Abstract
Scientific biological nomenclature underpins all knowledge exchange on biodiversity and works best when the names of organisms are stable. However, science-driven changes to zoological nomenclature are essential to reflect advances in knowledge. In contrast, process-driven changes resulting from historical discoveries followed by strict application of the Code can destabilise nomenclature without representing any gain in taxonomic knowledge. We illustrate this problem using the example of the European Nose-horned Viper, Vipera ammodytes (Linnaeus, 1758): its Linnaean type was recently reassessed and found not to originate from the western Balkans, as long assumed, but from near Istanbul, European Türkiye. The transfer of the name V. ammodytes ammodytes from its long-standing prevailing usage to the eastern subspecies widely known as V. a. montandoni Boulenger, 1904 would greatly impede the interpretation of a large body of literature and complicate communication about an iconic, widely known venomous snake of public health importance. We emphasise the imperative of nomenclatural parsimony (i.e., the need to minimise changes) when deciding between different possible courses of action required by historical discoveries. We urge researchers into nomenclatural history to safeguard long-established, widely used names, including through petitions to the International Commission on Zoological Nomenclature.
References
- Avella, I., Damm, M., Di Nicola, M.R., Dresler, J., İğci, N., Kariş, M., Kazemi, S.M., Kreuels, B., Paolino, G., Sarigiannis, Y., Vilcinskas, A., Wüster, W. & Lüddecke, T. (2025) The biology and toxinology of blunt-nosed vipers. npj Biodiversity, 4 (1), 21. https://doi.org/10.1038/s44185-025-00090-w
- Bauer, A.M. & Günther, R. (2013) Origin and identity of the von Borcke collection of amphibians and reptiles in the Museum für Naturkunde in Berlin: A cache of Seba specimens? Zoosystematics and Evolution, 89 (1), 167–185. https://doi.org/10.1002/zoos.201300005
- Böhme, W., Denzer, W. & Wüster, W. (in press) Noli tangere nomina probata! Taxonomists must prioritise stability and protection of well established scientific names in herpetology. Salamandra. [in press]
- Boulenger, G.A. (1896) Catalogue of the Snakes in the British Museum (Natural History). Vol. III. Containing the Colubridae (Opisthoglyphae and Proteroglyphae), Amblycephalidae, and Viperidae. British Museum (Natural History), London, xiv + 727 pp. https://doi.org/10.5962/bhl.title.54273
- Boulenger, G.A. (1903) On the geographical variants of the sand-viper, Vipera ammodytes. Proceedings of the Zoological Society of London, 1903 (1), 185–186. https://doi.org/10.1111/j.1469-7998.1903.tb08276.x
- Boulenger, G.A. (1904) On the sand-viper of Roumania (Vipera ammodytes var. Montandoni). Annals and Magazine of Natural History, Series 7, 14 (80), 134–135. https://doi.org/10.1080/03745480409442981
- Boulenger, G.A. (1913) On the geographical races of Vipera ammodytes. Annals and Magazine of Natural History, 11 (63), 283–297, 5 pl. https://doi.org/10.1080/00222931308693322
- Bruno, S. (1968) Sulla Vipera ammodytes (Linnaeus, 1758) in Italia. Memorie del Museo Civico di Storia Naturale di Verona, 15, 289–336.
- Carrasco, P.A., Venegas, P.J., Chaparro, J.C. & Scrocchi, G.J. (2016) Nomenclatural instability in the venomous snakes of the Bothrops complex: Implications in toxinology and public health. Toxicon, 119, 122–128. https://doi.org/10.1016/j.toxicon.2016.05.014
- Cattaneo, A. (2021) Variabilità di Vipera ammodytes (Linnaeus, 1758) (Reptilia Viperidae) in alcune isole egee, con descrizione di Vipera ammodytes buchholzi subsp. nova. Naturalista Siciliano, 45 (1–2), 119–152. https://doi.org/10.5281/ZENODO.5713465
- David, P. & Ineich, I. (1999) Les serpents venimeux du monde: systématique et répartition. Dumerilia, 3, 3–499.
