Abstract
The checklist of the millipedes of Vietnam was re-documented and updated after 20 years. A total of 280 species belonging to 97 genera, 31 families and 14 orders have been recorded. Of these, 246 species and 21 genera are considered to be endemic to the Vietnam fauna. Four previously listed species, Unixenus sp.; Platyrhacus marginellis Silvestri, 1911; Platyrhacus bouvieri Brölemann, 1896; and Oxidus gracilis (C.L. Koch, 1847) are removed from the current list. Sundanina cochinchina Nguyen, 2010 is transferred to the genus Enghoffosoma Golovatch, 1993 as Enghoffosoma cochinchina (Nguyen, 2010) comb. nov. Nesorthomorpha montana Nguyen, Tran & Le, 2018 is synonymized with Antheromorpha nguyeni Likhitrakarn, Golovatch, Semenyuk & Panha, 2019 syn. nov..
References
- Akkari, N., Macek, O. & Stoev, P. (2023) A revision of the millipede family Paracortinidae (Diplopoda, Callipodida). ZooKeys, 1187, 341–399. https://doi.org/10.3897/zookeys.1187.113473
- Attems, C. (1907) Javanische Myriopoden, gesammelt von Direktor K. Kraepelin im Jahre 1903. Mitteilungen aus dem Naturhistorischen Museum in Hamburg, 24, 77–142.
- Attems, C. (1914) Die indo-australischen Myriopoden. Archiv für Naturgeschichte, 80A (4), 1–398.
- Attems, C. (1930a) Myriopoden der Kleinen Sunda-Inseln, gesammelt von der Expedition Dr. Rensch. Mitteilungen aus dem Zoologischen Museum in Berlin, 16 (1), 117–184. https://doi.org/10.1002/mmnz.19300160103
- Attems, C. (1930b) Myriopoden von Java, Sumatra und Bali. Archiv für Hydrobiologie, Supplement8, 115–192.
- Attems, C. (1932) Neue Polydesmiden des Museums Buitenzorg. Treubia, 14 (1), 29–41.
- Attems, C. (1936) Diplopoda of India. Memoirs of the Indian Museum, 11 (4), 133–323.
- Attems, C. (1937) Myriapoda 3. Polydesmoidea I. Fam. Strongylosomidae. Das Tierreich, 68, 1–300. https://doi.org/10.1515/9783111567099
- Attems, C. (1938) Die von Dr. C. Dawydoff in französisch Indochina gesammelten Myriopoden. Mémoires du Muséum national d’histoire naturelle, New Series, 6 (2), 187–353.
- Attems, C. (1942) Zur Kenntnis der Indischen Harpagophoriden. Annalen des Naturhistorischen Museums in Wien, 52, 66–105.
- Attems, C. (1951) Revision systématique des Colobognata (Myriapodes Diplopodes) et description d’espèces nouvelles. Mémoires du Muséum national d’histoire naturelle, New Series, Série A, 3 (3), 193–231.
- Attems, C. (1953) Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938-1939). Mémoires du Muséum national d’histoire naturelle, New Series, Série A, 5 (3), 133–230.
- Brölemann, H.W. (1896) Myriapodes recueillis en Indo-Chine par M. Pavie et offerts au Museum d’Histoire Naturelle de Paris. Bulletin du Muséum national d’histoire naturelle, Series 2 (7), 332–333.
- Brölemann, H.W. (1904) Myriapodes recueillis par M. A. Pavie en Indo-Chine. In: Pavie, A. (Ed.), Mission Pavie Indo-Chine 1879–1895. A. Etudes diverses. III. Rechereches sur l’histoire naturelle de l’Indo-Chine Orientale. E. Leroux, Paris, pp. 296–308.
- Brölemann, H.W. (1916) Essai de classification des Polydesmiens. Annales de la Société entomologique de France, 84 (4), 523–608. https://doi.org/10.1080/21686351.1915.12279415
- Butler, A.G. (1876) Preliminary notice of new species of Arachnida and Myriapod from Rodriguez. Annals and Magazine of Natural History, Series 4, 17, 445. https://doi.org/10.1080/00222937608681989
- Carl, J. (1912) Sur quelques colobognathes du Muséum du Genève. Revue suisse de Zoologie, 20 (9), 507–518.
- Carl, J. (1917) Spirostreptides nouveaux ou peu connus du Museum de Geneve. Revue suisse de zoologie, 25, 383–409. https://doi.org/10.5962/bhl.part.31930
- Carl, J. (1941) Diplopoden aus Südindien und Ceylon. 2. Teil: Nematophora und Juliformia. Revue suisse de zoologie, 48 (22), 569–714. https://doi.org/10.5962/bhl.part.154483
- Chamberlin, R.V. & Wang, Y.-H.M. (1953) Records of millipeds (Diplopoda) from Japan and other oriental areas, with descriptions of new genera and species. American Museum Novitates, 1621, 1–13.
- Chen, C., Golovatch, S.I. & Chang, H. (2006) The millipede tribe Nedyopodini, with special reference to the fauna of Taiwan (Diplopoda: Polydesmida: Paradoxosomatidae). Journal of Natural History, 39 (47), 3997–4030. https://doi.org/10.1080/00222930600556112
- Demange, J.-M. (1961) Matériaux pour servir a une révision des Harpagophoridae (Myriapodes-Diplopodes). Mémoires du Muséum national d’histoire naturelle, New Series, Série A, 24, 1–274.
- Demange, J.-M. (1963) Données nouvelles sur la famille des Harpagophoridae (Myriapoda, Diplopoda). Bulletin du Muséum National d’Histoire Naturelle, Section A, Series 2 (4e), 561–584. https://doi.org/10.5962/p.285981
- Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschungsinstitut Senckenberg, 93, 241–331.
- Enghoff, H. (2001) Benoitolus flavicollis Mauries, 1980, another tropical tramp millipede, and notes on Pseudospirobolellus avernus (Butler, 1876) (Diplopoda, Spirobolida, Pseudospirobolellidae). Entomologist’s Monthly Magazine, 137 (1644–1647), 167–170.
