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Type: Article
Published: 2025-04-15
Page range: 211-230
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A new species of montane Prasinohaema (Sphenomorphini: Scincidae) from the South-fold Mountains of New Guinea

Cesar Australia, Brunswick, Victoria 3056, Australia
Sydney School of Veterinary Science, B01, University of Sydney, New South Wales 2006, Australia; Australian Museum Research Institute, Australian Museum, 10 William St, Sydney, New South Wales 2010, Australia
Australian Museum Research Institute, Australian Museum, 10 William St, Sydney, New South Wales 2010, Australia; Herpetology Department, South Australian Museum, North Terrace, Adelaide, South Australia 5000, Australia
School of Environment and Science, Griffith University, Nathan, Queensland, Australia, and Biodiversity and Geosciences, Queensland Museum, South Brisbane, Queensland 3300, Australia
Central Cordillera ecological plasticity Lobulia group skink taxonomy Reptilia

Abstract

Over the last two decades herpetofauna surveys in the South-fold mountains of central New Guinea have revealed many new and locally endemic species. Here, we describe a new species of skink in the genus Prasinohaema from three montane localities in Papua New Guinea’s central ranges spanning approximately 80 km between the Muller Range in the northwest and the Tari Gap in the southeast. Genetic evidence indicates that Prasinohaema scurrula sp. nov. is most closely related to two other predominantly montane species of Prasinohaema, including the type species for the genus P. flavipes. It can be readily distinguished from other Prasinohaema by aspects of body size and proportions, scalation of head and digits, and colouration. The new species is likely arboreal as the paratype was collected in a tree canopy. This could explain the scarcity of observations in a relatively well-sampled region. Despite the small number of known localities there appears to be extensive suitable habitat for this species in the region and there are no major current threats, so we suggest an IUCN status of Least Concern. To facilitate further taxonomic work on Australasian skinks we also provide preferred definitions and names for key scalation states which have been variably defined and/or assigned different names in previous literature.

References

  1. Adolph, S.C. (1990) Influence of behavioral thermoregulation on microhabitat use by two Sceloporus lizards. Ecology, 71, 315–327. https://doi.org/10.2307/1940271
  2. Allison, A. & Greer, A.E. (1986) Egg shells with pustulate surface structures: basis for a new genus of New Guinea skinks (Lacertilia: Scincidae). Journal of Herpetology, 20 (1), 116–119. https://doi.org/10.2307/1564142
  3. Amarasinghe, A.A.T., Campbell, P.D., Chandramouli, S.R., Deuti, K., Raha, S., Karunarathna, D.M.S.S. & Ineich, I. (2016) Taxonomy and natural history of Eutropis beddomei (Jerdon, 1870)(Reptilia: Scincidae), including a redescription of the holotype. Zootaxa, 4132 (4), 509–520. https://doi.org/10.11646/zootaxa.4132.4.3
  4. Austin, C.C. & Jessing, K.W. (1994) Green-blood pigmentation in lizards. Comparative Biochemistry and Physiology, Part A: Physiology, 109 (3), 619–626. https://doi.org/10.1016/0300-9629(94)90201-1
  5. Bell, T. (1833) Characters of two new genera of reptiles. Proceedings of the Zoological Society of London, 1, 98–99.
  6. Boulenger, G.A. (1895) Description of two new reptiles obtained by Mr. A.S. Meek in the Trobriand Islands, British New Guinea. Annals and Magazine of Natural History, Series 6, 16 (95), 408–409.
  7. https://doi.org/10.1080/00222939508680292
  8. Boulenger, G.A. (1897) Descriptions of new lizards and frogs from Mount Victoria, Owen Stanley Range, New Guinea, collected by Mr. A. S. Anthony. The Annals and Magazine of Natural History, Series 6, 19 (109), 6–13.
