) Sur divers Vespides Asiatiques et Africains du Musée de Leyden. Stettiner Entomologische Zeitung, 23, 177–207.
  27. de Saussure, H.F. (1863) Mélanges hyménoptérologiques. Mémoires de la Société de Physique et d’Histoire Naturelle de Genève, 17 (1), 171–244, pl. 2.
  28. Dusmet y Alonso, J.M. (1903) Véspidos, Euménidos y Masáridos de España. Memorias de la Real Sociedad Española de Historia Natural, 2, 119–225.
  29. Fateryga, A.V. (2024) On some new and little-known Palaearctic species of the genus Leptochilus de Saussure, 1853 (Hymenoptera: Vespidae: Eumeninae). Zootaxa, 5432 (1), 38–53. https://doi.org/10.11646/zootaxa.5432.1.3
  30. Fateryga, A.V. & Buyanjargal, B. (2023) A new species of the genus Onychopterocheilus Blüthgen, 1955 from the Altai Mountains, with the re-description and new records of the closely related O. kiritshenkoi (Kostylev, 1940) (Hymenoptera: Vespidae: Eumeninae). Zootaxa, 5323 (1), 94–104. https://doi.org/10.11646/zootaxa.5323.1.5
  31. Fateryga, A.V., Carpenter, J.M. & Fateryga, V.V. (2023a) Ancistrocerus capra (de Saussure, 1857), a valid species, not a synonym of A. antilope (Panzer, 1798) (Hymenoptera: Vespidae: Eumeninae). American Museum Novitates, 4002, 1–13. https://doi.org/10.1206/4002.1
  32. Fateryga, A.V., Mauss, V. & Proshchalykin, M.Yu. (2023b) Foraging behavior of two pollen wasp species of the genus Celonites Latreille, 1802 (Hymenoptera: Vespidae: Masarinae), from the Altai Mountains. Insects, 14 (5), 408. https://doi.org/10.3390/insects14050408
  33. Fateryga, A.V. & Mokrousov, M.V. (2019) New records of eumenine wasps (Hymenoptera: Vespidae: Eumeninae) from Russia with description of a new species of Leptochilus de Saussure, 1853. Zootaxa, 4612 (3), 412–422. https://doi.org/10.11646/zootaxa.4612.3.7
  34. Fateryga, A.V., Proshchalykin, M.Yu., Aliyev, Kh.A. & Maharramov, M.M. (2019) To the knowledge of eumenine wasps (Hymenoptera: Vespidae: Eumeninae) of Nakhchivan Autonomous Republic of Azerbaijan. Far Eastern Entomologist, 379, 25–32. https://doi.org/10.25221/fee.379.2
  35. Fateryga, A.V., Proshchalykin, M.Yu., Kochetkov, D.N. & Buyanjargal, B. (2020) New records of eumenine wasps (Hymenoptera, Vespidae, Eumeninae) from Russia, with description of a new species of Stenodynerus de Saussure, 1863. Journal of Hymenoptera Research, 79, 89–109. https://doi.org/10.3897/jhr.79.57887
  36. Fateryga, A.V., Proshchalykin, M.Yu., Maharramov, M.M., Astafurova, Yu.V. (2021) New records of solitary vespid wasps (Hymenoptera: Vespidae: Masarinae and Eumeninae s. l.) from the Nakhchivan Autonomous Republic of Azerbaijan. Zootaxa, 5027 (1), 36–60. https://doi.org/10.11646/zootaxa.5027.1.2
  37. Fateryga, A.V., Proshchalykin, M.Yu., Mokrousov, M.V. & Akhmedov, A.G. (2023c) To the knowledge of the solitary vespid wasps (Hymenoptera: Vespidae: Masarinae and Eumeninae s. l.) of Uzbekistan. Zootaxa, 5278 (3), 439–460. https://doi.org/10.11646/zootaxa.5278.3.2
  38. Germar, E.F. (1817) Reise nach Dalmatien und in das Gebiet von Ragusa. Brockhaus, Leipzig and Altenburg, xii + 323 pp. + 9 pls.
  39. Giordani Soika, A. (1943) Eumenini palearctici nuovi o poco noti (Hym.). Bollettino della Società Vineziana di Storia Naturale e del Museo Civico di Storia Naturale, 3 (1), 1–14.
  40. Giordani Soika, A. (1952a [1951]) Risultati della spedizione scientifica zoologica in Turchia del Museo Nazionale di Praha. Acta Entomologica Musei Nationalis Pragae, 27, 375–386.
  41. Giordani Soika, A. (1952b) Sulle caratteristiche biogeografiche della Palestina, Arabia ed Egitto con un contributo alla conoscenza degli Zethini ed Eumenini della Palestina. Bollettino della Società Vineziana di Storia Naturale e del Museo Civico di Storia Naturale, 6 (1), 5–62.