- Enghoff, H., Golovatch, S.I. & Nguyen, D.A. (2004) A review of the millipede fauna of Vietnam. Arthropoda Selecta, 13 (1–2), 25–43.
- Eydoux, F. & Souleyet, L.F.A. (1842) Voyage Autour du Monde, Exécuté Pendant les Années 1836 et 1837 sur la Corvette La Bonite, Commandée par M. Vaillant. Zoologie. Tome Premier. Seconde Partie. Arthus Bertrand, Paris, 202 pp. [pp. 133–334]
- Gervais, P. (1836) Études pour servir á l’histoire naturelle des Myriapodes. Annales des Sciences naturelles, Zoologie, Series 2, 7, 35–60.
- Gervais, P. (1847) Myriapodes. In: Walckenaer, L.B. & Gervais, P. (Eds.), Histoire naturelle des Insectes Aptères. Vol. 4. Librairie encyclopédique de Roret, Paris, pp. 8–330.
- Golovatch, S.I. (1983a) Contributions to the millipede fauna of Vietnam (Diplopoda) I. Acta Zoologica Hungarica, 29 (1–3), 123–127.
- Golovatch, S.I. (1983b) Millipedes (Diplopoda) of the fauna of Vietnam. In: Medvedev, L.N. (Eds.), Fauna and ecology of the animals of Vietnam. Nauka, Moscow, pp. 1–207.
- Golovatch, S.I. (1983c) On several new Glomeridae (Diplopoda) from Indochina. Annales historico-naturales Musei Nationalis Hungarici, 75, 107–116.
- Golovatch, S.I. (1984) Contributions to the millipede fauna of Vietnam (Diplopoda) II. Acta Zoologica Hungarica, 30 (1–2), 53–77.
- Golovatch, S.I. (1987) Diplopoda from the Nepal Himalayas. Opisotretidae, additional Polydesmidae and Fuhrmannodesmidae. Courier Forschungsinstitut Senckenberg, 933, 203–217.
- Golovatch, S.I. (1994) Diplopoda from the Himalayas. Two new Alogolykini (Polydesmida: Paradoxosomatidae). Senckenbergiana biologica, 73 (1–2), 183–187.
- Golovatch, S.I. (1998) On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda Polydesmida), VI. Golovatch, S.I., 6 (3–4), 35–46.
- Golovatch, S.I. (2009a) On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), IX. Arthropoda Selecta, 18 (3–4), 119–124.
- Golovatch, S.I. (2009b) On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), VIII. Arthropoda Selecta, 18 (1–2), 1–7. https://doi.org/10.15298/arthsel.32.1.01
- Golovatch, S.I. (2015) Review of the millipede genus Ophrydesmus Cook, 1896, with the description of a new species from southern Vietnam (Diplopoda: Polydesmida: Cryptodesmidae). Tropical Natural History, 15 (2), 155–165. https://doi.org/10.58837/tnh.15.2.103079
- Golovatch, S.I. (2016a) On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XIX. Arthropoda Selecta, 25 (2), 131–152. https://doi.org/10.15298/arthsel.25.2.01
- Golovatch, S.I. (2016b) The millipede family Cryptodesmidae in Indochina (Diplopoda, Polydesmida). ZooKeys, 578, 33–43. https://doi.org/10.3897/zookeys.578.7994
- Golovatch, S.I. (2017) Another two new species of the millipede family Glomeridae from Vietnam, with a new record of Rhopalomeris variegata Golovatch et Semenyuk, 2016 from southern Vietnam (Diplopoda: Glomerida). Russian Entomology Journal, 26 (2), 195–202. https://doi.org/10.15298/rusentj.26.2.10
- Golovatch, S.I. (2019) On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXVII. Arthropoda Selecta, 28 (4), 459–478. https://doi.org/10.15298/arthsel.28.4.01
- Golovatch, S.I. (2023) On the synonymy of the millipede genera Riukiaria Attems, 1938 and Parariukiaria Nguyen, 2016, with records of Riukiaria cf. davidiani Golovatch, 2014 from Sichuan, southern China (Diplopoda: Polydesmida: Xystodesmidae). Arthropoda Selecta, 32 (3), 231–238. https://doi.org/10.15298/arthsel.32.3.01
- Golovatch, S.I. (2024) A new species and some new records of millipedes (Diplopoda) from southern Vietnam. Russian Entomological Journal, 33 (3), 407–413. https://doi.org/10.15298/rusentj.33.3.14
- Golovatch, S.I. & Akkari, N. (2016) Identity of the millipede, Pseudoniponiella kometis (Attems, 1938) (Diplopoda: Polydesmida: Cryptodesmidae). Tropical Natural History, 16 (1), 1–6. https://doi.org/10.58837/tnh.16.1.103069
- Golovatch, S.I. & Enghoff, H. (1993) Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia, 19 (3), 85–125.
- Golovatch, S.I. & Enghoff, H. (1994) Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia, 20 (2), 45–71.
- Golovatch, S.I., Geoffroy, J.-J. & Akkari, N. (2017) Revision of the Vietnamese millipede genus Annamina Attems, 1937, with descriptions of three new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys, 669, 1–18. https://doi.org/10.3897/zookeys.669.12561
- Golovatch, S.I., Geoffroy, J.-J. & Mauriès, J.-P. (2006) Review of the millipede genus Hyleoglomeris Verhoeff, 1910 (Diplopoda, Glomerida, Glomeridae), with descriptions of new species from caves in Southeast Asia. Zoosystema, 28 (4), 887–915.
- Golovatch, S.I., Geoffroy, J.J., Mauries, J.P. & Van-den-Spiegel, D. (2007) Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group. Zoosystema, 29 (3), 417–456.