  9. https://doi.org/10.1080/00222939708680502
  10. Brongersma, L.D. (1949) Zoological results of the Dutch New Guinea expedition, 1939. No. 1. A new scincid lizard. Nova Guinea, New Series, 5, 272–283, pl. I.
  11. Cámara-Leret, R., Frodin, D.G., Adema, F., Anderson, C., Appelhans, M.S., Argent, G., Guerrero, S.A., Ashton, P., Baker, W.J., Barfod, A.S., Barrington, D., Borosova, R., Bramley, G.L.C., Briggs, M., Buerki, S., Cahen, D., Callmander, M.W., Cheek, M., Chen, C.-W., Conn, B.J., Coode, M.J.E., Darbyshire, I., Dawson, S., Dransfield, J., Drinkell, C., Duyfjes, B., Ebihara, A., Ezedin, Z., Fu, L.-F., Gideon, O., Girmansyah, D., Govaerts, R., Fortune-Hopkins, H., Hassemer, G., Hay, A., Heatubun, C.D., Hind, D.J.N., Hoch, P., Homot, P., Hovenkamp, P., Hughes, M., Jebb, M., Jennings, L., Jimbo, T., Kessler, M., Kiew, R., Knapp, S., Lamei, P., Lehnert, M., Lewis, G.P., Linder, H.P., Lindsay, S., Low, Y.W., Lucas, E., Mancera, J.P., Monro, A.K., Moore, A., Middleton, D.J., Nagamasu, H., Newman, M.F., Lughadha, E.N., Melo, P.H.A., Ohlsen, D.J., Pannell, C.M., Parris, B., Pearce, L., Penneys, D.S., Perrie, L.R., Petoe, P., Poulsen, A.D., Prance, G.T., Quakenbush, J.P., Raes, N., Rodda, M., Rogers, Z.S., Schuiteman, A., Schwartsburd, P., Scotland, R.W., Simmons, M.P., Simpson, D.A., Stevens, P., Sundue, M., Testo, W., Trias-Blasi, A., Turner, I., Utteridge, T., Walsingham, L., Webber, B.L., Wei, R., Weiblen, G.D., Weigend, M., Weston, P., de Wilde, W., Wilkie, P., Wilmot-Dear, C.M., Wilson, H.P., Wood, J.R.I., Zhang, L.-B. & van Welzen, P.C. (2020) New Guinea has the world’s richest island flora. Nature, 584, 579–583. https://doi.org/10.1038/s41586-020-2549-5
  12. Das, I. & Austin, C.C. (2007) New species of Lipinia (Squamata: Scincidae) from Borneo, revealed by molecular and morphological data. Journal of Herpetology, 41 (1), 61–71. https://doi.org/10.1670/0022-1511(2007)41[61:NSOLSS]2.0.CO;2
  13. Deshmukh, U.B. (2024) Neopalaia nom. nov., a new name for Palaia Slavenko, Tamar, Tallowin, Kraus, Allison, Carranza & Meiri, 2021 (Reptilia: Squamata: Scincidae). Zootaxa, 5481 (4), 495–496. https://doi.org/10.11646/zootaxa.5481.4.9
  14. Edgar, R.C. (2004) MUSCLE: multiple sequence alignment with high accuracy and high throughput. Nucleic Acids Research, 32 (5), 1792–1797. https://doi.org/10.1093/nar/gkh340
  15. Feria-Ortiz, M. & Garcia-Vázquez, U.O. (2012) A new species of Plestiodon (Squamata: Scincidae) from Sierra Madre del Sur of Guerreri, México. Zootaxa, 3339 (1), 57–68. https://doi.org/10.11646/zootaxa.3339.1.4
  16. Fitzinger, L. (1843) Systema Reptilium. Fasciculus Primus, Amblyglossae. Apud Braumüller & Seidel Bibliopolas, Vienna, 106 pp. https://doi.org/10.5962/bhl.title.4694
  17. Frith, C.B. & Frith, D.W. (1990) Discovery of the King of Saxony Bird of Paradise Pteridophora alberti nest, egg and nestling, with notes on parental care. Bulletin of the British Ornithologists’ Club, 110, 160–164.