  42. Giordani Soika, A. (1953 [1952]) Hyménoptères récoltés par une Mission suisse au Maroc (1947). Eumeninae. Bulletin de la Société des Sciences Naturelles du Maroc, 32, 235–267.
  43. Giordani Soika, A. (1970) Contributo alla conoscenza degli Eumenidi del Medio Oriente. Missione Giordani Soika in Iran 1965, III. Bollettino del Museo Civico di Storia Naturale di Venezia, 20/21, 27–183.
  44. Giordani Soika, A. (1973) Designazione di lectotipi ed elenco dei tipi di Eumenidi, Vespidi e Masaridi da me descritti negli anni 1934–1960. Bollettino del Museo Civico di Storia Naturale di Venezia, 24, 7–58.
  45. Giraud, J. (1866) Mémoire sur les insectes qui habitent les tiges sèches de la Ronce. Annales de la Société Entomologique de France, Série 4, 6, 443–500.
  46. Guiglia, D. (1951) Nuovi Eumenes Italiani (Hymen. Eumenidae). Bollettino della Società Entomologica Italiana, 81 (1–2), 27–30.
  47. Gusenleitner, J. (1970 [1969]) Bemerkenswertes über Faltenwespen I. (Diploptera, Hymenopt.). Mitteilungen der Münchner Entomologischen Gesellschaft, 59, 163–167.
  48. Gusenleitner, J. (1972) Übersicht über die derzeit bekannten westpaläarktischen Arten der Gattung Eumenes Latr. (Hymenoptera: Vespoidea). Bollettino del Museo Civico di Storia Naturale di Venezia, 22/23, 67–117.
  49. Gusenleitner, J. (1973 [1971–1972]) Beiträge zur Kenntnis der Fauna Afghanistans (Sammelergebnisse von O. Jakeš 1963–64, D. Povolný 1965, D. Povolný & Fr. Tenora 1966, J. Šimek 1965–66, D. Povolný, J. Gaisler, Z. Šebek & Fr. Tenora 1967). Vespoidea, Hymenoptera. Acta Musei Moraviae. Scientiae Naturales, 56/57, 315–334.
  50. Gusenleitner, J. (1988) Bemerkenswertes über Faltenwespen IX (Hymenoptera, Vespoidea). Nachrichtenblatt der Bayerischen Entomologen, 37 (4), 107–113.
  51. Gusenleitner, J. (1992) Bemerkenswerte Funde von Masaridae in Griechenland und im Nahen Osten (Vespoidea, Hymenoptera). Linzer Biologische Beiträge, 24 (1), 59–63.
  52. Gusenleitner, J. (1994) Über Eumeniden aus dem Nahen Osten und Arabien sowie Beschreibung einer neuen Subspecies aus Zanzibar (Vespoidea, Hymenoptera). Linzer Biologische Beiträge, 26 (1), 307–324.
  53. Gusenleitner, J. (2000) Bestimmungstabellen mittel- und südeuropäischer Eumeniden (Vespoidea, Hymenoptera). Teil 14. Die Gattungsschlüssel und die bisher in däeser Reihe nicht behandelten Gattungen und Arten. Linzer Biologische Beiträge, 32 (1), 43–65.
  54. Gusenleitner, J. (2001) Über eine Sammlung von Eumeniden-Arten im Museum d’Histoire naturelle Neuchâtel, welche Ed. André 1883 und 1884 beschrieben hat (Hymenoptera, Vespoidea, Eumenidae). Linzer Biologische Beiträge, 33 (1), 211–215.
  55. Gusenleitner, J. (2002) Über die Arten der Gattung Masaris Fabricius 1793 aus Zentralasien (Hymenoptera, Vespoidea, Masaridae). Linzer Biologische Beiträge, 34 (1), 321–333.
  56. Gusenleitner, J. (2013) Die Gattungen der Eumeninae im Nahen Osten, in Nordafrika und in Arabien (Hymenoptera: Vespidae: Eumeninae). Linzer Biologische Beiträge, 45 (1), 5–107.