- Golovatch, S.I., Geoffroy, J.-J., Mauriès, J.-P. & VandenSpiegel, D. (2009a) Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys, 7, 1–53. https://doi.org/10.3897/zookeys.7.117
- Golovatch, S.I., Geoffroy, J.-J., Mauriès, J.-P. & VandenSpiegel, D. (2009b) Review of the millipede genus Eutrichodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae), with descriptions of new species. ZooKeys, 12, 1–46. https://doi.org/10.3897/zookeys.12.167
- Golovatch, S.I., Geoffroy, J.-J., Mauriès, J.-P. & VandenSpiegel, D. (2009c) Review of the millipede genus Plusioglyphiulus Silvestri, 1923, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema, 31 (1), 71–116. https://doi.org/10.5252/z2009n1a5
- Golovatch, S.I., Geoffroy, J.-J., Mauriès, J.-P. & VandenSpiegel, D. (2010) Two new species of the millipede genus Trichopeltis Pocock, 1994 (Diplopoda: Polydesmida: Cryptodesmidae) from Vietnam and China. Arthropoda Selecta, 19 (2), 63–72. https://doi.org/10.15298/arthsel.19.2.02
- Golovatch, S.I., Geoffroy, J.-J., Mauriès, J.-P. & VandenSpiegel, D. (2011a) New species of the millipede genus Glyphiulus Gervais, 1847 from the javanicus-group (Diplopoda: Spirostreptida: Cambalopsidae). Arthropoda Selecta, 20 (3), 149–165. https://doi.org/10.15298/arthsel.20.3.02
- Golovatch, S.I., Geoffroy, J.-J., Mauriès, J.-P. & VandenSpiegel, D. (2011b) Two new species of the millipede genus Hypocambala Silvestri, 1895 from China and Vietnam (Diplopoda: Spirostreptida: Cambalopsidae). Arthropoda Selecta, 20 (3), 167–174. https://doi.org/10.15298/arthsel.20.3.03
- Golovatch, S.I., Geoffroy, J.-J., Mauriès, J.-P. & VandenSpiegel, D. (2012a) New or poorly-known species of the millipede genus Trachyjulus Peters, 1864 (Diplopoda: Spirostreptida: Cambalopsidae). Arthropoda Selecta, 21 (2), 103–129. https://doi.org/10.15298/arthsel.21.2.01
- Golovatch, S.I., Geoffroy, J.-J., Mauriès, J.-P. & VandenSpiegel, D. (2016a) Four new species of the millipede genus Eutrichodesmus Silvestri, 1910, from caves in Indochina (Diplopoda: Polydesmida: Haplodesmidae). Arthropoda Selecta, 25 (3), 247–256. https://doi.org/10.15298/arthsel.25.3.03
- Golovatch, S.I., Geoffroy, J.-J., Stoev, P. & Vanden Spiegel, D. (2013a) Review of the millipede family Opisotretidae (Diplopoda, Polydesmida), with descriptions of new species. ZooKeys, 302, 13–77. https://doi.org/10.3897/zookeys.302.5357
- Golovatch, S.I., Geoffroy, J.J. & VandenSpiegel, D. (2013b) On several new species of the millipede family Glomeridae from Vietnam (Diplopoda: Glomerida). Arthropoda Selecta, 22 (3), 201–206. https://doi.org/10.15298/arthsel.22.3.02
- Golovatch, S.I., Geoffroy, J.-J. & VandenSpiegel, D. (2014) Review of the millipede family Trichopolydesmidae in the Oriental realm (Diplopoda, Polydesmida), with descriptions of new genera and species. ZooKeys, 414, 19–65. https://doi.org/10.3897/zookeys.414.7671
- Golovatch, S.I. & Korsós, Z. (1990) Contributions to the millipede fauna of Vietnam (Diplopoda) 3. Spirobolida. Acta Zoologica Hungarica, 36 (1–2), 25–36.
- Golovatch, S.I., Mikhaljova, E.V. & Chang, H.-W. (2011c) The millipede families Cryptodesmidae, Haplodesmidae, Pyrgodesmidae, Opisotretidae and Xystodesmidae in Taiwan (Diplopoda, Polydesmida). Tropical Natural History, 11 (2), 129–143. https://doi.org/10.58837/tnh.11.2.102999
- Golovatch, S.I. & Nguyen, A.D. (2007) Two new species of the millipede genus Platyrhacus C.L. Koch, 1847 from Vietnam (Diplopoda: Polydesmida: Platyrhacidae). Arthropoda Selecta, 15 (3), 215–224.
- Golovatch, S.I., Semenyuk, I., VandenSpiegel, D. & Anichkin, A.E. (2011d) Three new species of the millipede family Pyrgodesmidae from Nam Cat Tien National Park, southern Vietnam (Diplopoda: Polydesmida). Arthropoda Selecta, 20 (1), 1–9. https://doi.org/10.15298/arthsel.20.1.01
- Golovatch, S.I. & Semenyuk, I.I. (2010) On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), X. Arthropoda Selecta, 19 (3), 123–127. https://doi.org/10.15298/arthsel.19.3.02
- Golovatch, S.I. & Semenyuk, I.I. (2016a) Two new species of the millipede family Glomeridae from Vietnam (Diplopoda: Glomerida). Russian Entomology Journal, 25 (4), 411–416. https://doi.org/10.15298/rusentj.25.4.11
- Golovatch, S.I. & Semenyuk, I.I. (2016b) Two new species of the millipede family Glomeridae from Vietnam (Diplopoda: Glomerida). Russian Entomology Journal, 25 (4), 411–416. https://doi.org/10.15298/rusentj.25.4.11
- Golovatch, S.I. & Semenyuk, I.I. (2018a) On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXIII. Arthropoda Selecta, 27 (1), 1–21. https://doi.org/10.15298/arthsel.27.1.01
- Golovatch, S.I. & Semenyuk, I.I. (2018b) On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXIV. Arthropoda Selecta, 27 (1), 187–200. https://doi.org/10.15298/arthsel.27.3.01
- Golovatch, S.I. & Semenyuk, I.I. (2021) On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXX. Arthropoda Selecta, 30 (4), 473–496. https://doi.org/10.15298/arthsel.30.4.04
- Golovatch, S.I., Tiunov, A.V. & Anichkin, A.E. (2011e) Millipedes (Diplopoda). In: Tiunov, A.V. (Ed.), Structure and functions of soil communities of a monsoon tropical forest (Cat Tien National Park, southern Vietnam). KMK Scientific Press, Moscow, pp. 76–90.