  18. Gray, J.E. (1845) Catalogue of the Specimens of Lizards in the Collection of the British Museum. Printed by order of the Trustees, London, xxvii + 289 pp.
  19. Greer, A.E. (1974) The generic relationships of the scincid lizard genus Leiolopisma and its relatives. Australian Journal of Zoology Supplementary Series, 22 (31), 1–67. https://doi.org/10.1071/AJZS031
  20. Greer, A.E. (1982) A new species of Leiolopisma (Lacertilia: Scincidae) from Western Australia, with notes on the biology and relationships of other Australian species. Records of the Australian Museum, 34 (12), 549–573. https://doi.org/ 10.3853/j.0067-1975.34.1982.242
  21. Greer, A.E. (1983) A new species of Lerista from Groote Eylandt and the Sir Edward Pellew Group of northern Australia. Journal of Herpetology, 17 (1), 48–53. https://doi.org/10.2307/1563780
  22. Greer, A.E., Allison, A. & Cogger, H.G. (2005) Four new species of Lobulia (Lacertilia: Scincidae) from high altitude in New Guinea. Herpetological Monographs, 19 (1), 153–179. https://doi.org/10.1655/0733-1347(2005)019[0153:FNSOLL]2.0.CO;2
  23. Greer, A.E. & Raizes, G. (1969) Green blood pigment in lizards. Science, 166 (3903), 392–393. https://doi.org/10.3853/j.0067-1975.34.1982.242
  24. Greer, A.E. & Shea, G. (2004) A new character within the taxonomically difficult Sphenomorphus group of lygosomine skinks, with a description of a new species from New Guinea. Journal of Herpetology, 38 (1), 79–87. https://doi.org/10.1670/78-03A
  25. Greer, A.E. & Simon, M. (1982) Fojia bumui, an unusual new genus and species of scincid lizard from New Guinea. Journal of Herpetology, 16, 131–139. https://doi.org/10.2307/1563805
  26. Grismer, L.L., Huatt, E.Q.S., Siler, C.D., Chan, K.O., Wood, P.L., Grismer, J.L., Sah, S.A.M. & Ahmad, N. (2011) Peninsular Malaysia’s first limbless lizard: a new species of skink of the genus Larutia (Böhme) from Pulau Pinang with a phylogeny of the genus. Zootaxa, 2799 (1), 29–40. https://doi.org/10.11646/zootaxa.2799.1.3
  27. Günther, R. & Richards, S. (2021) Description of six new species of Xenorhina Peters, 1863 from southern Papua New Guinea (Amphibia, Anura, Microhylidae). Zoosystematics and Evolution, 97 (2), 355–382. https://doi.org/10.3897/zse.97.59696
  28. Hardy, G.S. (1977) The New Zealand Scincidae (Reptilia: Lacertilia); a taxonomic and zoogeographic study. New Zealand Journal of Zoology, 4, 221–325. https://doi.org/10.1080/03014223.1977.9517956
  29. Hedges, S.B. & Conn, C.E. (2012) A new skink fauna from Caribbean islands (Squamata, Mabuyidae, Mabuyinae). Zootaxa, 3288 (1), 1–244. https://doi.org/10.11646/zootaxa.3288.1.1
  30. Horner, P. (2007) Systematics of the snake-eyed skinks, Cryptoblepharus Wiegmann (Reptilia: Squamata: Scincidae)—an Australian-based review. The Beagle: Records of the Museums and Art Galleries of the Northern Territory, 3, 21–198.