  57. Gussakovskij, V.V. (1935) Sphecodea und Vespodea von Tajikistan. Transactions of the Tajik Base of the Academy of Sciences of the USSR, 5, 409–467. [in Russian]
  58. Haeseler, V. (1975) Pterocheilus phaleratus (Hymenoptera: Vespoidea), ein Nektardieb an den Blüten von Lotus corniculatus (Fabales: Fabaceae). Entomologica Germanica, 1 (3–4), 213–221. https://doi.org/10.1127/entom.germ/1/1975/213
  59. Harris, M. (1782) An exposition of English insects including the several classes of Neuroptera, Hymenoptera, & Diptera, or bees, flies, & Libellulae. Exhibiting on 51 copper plates near 500 figures, accurately drawn & highly finished in colours, from nature. The whole minutely described, arranged, & named, according to the Linnean system, with remarks. The figures of a great number of moths not in the Aurelian collection. Formerly published by the same author and a plate with an explanation of colours, are likewise given in the work. White & Robson, London, 166 pp., pls. I–L, 4 unnumbered text pages. https://doi.org/10.5962/bhl.title.168994
  60. ICZN (1999) International code of zoological nomenclature. 4th Edition. International Trust for Zoological Nomenclature, London, xxix + 306 pp. https://doi.org/10.5962/bhl.title.50608
  61. Inouye, D.W. (1980) The terminology of flower larceny. Ecology, 61 (5), 1251–1253. https://doi.org/10.2307/1936841
  62. Kostylev, G. (1925) Zur Kenntnis der Gattung Masaris F. (Hym). Entomologische Mitteilungen, 14 (2), 150–153.
  63. Kostylev, G. (1935a [1934]) Neue und wenig bekannte Mittelasiatische Wespidenarten (Hym.). Archives du Muzée Zoologique de l’Université de Moscou, 1, 117–146. [in Russian and German]
  64. Kostylev, G. (1935b) Materialien zur Kenntnis der Masariden-Fauna der Paläarktis. Archives du Muzée Zoologique de l’Université de Moscou, 2, 85–116. [in Russian and German]
  65. Kostylev, G. (1940a) Espèces novivelles et peu connues de Vespides, d’Euménides et de Masarides paléarctiques (Hymenoptera) I. Bulletin de la Société des Naturalistes de Moscou. Section Biologique, 49 (3–4), 137–154.
  66. Kostylev, G. (1940b) Espèces novivelles et peu connues de Vespides, d’Euménides et de Masarides paléarctiques (Hymenoptera) II. Bulletin de la Société des Naturalistes de Moscou. Section Biologique, 49 (5–6), 24–42.
  67. Kurzenko, N.V. (1976) New species of vespoid solitary wasps (Hymenoptera, Eumenidae) in the fauna of the USSR. Entomologicheskoe Obozrenie, 55 (2), 434–437. [in Russian]
  68. Kurzenko, N.V. (1977) Eumenid wasps (Hymenoptera, Eumenidae) of the Mongolian People’s Republic and adjacent regions of China and Southern Siberia. In: Kerzhner, I.M. (Ed.), Insects of Mongolia. No. 5. Nauka, Leningrad, pp. 537–582. [in Russian]
  69. Kurzenko, N.V. (1978) Solitary wasps of the family Eumenidae (Hymenoptera, Vespoidea) in the fauna of the USSR. 1. Subfamilies Raphiglossinae and Discoeliinae. Zoologicheskii Zhurnal, 57 (6), 867–872. [in Russian]
  70. Kurzenko, N.V. (1982 [1981]) [Review of the genera of the vespid wasps of the family Eumenidae (Hymenoptera, Vespoidea) of the fauna of the USSR]. In: Lehr, P.A. (Ed.), [Hymenoptera of the Far East]. Far Eastern Center of the Academy of Sciences of USSR Press, Vladivostok, pp. 81–112. [in Russian]
  71. Li, T.-J., Bai, Y. & Chen, B. (2022) A revision of the genus Jucancistrocerus Blüthgen, 1938 from China, with review of three related genera (Hymenoptera: Vespidae: Eumeninae). Zootaxa, 5105 (3), 401–420. https://doi.org/10.11646/zootaxa.5105.3.4
  72. Luo, L., Carpenter, J.M., Chen, B. & Li, T.-J. (2022) First comprehensive analysis of both mitochondrial characteristics and mitogenome-based phylogenetics in the subfamily Eumeninae (Hymenoptera: Vespidae). Insects, 13 (6), 529. https://doi.org/10.3390/insects13060529
  73. Mauss, V., Kuba, K. & Krenn, H.W. (2019) Evolution of the multifunctional mouthparts of adult Vespidae. In: Krenn, H.W. (Ed.), Insect Mouthparts. Zoological Monographs 5. Springer, Cham, pp. 443–478. https://doi.org/10.1007/978-3-030-29654-4_14
  74. Mauss, V. & Müller, A. (2014) First contribution to the bionomics of the pollen wasp Celonites fischeri Spinola, 1838 (Hymenoptera, Vespidae, Masarinae) in Cyprus. Journal of Hymenoptera Research, 39, 119–153. https://doi.org/10.3897/JHR.39.7841
  75. Morawitz, F. (1867) Übersicht der im Gouvernement von Saratow und um St. Petersburg vorkommenden Odynerus-Arten. Horae Societatis Entomologicae Rossicae, 4 (3–4), 109–144.