- Golovatch, S.I., Vandenspiegel, D. & Semenyuk, I.I. (2016b) On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXI. Arthropoda Selecta, 25 (4), 335–354. https://doi.org/10.15298/arthsel.25.4.01
- Golovatch, S.I., Wesener, T., Mauriès, J.P. & Semenyuk, I.I. (2012b) On the identities of Cryxus Leach, 1814 and Zephronia Gray, 1832, the oldest generic names in the millipede order Sphaerotheriida (Diplopoda). Arthropoda Selecta, 21 (No. 4) 273–294. https://doi.org/10.15298/arthsel.21.4.01
- Golovatch, S.I., Wytwer, J. & Jedryczkowski, W. (2003) The millipede Vaulogerodesmus pictus (Brölemann, 1916) rediscovered, with a review of this Southeast Asian genus (Diplopoda: Polydesmida: Paradoxosomatidae). Arthropoda Selecta, 11 (4), 259–264.
- Gray, J.E. (1832) The myriapods. In: Griffith, E. (Ed.), The Animal Kingdom II. Whittaker & Co., London, pp. 135.
- Hoffman, R.L. (1963a) A contribution to the knowledge of asiatic strongylosomoid Diplopoda (Polydesmida: Strongylosomatidae). Annals and Magazine of Natural History, Series 13, 5, 577–593. https://doi.org/10.1080/00222936208651289
- Hoffman, R.L. (1963b) A contribution to the knowledge of asiatic strongylosomoid Diplopoda (Polydesmida: Strongylosomatidae). Annals and Magazine of Natural History, Series13, 5, 577–593. https://doi.org/10.1080/00222936208651289
- Hoffman, R.L. (1973) Descriptions and allocation of new or poorly known genera and species of Paradoxosomatidae from south-eastern Asia (Diplopoda:Polydesmida). Journal of Natural History, 7 (4), 361–389. https://doi.org/10.1080/00222937300770281
- Hoffman, R.L. (1975) Studies on spirostreptoid millipeds. XI. A review of some Indonesian genera of the family Harpagophoridae. Journal of Natural History, 9 (2), 121–152. https://doi.org/10.1080/00222937500770101
- Hoffman, R.L. (1977) Diplopoda from Malayan caves, collected by M. Pierre Strinati. Revue suisse de Zoologie, 84 (3), 699–719.
- Hoffman, R.L. (1980) Classification of the Diplopoda. Muséum d’Histoire Naturelle, Genève, 237 pp.
- Hoffman, R.L. (1981) Studies on spiroboloid millipeds. XIV. Notes on the family Pseudospirobolellidae, and description og a new genus and species from Thailand. Steenstrupia, 7 (7), 181–190.
- Hoffman, R.L. & Keeton, W.T. (1960) A list of the generic names proposed in the Diplopod order Spirobolida, with their type species. Transactions of the American Entomological Society, 86, 1–26.
- Hoffman, R.L. & Payne, J.A. (1969) Diplopods as carnivores. Ecology, 50 (6), 1096–1098. https://doi.org/10.2307/1936905
- Hopkin, S.P. & Read, H.J. (1992) The biology of millipedes. Oxford, New York and Tokyo, 233 pp. https://doi.org/10.1093/oso/9780198576990.001.0001
- Huynh, C. & Veenstra, A.A. (2015) Description of a new species of Penicillate Millipede from the genus Monographis (Diplopoda: Polyxenidae) found in Vietnam. Zootaxa, 3964 (4), 460–474. https://doi.org/10.11646/zootaxa.3964.4.5
- Huynh, C. & Veenstra, A.A. (2018) Two new species of penicillate millipedes (Diplopoda, Polyxenidae) from Phu Quoc Island in southern Vietnam. Zootaxa, 4402 (2), 283–302. https://doi.org/10.11646/zootaxa.4402.2.3
- Huynh, C. & Veenstra, A.A. (2020) Three new species of penicillate millipedes from the Con Dao Islands of southeast Vietnam (Diplopoda, Lophoproctidae and Polyxenidae). Zootaxa, 4759 (1), 1–30. https://doi.org/10.11646/zootaxa.4759.1.1
- Huynh, C. & Veenstra, A.A. (2022) Four new species of Monographis Attems, 1907 (Diplopoda, Polyxenida, Polyxenidae) from Vietnam. Zootaxa, 5214 (3), 393–420. https://doi.org/10.11646/zootaxa.5214.3.4
- Jeekel, C.A.W. (1953) Two new Strongylosomidae from Indochina (Diplopoda, Polydesmida). Beaufortia, 2 (29), 1–8.
- Jeekel, C.A.W. (1963a) Diplopoda of Guiana. Studies on the Fauna of Suriname and other Guyanas, 4 (11), 1–157.
- Jeekel, C.A.W. (1963b) Paradoxosomatidae from Borneo (Diplopoda, Polydesmida). Tijdschrift voor Entomologie, 106 (6), 205–283.
- Jeekel, C.A.W. (1964a) A new species of Orthomorpha Bollman from Thailand observed in migration, with taxonomic notes on the genus (Diplopoda). Tijdschrift voor Entomologie, 107, 355–364.
- Jeekel, C.A.W. (1964b) Two new species of Pratinus Attems, with taxonomoc notes on the genus and a redescription of its type-species (Diplopoda, Polydesmida). Beaufortia, 11 (137), 61–73.
- Jeekel, C.A.W. (1965) A revision of the Burmese Paradoxosomatidae (Diplopoda, Polydesmida) in the Museo Civico di Storia Naturale at Genoa (Part I). Tijdschrift voor Entomologie, 108 (5), 95–144.