  31. Iannella, A., Richards, S. & Oliver, P. (2014) A new species of Choerophryne (Anura, Microhylidae) from the central cordillera of Papua New Guinea. Zootaxa, 3753 (5), 483–493. https://doi.org/10.11646/zootaxa.3753.5.6
  32. Kalyaanamoorthy, S., Minh, B.Q., Wong, T.K.F., von Haeseler, A. & Jermiin, L.S. (2017) ModelFinder: Fast model selection for accurate phylogenetic estimates. Nature Methods, 14, 587–589. https://doi.org/10.1038/nmeth.4285
  33. Kraus, F. (2007) Taxonomic partitioning within Papuan members of the Carlia novaeguineae complex (Squamata: Scincidae). Journal of Herpetology, 41 (3), 410–423. https://doi.org/10.1670/0022-1511(2007)41[410:TPWPMO]2.0.CO;2
  34. Kraus, F. (2010) Colour-pattern polymorphism in lizards of the genus Prasinohaema (Squamata: Scincidae). Herpetofauna, 40, 30–36.
  35. Kraus, F. (2018) A new species of Emoia (Squamata: Scincidae) from Papua New Guinea. Journal of Herpetology, 52 (4), 430–436. https://doi.org/10.1670/18-014
  36. Kraus, F. (2020) A new species of Lobulia (Squamata: Scincidae) from Papua New Guinea. Zootaxa, 4779 (2), 201–214. https://doi.org/10.11646/zootaxa.4779.2.3
  37. Lanfear, R., Calcott, B., Kainer, D., Mayer, C. & Stamatakis, A. (2014) Selecting optimal partitioning schemes for phylogenomic datasets. BMC Ecology and Evolution, 114, 82. https://doi.org/10.1186/1471-2148-14-82
  38. Losos, J.B. (2009) Lizards in an evolutionary tree: ecology and adaptive radiation of anoles. University of California Press, Berkeley, California, 528 pp. https://doi.org/10.1525/california/9780520255913.001.0001
  39. Loveridge, A. (1945) New scincid lizards of the genera Tropidophorus and Lygosoma from New Guinea. Proceedings of the Biological Society of Washington, 58, 47–52.
  40. Meyer, A.B. (1895) Zwei neue Paradiesvögel. Abhandlungen und Berichte des Königlichen Zoologischen und Anthropologisch-Ethnographischen Museums zu Dresden, 5, 1–11.
  41. Minh, B.Q., Schmidt, H.A., Chernomor, O., Schrempf, D., Woodhams, A.D., von Haeseler, A. & Lanfear, R. (2020) IQ-TREE 2: New models and efficient methods for phylogenetic inference in the genomic era. Molecular Biology and Evolution, 37, 1530–1534. https://doi.org/10.1093/molbev/msaa015
  42. Miralles, A. (2006) A new species of Mabuya (Reptilia, Squamata, Scincidae) from the Caribbean Island of San Andrés, with a new interpretation of nuchal scales: character of taxonomic importance. Herpetological Journal, 16, 1–7.
  43. Moen, D.S. & Wiens, J.J. (2017) Microhabitat and climatic niche change explain patterns of diversification among frog families. American Naturalist, 190, 29–44. https://doi.org/10.1086.692065
  44. Mys, B. (1988) The zoogeography of the scincid lizards from North Papua New Guinea (Reptilia : Scincidae). I. The distribution of the species. Bulletin de L’institut Royal Des Sciences Naturelles de Belgique Biologie, 58, 127–183.
  45. Oliver, P.M., Bower, D.S., McDonald, P.J., Kraus, F., Luedtke, J., Neam, K., Hobin, L., Chauvenet, A.L.M., Allison, A., Arida, E., Clulow, S., Günther, R., Nagombi, E., Tjaturadi, B., Travers, S.L. & Richards, S.J. (2022) Melanesia holds the world’s most diverse and intact insular amphibian fauna. Communications Biology, 5 (1), 1182. https://doi.org/10.1038/s42003-022-04105-1
  46. Oliver, P.M. & Richards, S.J. (2019) Two new species of treefrog (Pelodrydidae: Litoria) from southern New Guinea elucidated by DNA barcoding. Zootaxa, 4609 (3), 469–484.