- De Carvalho, M.R., Bockmann, F.A., Amorim, D.S., De Vivo, M., De Toledo-Piza, M., Menezes, N.A., De Figueiredo, J.L., Castro, R.M.C., Gill, A.C., McEachran, J.D., Compagno, L.J.V., Schelly, R.C., Britz, R., Lundberg, J.G., Vari, R.P. & Nelson, G. (2005) Revisiting the taxonomic impediment. Science, 307 (5708), 353–353. https://doi.org/10.1126/science.307.5708.353b
- Denzer, W., Dondorp, E., Kaiser, H. & Wüster, W. (2025) Notes on the type series of the Javan Spitting Cobra, Naja sputatrix Boie, 1827, and designation of a lectotype for Naja tripudians sondaica Schlegel, 1844. Zootaxa, 5679 (4), 585–599. https://doi.org/10.11646/zootaxa.5679.4.7
- Di Nicola, M.R., Pontara, A., Kass, G.E.N., Kramer, N.I., Avella, I., Pampena, R., Mercuri, S.R., Dorne, J.L.C.M. & Paolino, G. (2021) Vipers of major clinical relevance in Europe: taxonomy, venom composition, toxicology and clinical management of human bites. Toxicology, 453, 152724. https://doi.org/10.1016/j.tox.2021.152724
- Dobaja Borak, M., Babić, Ž., Caganova, B., Grenc, D., Karabuva, S., Kolpach, Z., Krakowiak, A., Kolesnikova, V., Lukšić, B., Pap, C., Puljiz, I., Piekarska-Wijatkowska, A., Radenkova-Saeva, J., Vučinić, S., Zacharov, S., Eddleston, M. & Brvar, M. (2023) Viper envenomation in Central and Southeastern Europe: a multicentre study. Clinical Toxicology, 61 (9), 656–664. https://doi.org/10.1080/15563650.2023.2273761
- Dubois, A. (2010) Zoological nomenclature in the century of extinctions: priority vs. ‘usage’. Organisms Diversity & Evolution, 10 (3), 259–274. https://doi.org/10.1007/s13127-010-0021-3
- Dubois, A., Denzer, W., Entiauspe-Neto, O.M., Frétey, T., Ohler, A., Bauer, A.M. & Pyron, R.A. (2024) Nomenclatural problems raised by the recent description of a new anaconda species (Squamata, Serpentes, Boidae), with a nomenclatural review of the genus Eunectes. Bionomina, 37, 8–58. https://doi.org/10.11646/bionomina.37.1.2
- Dufresnes, C., Ghielmi, S., Halpern, B., Martínez-Freiría, F., Mebert, K., Jelić, D., Crnobrnja-Isailović, J., Gippner, S., Jablonski, D., Joger, U., Laddaga, L., Petrovan, S., Tomović, L., Vörös, J., İğci, N., Kariş, M., Zinenko, O. & Ursenbacher, S. (2024) Phylogenomic insights into the diversity and evolution of Palearctic vipers. Molecular Phylogenetics and Evolution, 197, 108095. https://doi.org/10.1016/j.ympev.2024.108095
- Dunn, E.R. & Stuart, L.C. (1951) On the legality of restriction of type locality. Science, 113 (2946), 677–678. https://doi.org/10.1126/science.113.2946.677
- Ebach, M.C., Valdecasas, A.G. & Wheeler, Q.D. (2011) Impediments to taxonomy and users of taxonomy: accessibility and impact evaluation. Cladistics, 27 (5), 550–557. https://doi.org/10.1111/j.1096-0031.2011.00348.x
- Ferquel, E., de Haro, L., Jan, V., Guillemin, I., Jourdain, S., Teynié, A., d’Alayer, J. & Choumet, V. (2007) Reappraisal of Vipera aspis venom neurotoxicity. PLoS ONE, 2 (11), e1194. https://doi.org/10.1371/journal.pone.0001194
- Fritz, U. & Schmidtler, J.F. (2020) The Fifth Labour of Heracles: Cleaning the Linnean stable of names for grass snakes (Natrix astreptophora, N. helvetica, N. natrix sensu stricto). Vertebrate Zoology, 70 (4), 621–665. https://doi.org/10.26049/VZ70-4-2020-07
- García-Arredondo, A., Martínez, M., Calderón, A., Saldívar, A. & Soria, R. (2019) Preclinical assessment of a new polyvalent antivenom (Inoserp Europe) against several species of the subfamily Viperinae. Toxins, 11 (3), 149. https://doi.org/10.3390/toxins11030149
- Garnett, S.T. & Christidis, L. (2017) Taxonomy anarchy hampers conservation. Nature News, 546 (7656), 25. https://doi.org/10.1038/546025a
- Garnett, S.T., Christidis, L., Conix, S., Costello, M.J., Zachos, F.E., Bánki, O.S., Bao, Y., Barik, S.K., Buckeridge, J.S., Hobern, D., Lien, A., Montgomery, N., Nikolaeva, S., Pyle, R.L., Thomson, S.A., van Dijk, P.P. & Whalen, A., Zhang, Z.-Q. & Thiele, K.R. (2020) Principles for creating a single authoritative list of the world’s species. PLoS Biology, 18 (7), e3000736. https://doi.org/10.1371/journal.pbio.3000736
- Geniez, P. (2018) Snakes of Europe, North Africa & the Middle East: A Photographic Guide. Princeton University Press, Princeton, New Jersey, 379 pp.