- Jeekel, C.A.W. (1968) On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida). Privately published, Rotterdam, 162 pp.
- Jeekel, C.A.W. (1980) The generic allocation of some little-known Paradoxosomatidae from South-East Asia (Diplopoda, Polydesmida). Revue Suisse de Zoologie, 87 (3), 651–670. https://doi.org/10.5962/bhl.part.85538
- Jeekel, C.A.W. (2001a) A bibliographic catalogue of the Asiatic Sphaerotheriida (Diplopoda). Myriapod memoranda, 3, 5–38.
- Jeekel, C.A.W. (2001b) A bibliographic catalogue of the Siphonophorida (Diplopoda). Myriapod Memoranda, 3, 44–71.
- Jeekel, C.A.W. (2001c) A bibliographic catalogue of the Spirobolida of the Oriental and Australian regions (Diplopoda). Myriapod memoranda, 4, 5–104.
- Jeekel, C.A.W. (2002) On the year of publication of the Myriapoda in Eydoux & Souleyet, Zoology of La Bonite. Myriapod memoranda, 5, 35–39.
- Jeekel, C.A.W. (2006) A bibliographic catalogue of the Oriental Harpagophoridae (Diplopoda, Spirostreptida). Myriapod memoranda, 9, 5–58.
- Kime, R.D. & Golovatch, S.I. (2000) Trends in the ecological strategies and evolution of millipedes (Diplopoda). Biological Journal of the Linnean Society, 69, 333–349. https://doi.org/10.1111/j.1095-8312.2000.tb01209.x
- Korsós, Z. & Golovatch, S.I. (1989) Addenda to the millipede fauna of Vietnam (Diplopoda). Acta Zoologica Hungarica, 35 (3–4), 211–220.
- Kuroda, M., Eguchi, K., Oguri, E. & Nguyen, A.D. (2022) Two new cave Hyleoglomeris species (Glomerida, Glomeridae) from northern Vietnam. ZooKeys, 1108, 161–174. https://doi.org/10.3897/zookeys.1108.85423
- Likhitrakarn, N., Golovatch, S.I. & Panha, S. (2011) Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys, 131, 1–161. https://doi.org/10.3897/zookeys.131.1921
- Likhitrakarn, N., Golovatch, S.I. & Panha, S. (2014) Review of the Southeast Asian millipede genus Enghoffosoma Golovatch, 1993 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of new species. Zootaxa, 3811 (4), 491–514. https://doi.org/10.11646/zootaxa.3811.4.4
- Likhitrakarn, N., Golovatch, S.I. & Panha, S. (2015) Two new species of dragon millipedes, genus Desmoxytes Chamnerlin, 1923, from Laos (Diplopoda: Polydesmida: Paradoxosomatidae), with redescriptions of all four species of Attems from Vietnam. Zootaxa, 3931 (1), 483–504. https://doi.org/10.11646/zootaxa.3931.4.2
- Likhitrakarn, N., Golovatch, S.I. & Panha, S. (2016) Review of the Southeast Asian millipede genus Antheromorpha Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys, 571, 21–57. https://doi.org/10.3897/zookeys.571.7566
- Likhitrakarn, N., Golovatch, S.I., Semenyuk, I., Efeykin, B.D. & Panha, S. (2019a) Review of the millipede genus Orthomorpha bollman, 1893 (Diplopoda, Polydesmida, Paradoxosomatidae) in Vietnam, with several new records and descriptions of two new species. ZooKeys, 898, 121–158. https://doi.org/10.3897/zookeys.898.39265
- Likhitrakarn, N., Golovatch, S.I., Semenyuk, I.I. & Panha, S. (2019b) A new species and a new record of the Southeast Asian millipede genus Antheromorpha Jeekel, 1968 (Polydesmida, Paradoxosomatidae) from Vietnam. ZooKeys, 832, 77–89. https://doi.org/10.3897/zookeys.832.32596
- Marek, P.E., Krejca, J.K. & Shear, W.A. (2016) A new species of Illacme Cook Loomis, 1928 from Sequoia National Park, California, with a world catalog of the Siphonorhinidae (Diplopoda, Siphonophorida). ZooKeys, 626, 1–43. https://doi.org/10.3897/zookeys.626.9681
- Mauriès, J.-P. (1977) Le genre Glyphiulus Gervais, 1847, et sa place dans la classification des Cambalides, à propos de la description d’une nouvelle espèce du Viêt-Nam (Diplopoda, Iulida, Cambalidea). Bulletin du Muséum national d’histoire naturelle, 3e Série, Zoologie, 301 (431), 243–250. https://doi.org/10.5962/p.282031
- Mauriès, J.-P. & Enghoff, H. (1990) A new genus of cambaloid millipedes from Vietnam (Diplopoda: Spirostreptida: Cambalopsidae). Entomologica scandinavica, 21, 91–96. https://doi.org/10.1163/187631290X00076
- Mauriès, J.-P., Golovatch, S.I. & Geoffroy, J.-J. (2010) A new genus and species of the order Stemmiulida from Vietnam (Diplopoda). Arthropoda Selecta, 19 (2), 73–80. https://doi.org/10.15298/arthsel.19.2.03
- Mauriès, J.-P., Golovatch, S.I. & Hoffman, R.L. (2001) On type material and the identity of several Iulus species described by Paul Gervais, in the collection of the Muséum national d’ Histoire naturelle in Paris (Diplopoda, Spirostreptida, Spirobolida). Zoosystema, 23 (3), 579–589.
- Minelli, A. (2014) Treatise on Zoology: Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 2. Brill, Leiden, 482 pp. https://doi.org/10.1163/9789004188273_002
- Nguyen, A.D. (2009) A new species of the family Polydesmidae (Diplopoda: Polydesmida) from Vietnam. The International Journal of Myriapodology, 2 (1), 63–68.