  47. https://doi.org/10.11646/zootaxa.4609.3.4
  48. Oliver, P.M., Richards, S.J. & Sistrom, M. (2012a) Phylogeny and systematics of Melanesia’s most diverse gecko lineage (Cyrtodactylus, Gekkonidae, Squamata). Zoologica Scripta, 41 (5), 437–454. https://doi.org/10.1111/j.1463-6409.2012.00545.x
  49. Oliver, P.M., Richards, S.J. & Tjaturadi, B. (2012b) Two new species of Callulops (Anura: Microhylidae) from montane forests in New Guinea. Zootaxa, 3178 (1), 33–44. https://doi.org/10.11646/zootaxa.3178.1.3
  50. Oudemans, J.T. (1894) Eidechsen und Schildkröten. In: Zoologische Forschungsreisen in Australien und dem Malayischen Archipel. Denkschriften der Medicinisch-Naturwissenschaftlichen Gesellschaft zu Jena. Vol. 8. Fischer, Jena, pp. 127–149.
  51. Parker, H.W. (1936) V.—A Collection of Reptiles and Amphibians from the Mountains of British New Guinea. Annals and Magazine of Natural History, Series 10, 17 (97), 66–93. https://doi.org/10.1080/03745481.1936.10801389
  52. Parker, H.W. (1940) Undescribed anatomical structures and new species of reptiles and amphibians. Annals and Magazine of Natural History, Series 11, 5 (27), 257–274.
  53. https://doi.org/10.1080/00222934008527045
  54. Peters, W.C.H. (1881) Drei neue Eidechsen, zu der Familie der Scincoiden gehörig, eine Lipinia (mit geckonenuahnlicher Bildung der Zehen!) aus Neu-Guinea und wie Mocoa aus Neuholland. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin, 5, 81–85.
  55. Prasetya, A.M., Moritz, C., Joseph, L., Stelling, M.W. & Oliver, P.M. (2023) Birds and barriers: present and past seas are dominant correlates of avian turnover in the Indo-Australian Archipelago. Frontiers of Biogeography, 15 (2), e58189. https://doi.org/10.21425/F5FBG58189
  56. Quarles van Ufford, A. & Cloos, M. (2005) Cenozoic tectonics of New Guinea. AAPG bulletin, 89 (1), 119–140. https://doi.org/10.1306/08300403073
  57. Richards, S.J. & Oliver, P.M. (2010) A new scansorial species of Cophixalus (Anura: Microhylidae) from the Kikori Rivan Basin, Papua New Guinea. Journal of Herpetology, 44 (4), 555–562. https://doi.org/10.1670/09-044.1
  58. Rodriguez, Z.B., Perkins, S.L. & Austin, C.C. (2018) Multiple origins of green blood in New Guinea lizards. Science Advances, 4 (5), eaao5017. https://doi.org/10.1126/sciadv.aao5017
  59. Scheffers, B.R., Phillips, B.L., Laurance, W.F., Sodhi, N.S., Diesmos, A. & Williams, S.E. (2013) Increasing arboreality with altitude: a novel biogeographic dimension. Proceedings of the Royal Society B: Biological Sciences, 280 (1770), 20131581. https://doi.org/10.1098/rspb.2013.1581
  60. Scheffers, B.R., Shoo, L., Phillips, B., Macdonald, S.L., Anderson, A., VanDerWal, J., Storlie, C., Gourret, A. & Williams, S.E. (2017) Vertical (arboreality) and horizontal (dispersal) movement increase the resilience of vertebrates to climatic instability. Global Ecology and Biogeography, 26 (7), 787–798. https://doi.org/10.1111/geb.12585
  61. Scheffers, B.R. & Williams, S.E. (2018) Tropical mountain passes are out of reach - but not for arboreal species. Frontiers in Ecology and the Environment, 16 (2), 101–108. https://doi.org/10.1002/fee.1764
  62. Shea, G. & Greer, A.E. (2002) From Spehnomorphus to Lipinia: generic reassignment of two poorly known New Guinea skinks. Journal of Herpetology, 36 (2), 148–156. https://doi.org/10.2307/1565985
  63. Shea, G.M. (2017) A new species of Sphenomorphus (Squamata: Scincidae) from the Doberai Peninsula of New Guinea, with a redescription of Sphenomorphus consobrinus (Peters et Doria, 1878). In: Telnov, D., Barclay, M.V.L. & Pauwels, O.S.G. (Eds.), Biodiversity, biogeography and nature conservation in Wallacea and New Guinea. Vol.III The Entomological Society of Latvia, Rīga, pp. 35–48.