  76. Morawitz, F. (1885) Eumenidarum species novae. Horae Societatis Entomologicae Rossicae, 19 (1–2), 135–181.
  77. Morawitz, F. (1888) Hymenoptera aculeata nova. Horae Societatis Entomologicae Rossicae, 22 (3–4), 224–302.
  78. Morawitz, F. (1895) Materialien zu einer Vespidenfauna des Russischen Reiches. Horae Societatis Entomologicae Rossicae, 29 (3–4), 407–493.
  79. Oehlke, J. (2012) Beitrag zur Kenntnis der Faltenwespen-Fauna von Kyrgyzstan (Hymenoptera, Vespidae & Eumenidae). Linzer Biologische Beiträge, 44 (1), 595–600.
  80. Panzer, G.W.F. (1798) Fauna insectorum Germanicae initia, oder, Deutschlands Insecten. Heft 53. Felseckerschen Buchhandlung, Nürnberg, 24 pls., unnumbered text pages. https://doi.org/10.5962/bhl.title.15007
  81. Piekarski, P.K., Carpenter, J.M., Lemmon, A.R., Lemmon, E.M. & Sharanowski, B.J. (2018) Phylogenomic evidence overturns current conceptions of social evolution in wasps (Vespidae). Molecular Biology and Evolution, 35 (9), 2097–2109.
  82. https://doi.org/10.1093/molbev/msy124
  83. Radoszkowski, O. 1893. Faune hyménoptèrologique transcaspienne. Horae Societatis Entomologicae Rossicae, 27 (1–2), 38–81.
  84. Rahmani, Z., Rakhshani, E. & Carpenter, J.M. (2020) Updated checklist of Vespidae (Hymenoptera: Vespoidea) in Iran. Journal of Insect Biodiversity and Systematics, 6 (1), 27–86. https://doi.org/10.52547/jibs.6.1.27
  85. Richards, O.W. (1962) A revisional study of the masarid wasps (Hymenoptera, Vespoidea). British Museum of Natural History, London, vi + 294 pp.
  86. Saunders, S.S. (1850) Descriptions of some new aculeate Hymenoptera from Epirus. Transactions of the Entomological Society of London, 6 (3), 69–75, pls. 5–6. https://doi.org/10.1111/j.1365-2311.1850.tb02485.x
  87. Schmid-Egger, C., Gusenleitner, J., Liebig, W.-H. & Jacobs, H.-J. (2021) New records of Vespidae (Hymenoptera) from Iran with descriptions of three species. Linzer Biologische Beiträge, 52 (2), 1105–1121.
  88. Selis, M. (2019) A new species of Masaris Fabricius from Iran (Hymenoptera: Vespidae: Masarinae). Zootaxa, 4567 (2), 395–400. https://doi.org/10.11646/zootaxa.4567.2.12
  89. Smith, F. (1852) Descriptions of some hymenopterous insects captured in India, with notes on their oeconomy, by Ezra T. Downes, Esq., who presented them to the Honourable the East India Company. Annals and Magazine of Natural History, Series 2, 9 (49), 44–50.
  90. Thomson, C.G. (1870) Opuscula entomologica. Fasc. 2. Tryckt i Lundbergska Boktryckeriet, Lund, pp. 83–304. https://doi.org/10.5962/bhl.title.8248
  91. van Achterberg, C., Smit, J.T. & Ljubomirov, T. (2023) Review of the European Eumenes Latreille (Hymenoptera, Vespidae) using morphology and DNA barcodes, with an illustrated key to species. ZooKeys, 1143, 93–163. https://doi.org/10.3897/zookeys.1143.94951
  92. van der Vecht, J. & Fischer, F.C.J. (1972) Hymenopterorum Catalogus. Pars 8. Palaearctic Eumenidae. Junk, ‘s-Gravenhage, vi + 199 pp.
  93. von Schulthess, A. (1922) Vespiden aus Südafrika (Hym.), gesammelt von Professor Dr. Leonhard Schultze, Jena. Deutsche Entomologische Zeitschrift, 1922 (4), 399–405.
  94. Yildirim, E. & Gusenleitner, J. (2012) Contribution to the knowledge of the Vespidae (Hymenoptera, Aculeata) of Turkey, with a checklist of the Turkish species. Turkish Journal of Zoology, 36 (3), 361–374. https://doi.org/10.3906/zoo-1012-62
  95. Zhang, X., Chen, B. & Li, T.-J. (2020) A taxonomic revision of Allodynerus Blüthgen (Hymenoptera: Vespidae: Eumeninae) from China. Zootaxa, 4750 (4), 545–559. https://doi.org/10.11646/zootaxa.4750.4.5
  96. ZOBODAT (2024a) Biogeographical data record. Available from: https://www.zobodat.at/belege.php?id=101800442 (accessed 24 September 2024)
  97. ZOBODAT (2024b) Biogeographical data record. Available from: https://www.zobodat.at/belege.php?id=101800443 (accessed 24 September 2024)