- Nguyen, A.D. (2010a) The millipede genus Anoplodesmus Pocock, 1895 in Vietnam (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa, 2649 (1), 52–60. https://doi.org/10.11646/zootaxa.2649.1.2
- Nguyen, A.D. (2010b) The millipede tribe Sundaninini in Vietnam (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa, 2479 (1), 59–68. https://doi.org/10.11646/zootaxa.2479.1.4
- Nguyen, A.D. (2011) A review of the millipede tribe Tonkinosomatini (Diplopoda: Polydesmida: Paradoxosomatidae) from Vietnam. Zootaxa, 3036 (1), 58–68. https://doi.org/10.11646/zootaxa.3036.1.4
- Nguyen, A.D. (2012) Tylopus Millipedes in Vietnam (Diplopoda: Polydesmida: Paradoxosomatidae: Sulciferini), with descriptions of five new species. Raffles Bulletin of Zoology, 60 (2), 289–311.
- Nguyen, A.D. (2016) Discovery of a new millipede species in northern Vietnam, and the proposal of a new genus, Parariukiaria (Diplopoda, Polydesmida, Xystodesmidae). Zootaxa, 4121 (3), 331–336. https://doi.org/10.11646/zootaxa.4121.3.7
- Nguyen, A.D. (2017) A second species of the genus Vietnamorpha Golovatch, 1984 (Polydesmida: Paradoxosomatidae) and notes on the generic relationship. Journal of Natural History, 51 (39–40), 2331–2343. https://doi.org/10.1080/00222933.2017.1355491
- Nguyen, A.D. & Eguchi, K. (2022) Mountainous millipedes in Vietnam. II. A conspicuous Tylopus species from Northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys, 1091, 1–13. https://doi.org/10.3897/zookeys.1091.80094
- Nguyen, A.D., Eguchi, K. & Hwang, U.W. (2019a) Two new pill millipedes (Diplopoda: Glomerida: Glomeridae) from high mountains of Vietnam. Journal of Natural History, 53 (21–22), 1369–1384. https://doi.org/10.1080/00222933.2019.1646338
- Nguyen, A.D. & Golovatch, S.I. (2016) The millipede genus Enghoffosoma Golovatch, 1993 recorded in Vietnam for the first time, with descriptions of three new species (Diplopoda, Polydesmida, Paradoxosomatidae). Zootaxa, 4139 (2), 151–166. https://doi.org/10.11646/zootaxa.4139.2.1
- Nguyen, A.D., Golovatch, S.I. & Anichkin, A.E. (2006) The dragon millipedes in Vietnam (Polydesmida: Paradoxosomatidae, genus Desmoxytes Chamberlin, 1923). Arthropoda Selecta, 14 (3), 251–257.
- Nguyen, A.D., Jang, K.-H. & Hwang, U.-W. (2016) The first record of the millipede genus Streptogonopus Attems, 1914 from Vietnam, with description of a new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys, 601, 111–118. https://doi.org/10.3897/zookeys.601.9165
- Nguyen, A.D. & Korsós, Z. (2011) A revision of the millipede genus Riukiupeltis Verhoeff, 1939 (Diplopoda, Polydesmida, Paradoxosomatidae), with comments on the status of related species. ZooKeys, 156, 25–40. https://doi.org/10.3897/zookeys.156.2009
- Nguyen, A.D., Korsós, Z., Jang, K.H. & Hwang, U.W. (2017) A revision and phylogenetic analysis of the millipede genus Oxidus Cook, 1911 (Polydesmida: Paradoxosomatidae). European Journal of Taxonomy, 293, 1–22. https://doi.org/10.5852/ejt.2017.293
- Nguyen, A.D., Nguyen, D.D. & Eguchi, K. (2021a) Mountainous millipedes in Vietnam. I. Two new species of the family Paradoxosomatidae from Mount Fansipan (Diplopoda, Polydesmida). ZooKeys, 1032, 1–15. https://doi.org/10.3897/zookeys.1032.64917
- Nguyen, A.D., Nguyen, L.T.P. & Korsós, Z. (2024a) A review of the millipede genus Pseudodesmus Pocock, 1887 (Diplopoda, Platydesmida, Andrognathidae) from Vietnam, with descriptions of five new species and notes on its phylogeny. Zoosystematics and Evolution, 100 (4), 1515–1541. https://doi.org/10.3897/zse.100.132111
- Nguyen, A.D., Nguyen, M.-H., Nguyen, T.-A.T. & Phung, H.-L.T. (2019b) Review of dragon millipedes (Diplopoda, Polydesmida, Paradoxosomatidae) in the fauna of Vietnam, with descriptions of three new species. Zoological Studies, 58, 14. https://doi.org/10.6620/ZS.2019.58-14
- Nguyen, A.D., Nguyen, S.G. & Eguchi, K. (2021b) A new Rhopalomeris species (Diplopoda: Glomerida: Glomeridae), and notes on the phylogenetic relationships between glomeridans in Vietnam. ZooKeys, 4927 (2), 259–263. https://doi.org/10.11646/zootaxa.4927.2.5
- Nguyen, A.D., Nguyen, S.G. & Le, S.X. (2018a) On the millipede genus Antheromorpha Jeekel, 1968 (Diplopoda: Polydesmida: Paradoxosomatidae) from Vietnam, with a description of a new species. Raffles Bulletin of Zoology, 66.