  64. Shea, G.M. & Allison, A. (2021) A new species of Sphenomorphus (Squamata: Scincidae) from Mount Kaindi, Morobe Province, Papua New Guinea. In: Telnov, D., Barclay, M.V.L. & Pauwels, O.S.G. (Eds.), Biodiversity, biogeography and nature conservation in Wallacea and New Guinea. Vol.VI The Entomological Society of Latvia, Rīga, pp. 49–60.
  65. Skinner, A. (2007) Phylogenetic relationships and rate of early diversification of Australian Sphenomorphus group scincids (Scincoidea, Squamata). Biological Journal of the Linnean Society, 92 (2), 347–366. https://doi.org/10.1111/j.1095-8312.2007.00843.x
  66. Slavenko, A., Allison, A., Austin, C.C., Bauer, A.M., Brown, R.M., Fisher, R.N., Ineich, I., Iova, B., Karin, B.R., Kraus, F., Mecke, S., Meiri, S., Morrison, C., Oliver, P.M., O’Shea, M., Richmond, J.Q., Shea, G.M., Tallowin, O.J.S. & Chapple, D.G. (2023) Skinks of Oceania, New Guinea, and Eastern Wallacea: an underexplored biodiversity hotspot. Pacific Conservation Biology. https://doi.org/10.1071/PC22034
  67. Slavenko, A., Richards, S.J., Donnellan, S.C., Allison, A. & Oliver, P.M. (2024) Gold in the mountains: striking new species of Papuascincus (Sphenomorphini: Scincidae) from New Guinea. Vertebrate Zoology, 74, 133–149. https://doi.org/10.3897/vz.74.e112782
  68. Slavenko, A., Tamar, K., Tallowin, O.J.S., Allison, A., Kraus, F., Carranza, S. & Meiri, S. (2020) Cryptic diversity and non-adaptive radiation of montane New Guinea skinks (Papuascincus; Scincidae). Molecular Phylogenetics and Evolution, 146, 106749. https://doi.org/10.1016/j.ympev.2020.106749
  69. Slavenko, A., Tamar, K., Tallowin, O.J.S., Kraus, F., Allison, A., Carranza, S. & Meiri, S. (2022) Revision of the montane New Guinean skink genus Lobulia (Squamata: Scincidae), with the description of four new genera and nine new species. Zoological Journal of the Linnean Society, 195 (1), 220–278. https://doi.org/10.1093/zoolinnean/zlab052
  70. Smith, M.A. (1937) A review of the genus Lygosoma (Scincidae: Reptilia) and its allies. Records of the Indian Museum, 39, 213–234.