- Nguyen, A.D. & Sierwald, P. (2013) A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida). Check List, 9 (6), 1132–1353. https://doi.org/10.15560/9.6.1132
- Nguyen, A.D. & Sierwald, P. (2019) Distribution and descriptive characteristics of the two giant pill millipedes of the genus Sphaerobelum Verhoeff, 1924 (Diplopoda, Sphaerotheriida, Zephroniidae) in Vietnam. Tap Chi Sinh Hoc, 41 (1), 9–17. https://doi.org/10.15625/0866-7160/v41n1.11400
- Nguyen, A.D., Sierwald, P. & Marek, P.E. (2019c) The pill millipedes of Vietnam: A key to genera and descriptions of five new species (Diplopoda: Glomerida: Glomeridae). Raffles Bulletin of Zoology, 67, 260–297. https://doi.org/10.26107/RBZ-2019-0020
- Nguyen, A.D., Stoev, P., Nguyen, L.T.P. & Vu, T.T. (2023) A new species of Paracortina from a Vietnamese cave, with remarkable secondary sexual characters in males (Callipodida, Paracortinidae). ZooKeys, 1149, 181–195. https://doi.org/10.3897/zookeys.1149.99651
- Nguyen, A.D., Stoev, P. & Vu, T.T.T. (2025a) Integrative data reveal a new millipede species of Sinocallipus Zhang, 1993 (Callipodida, Sinocallipodidae) from Vietnam, with notes on its phylogeny. Zoosystematics and Evolution, 101 (1), 69–80. https://doi.org/10.3897/zse.101.138716
- Nguyen, A.D., Tran, B.T.T. & Minh, D.L.E. (2018b) First record of the millipede genus Nesorthomorpha Jeekel, 1980 in Vietnam with description of a new species (Diplopoda, Polydesmida, Paradoxosomatidae). Zootaxa, 4462 (3), 426–434. https://doi.org/10.11646/zootaxa.4462.3.8
- Nguyen, A.D., Vu, T.T.T. & Eguchi, K. (2024b) The millipede family Polydesmidae Leach, 1816 (Diplopoda, Polydesmida) from Vietnam, with a description of a new cavernicolous species. ZooKeys, 1190, 259–280. https://doi.org/10.3897/zookeys.1190.114958
- Nguyen, A.D., Vu, T.T.T. & Nguyen, T.-A.T. (2025b) Mountainous millipedes in Vietnam. III. Two new dragon millipedes from limestone mountains in northern Vietnam (Polydesmida, Paradoxosomatidae, Hylomus), with an identification key to Vietnamese Hylomus species. ZooKeys, 1223, 247–262. https://doi.org/10.3897/zookeys.1223.139649
- Nguyen Duy-Jacquemin, M. & Condé, B. (1967) Morphologie et géonémie du gere Monographis Attems. itteilungen aus dem Hamburgischen zoologischen Museum und Institut, 64, 43–81.
- Nguyen Duy-Jacquemin, M. & Condé, B. (1982) Lophoproctidés insulaires de l’océan Pacifique (Diplopodes Pénicillates). Bulletin du Muséum national d’histoire naturelle, 4e Série, Section A, Zoologie, 4 (1–2), 95–118. https://doi.org/10.5962/p.286031
- Pimvichai, P., Akkari, N., Semenyuk, I., Constant, J. & Enghoff, H. (2020) A strikingly coloured new giant millipede from Vietnam has copycat in Borneo (Diplopoda, Spirostreptida, Harpagophoridae). European Journal of Taxonomy, 722, 1–15. https://doi.org/10.5852/ejt.2020.722.1129
- Pimvichai, P., Enghoff, H. & Panha, S. (2009a) A revision of the Thyropygus allevatus group. Part 1: the T. opinatus subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa, 2016 (1), 17–50. https://doi.org/10.11646/zootaxa.2016.1.2
- Pimvichai, P., Enghoff, H. & Panha, S. (2009b) A revision of the Thyropygus allevatus group. Part 2: the T. bifurcus subgroup (Diplopoda, Spirostreptida, Harpagophoridae). Zootaxa, 2165 (1), 1–15. https://doi.org/10.11646/zootaxa.2165.1.1
- Pimvichai, P., Enghoff, H. & Panha, S. (2010) The Rhynchoproctinae, a south-east Asiatic subfamily of giant millipedes: cladistic analysis, classification, four new genera and a deviating new species from north-west Thailand (Diplopoda:Spirostreptida:Harpagophoridae). Invertebrate Systematics, 24 (1), 51–80. https://doi.org/10.1071/IS09052
- Pimvichai, P., Enghoff, H. & Panha, S. (2011a) A revision of the Thyropygus allevatus group. Part 3: the T. induratus subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa, 2941 (1), 47–68. https://doi.org/10.11646/zootaxa.2941.1.3
- Pimvichai, P., Enghoff, H. & Panha, S. (2011b) A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa, 2980 (1), 37–48. https://doi.org/10.11646/zootaxa.2980.1.3
- Pimvichai, P., Enghoff, H. & Panha, S. (2014) Molecular phylogeny of the Thyropygus allevatus group of giant millipedes and some closely related groups. Molecular Phylogenetics and Evolution, 71, 170–183. https://doi.org/10.1016/j.ympev.2013.11.006
- Pimvichai, P., Enghoff, H., Panha, S. & Backeljau, T. (2018) Morphological and mitochondrial DNA data reshuffle the taxonomy of the genera Atopochetus Attems, Litostrophus Chamberlin and Tonkinbolus Verhoeff (Diplopoda: Spirobolida: Pachybolidae), with descriptions of nine new species. Invertebrate Systematics, 32 (1), 159. https://doi.org/10.1071/IS17052
- Pimvichai, P., Panha, S. & Backeljau, T. (2022) Combining mitochondrial DNA and morphological data to delineate four new millipede species and provisional assignment to the genus. Invertebrate Systematics, 36 (2), 91–112. https://doi.org/10.1071/IS21038
- Pocock, R.I. (1895a) Description of a new suctorial Millipede sent from Trinidad by Mr J. H. Hart of the Royal Botanica) Gardens, Trinidad. Trinidad. Annals and Magazine of Natural History, Series 6, 15 (88), 375. https://doi.org/10.1080/00222939508677900
- Pocock, R.I. (1895b) Descriptions of new genera of Zephroniidae, with brief preliminary Diagnoses of some new Species. Annals and Magazine of Natural History, Series 6, 16, 409–415. https://doi.org/10.1080/00222939508680293
- Pocock, R.L. (1893) Viaggio di Leonardo Fea in Birmania e regioni vicine. LV. On the Myriapoda of Burma. Pt. 3. Report upon the Julidae, Chordeumidae and Polyzonidae collected by Sig. L. Fea and Mr. E. W. Oates. Annali del Museo civico di storia naturale di Genova, Series 2, 13, 386–406.