  71. https://doi.org/10.26515/rzsi/v39/i3/1937/162283
  72. Takeuchi, W. (2011) Additional notes of Psychotria (Rubiaceae) from the southern karst of Papua New Guinea: P. defretesiana comb. et stat. nov., P. dieniensis var. nov., and P. stevedarwiniana sp. nov. Phytotaxa, 24, 19–27. https://doi.org/10.11646/phytotaxa.24.1.3
  73. Tallowin, O., Allison, A., Algar, A.C., Kraus, F. & Meiri, S. (2017) Papua New Guinea terrestrial‐vertebrate richness: elevation matters most for all except reptiles. Journal of Biogeography, 44, 1734–1744. https://doi.org/10.1111/jbi.12949
  74. Tallowin, O.J.S., Tamar, K., Meiri, S., Allison, A., Kraus, F., Richards, S.J. & Oliver, P.M. (2018) Early insularity and subsequent mountain uplift were complementary drivers of diversification in a Melanesian lizard radiation (Gekkonidae: Cyrtodactylus). Molecular Phylogenetics and Evolution, 125, 29–39. https://doi.org/10.1016/j.ympev.2018.03.020
  75. Tamura, K., Stecher, G. & Kumar, S. (2021) MEGA11: molecular evolutionary genetics analysis version 11. Molecular Biology and Evolution, 38 (7), 3022–3027. https://doi.org/10.1093/molbev/msab120
  76. Taylor, E.H. (1936) A taxonomic study of the cosmopolitan scincoid lizards of the genus Eumeces with an account of the distribution and relationships of its species. University of Kansas Bulletin, 23, 1–643.
  77. Title, P.O., Singhal, S., Grundler, M.C., Costa, G.C., Pyron, R.A., Colston, T.J., Grundler, M.R., Prates, I., Stepanova, N., Jones, M.E.H., Cavalcanti, L.B.Q., Colli, G.R., Di-Poï, N., Donnellan, S.C., Moritz, C., Mesquita, D.O., Pianka, E.R., Smith, S.A., Vitt, L.J. & Rabosky, D.L. (2024) The macroevolutionary singularity of snakes. Science, 383 (6685), 918–923. https://doi.org/10.1126/science.adh2449
  78. Toussaint, E.F.A., White, L.T., Shaverdo, H., Lam, A., Surbakti, S., Panjaitan, R., Sumoked, B., von Rintelen, T., Sagata, K. & Balke, M. (2021) New Guinean orogenic dynamics and biota evolution revealed using a custom geospatial analysis pipeline. BMC Ecology and Evolution, 21 (1), 1–28. https://doi.org/10.1186/s12862-021-01764-2
  79. Wikramanayake, E.D. & Dryden, G.L. (1993) Thermal ecology of habitat and microhabitat use by sympatric Varanus bengalensis and V salvator in Sri Lanka. Copeia, 1993, 709–714. https://doi.org/10.2307/1447231
  80. Williams, E.E. (1983) Ecomorphs, faunas, island size, and diverse end points in island radiation of Anolis. In: Huey, R.B., Pianka, E.R. & Schoener, T.W. (Eds.), Lizard ecology: studies of a model organism. Harvard University Press, Cambridge, Massachusetts, pp. 326–370. https://doi.org/10.4159/harvard.9780674183384.c18
  81. Withers, P.C. (1981) Physiological correlates of limblessness and fossoriality in scincid lizards. Copeia, 1981, 197–204. https://doi.org/10.2307/1444055
  82. Woodruff, D.S. (1972) Amphibians and reptiles from Simbai, Bismark-Schrader Range, New Guinea. Memoirs of the National Museum of Victoria, 33, 57–64. https://doi.org/10.24199/j.mmv.1972.33.08
  83. Zug, G.R. (2004) Systematics of the Carlia ‘fusca’ lizards (Squamata: Scincidae) of New Guinea and nearby islands. Bishop Museum Bulletin in Zoology, 5, 1–83.

How to Cite

Slavenko, A., Shea, G.M., Richards, S.J. & Oliver, P.M. (2025) A new species of montane Prasinohaema (Sphenomorphini: Scincidae) from the South-fold Mountains of New Guinea. Zootaxa, 5621 (2), 211–230. https://doi.org/10.11646/zootaxa.5621.2.3