- de Saussure, H.L.F. (1860) Essai d’une faune des Myriapodes du Mexique avec la description de quelques espèces des autres parties de l’Amérique. Mémoires de la Société de Physiques et d’Histoire naturelle de Genève, 15 (2), 259–394.
- Semenyuk, I., Golovatch, S.I. & Wesener, T. (2020) Some new or poorly-known Zephroniidae (Diplopoda, Sphaerotheriida) from Vietnam. ZooKeys, 930, 37–60. https://doi.org/10.3897/zookeys.930.47742
- Shear, W. (2000) A new genus and species of callipodidan milliped from Vietnam (Callipodida, Schizopetalidae). Myriapodologica, 6 (11), 95–100.
- Shear, W.A. (2002) The millipede genus Metopidiothrix Attems (Diplopoda : Chordeumatida : Metopidiotrichidae). Invertebrate Systematics, 16 (6), 849–892. https://doi.org/10.1071/IS02005
- Shelley, R.M. & Lehtinen, P.T. (1999) Diagnoses, synonymies and occurrences of the pantropical millipeds, Leptogoniulus sorornus (Butler) and Trigoniulus corallinus (Gervais) (Spirobolida: Pachybolidae: Trigoniulinae). Journal of Natural History, 33 (9), 1379–1401. https://doi.org/10.1080/002229399299932
- Sierwald, P. & Bond, J.E. (2007) Current status of the myriapod class Diplopoda (Millipedes): Taxonomic diversity and phylogeny. Annual Review of Entomology, 52, 401–420. https://doi.org/10.1146/annurev.ento.52.111805.090210
- Silvestri, F. (1910a) Descrizione di una nuova famiglia di Diplopoda Cambaloidea del Tonkino. Bollettino del Laboratorio di Zoologia generale e agraria della R. Scuola superiore d’ Agricoltura in Portici, 4, 66–70.
- Silvestri, F. (1910b) Descrizioni preliminari di novi generi di Diplopodi. Zoologischer Anzeiger, 35 (12–13), 3357–364.
- Silvestri, F. (1917) Contributions to a knowledge of the oriental Diplopoda Oniscomorpha. I, The family Glomeridae. Records of the Indian Museum, 13 (3.9), 103–151. https://doi.org/10.26515/rzsi/v13/i3/1917/163604
- Silvestri, F. (1923) Descriptions of some Indian and Malayan Myriapoda Cambaloidea. Records of the Indian Museum, 25, 181–193. https://doi.org/10.26515/rzsi/v25/i2/1923/162702
- Srisonchai, R., Enghoff, H., Likhitrakarn, N. & Panha, S. (2018) A revision of dragon millipedes I: genus Desmoxytes Chamberlin, 1923, with the description of eight new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys, 761, 1–177. https://doi.org/10.3897/zookeys.761.24214
- Sterling, E.J., Hurley, M.M. & Le, D.M. (2006) Vietnam a natural history. Yale University Press, New Haven, Connecticut, 423 pp.
- Stoev, P. & Enghoff, H. (2011) A review of the millipede genus Sinocallipus Zhang, 1993 (Diplopoda: Callipodida: Sinocallipodidae), with notes on gonopods monotony vs. peripheral diversity in millipedes. ZooKeys, 90, 13–34. https://doi.org/10.3897/zookeys.90.1291
- Stoev, P. & Geoffroy, J.-J. (2004) Review of the family Paracortinidae Wang & Zhang, 1993 (Diplopoda: Callipodida). Acta Arachnologica, 53 (2), 93–103. https://doi.org/10.2476/asjaa.53.93
- Verhoeff, K.W. (1915) Zur Kenntnis der Plesiocerata. (Über Diplopoden, 82. Aufsatz). Zoologischer Anzeiger, 46 (1), 16–48.
- Verhoeff, K.W. (1920) Chilognathen-Studien. (91. Diplopoden-Aufsatz). Archiv für Naturgeschichte, 86A (12), 23–80.
- Verhoeff, K.W. (1924) Results of Dr. E. Mjöberg’s Swedish Scientific Expeditions to Australia 1910-1913. 34. Myriapoda: Diplopoda. Arkiv för Zoologi, 16 (5), 1–142.
- Verhoeff, K.W. (1938) Über Diplopoden des Zoologischen Museums in München. Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere, 71 (1–2), 1–54.
- Wellman, C.H., Lopes, G., McKellar, Z. & Hartley, A. (2024) Age of the basal ‘Lower Old Red Sandstone’ Stonehaven Group of Scotland: the oldest reported air-breathing land animal is Silurian (late Wenlock) in age. Journal of the Geological Society, 181 (1), 1–7. https://doi.org/10.1144/jgs2023-138
- Wesener, T. (2016) The Giant Pill-Millipedes, order Sphaerotheriida – An annotated species catalogue with morphological atlas and list of apomorphies. Bonn zoological Bulletin, Supplementum, 63, 1–107.
- Wesener, T., Moritz, L. & Akkari, N. (2023) Integrative redescription of the sucking millipede genus Dawydoffia Attems, 1953, with a description of a new species and a transfer to the family Hirudisomatidae (Diplopoda, Polyzoniida). ZooKeys, 5263 (3), 411–429. https://doi.org/10.11646/zootaxa.5263.3.6
- Xiong, Y., Chen, H., Jiang, X. & Jiang, C. (2025) Topotypes of the millipede species Kronopolites swinhoei (Pocock, 1895) reveal a new synonym with revalidation of Kronopolites svenhedini (Verhoeff, 1934) (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys, 1231, 85–98. https://doi.org/10.3897/zookeys.1231.137769