- Georgieva, D., Risch, M., Kardas, A., Buck, F., Von Bergen, M. & Betzel, C. (2008) Comparative analysis of the venom proteomes of Vipera ammodytes ammodytes and Vipera ammodytes meridionalis. Journal of Proteome Research, 7 (3), 866–886. https://doi.org/10.1021/pr070376c
- Giangrande, A. (2003) Biodiversity, conservation, and the ‘taxonomic impediment’. Aquatic Conservation: Marine and Freshwater Ecosystems, 13 (5), 451–459. https://doi.org/10.1002/aqc.584
- Golay, P., Smith, H.M., Broadley, D.G., Dixon, J.R., McCarthy, C.J., Rage, J.-C., Schätti, B. & Toriba, M. (1993) Endoglyphs and Other Major Venomous Snakes of the World: a Checklist. Azemiops, Geneva, Switzerland, 478 pp.
- Happ, H. & Wieser, D. (2008) Die Hornotter Vipera ammodytes (Linnaeus, 1758) – in Kärnten vom Aussterben bedroht! Ist Hilfe noch möglich? Carinthia II, 198 (118), 83–94.
- Happ, U., Smole-Wiener, A.K. & Gutleb, B. (1999) Rote Liste der Kriechtiere Kärntens (Vertebrata: Reptilia). Publikationen Naturschutz Kärnten, 1999, 113–116.
- Heckes, U., Gruber, H.-J. & Stümpel, N. (2005) Vipera (Vipera) ammodytes (Linnaeus, 1758) – Hornotter, Sandviper. In: Joger, U. & Stümpel, N. (Eds.), Handbuch der Reptilien und Amphibien Europas. Schlangen (Serpentes) III. AULA-Verlag, Wiebelsheim, pp. 81–150.
- Hillis, D.M. (2019) Species delimitation in herpetology. Journal of Herpetology, 53 (1), 3–12. https://doi.org/10.1670/18-123
- Hillis, D.M. (2020) The detection and naming of geographic variation within species. Herpetological Review, 51 (1), 52–56.
- Holycross, A.T., Anton, T.G., Douglas, M.E. & Frost, D.R. (2008) The type localities of Sistrurus catenatus and Crotalus viridis (Serpentes: Viperidae), with the unraveling of a most unfortunate tangle of names. Copeia, 2008 (2), 421–424. https://doi.org/10.1643/CH-07-095
- International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. 4th Edition. The International Trust for Zoological Nomenclature, London, 306 pp. https://doi.org/10.5962/bhl.title.50608
- International Commission on Zoological Nomenclature (2012) Amendment of Articles 8, 9, 10, 21 and 78 of the International Code of Zoological Nomenclature to expand and refine methods of publication. The Bulletin of Zoological Nomenclature, 69, 161–169. https://doi.org/10.21805/bzn.v69i3.a8.161
- International Commission on Zoological Nomenclature (2018) Opinion 2426 (Case 3703) – Nautilus pompilius Linnaeus, 1758 (Mollusca, Cephalopoda, Nautilida): neotype designated. The Bulletin of Zoological Nomenclature, 75 (1), 288. https://doi.org/10.21805/bzn.v75.a063
- International Commission on Zoological Nomenclature (2021) Opinion 2468 (Case 3601) – Spracklandus Hoser, 2009 (Reptilia, Serpentes, Elapidae) and Australasian Journal of Herpetology issues 1–24: confirmation of availability declined; Appendix A (Code of Ethics): not adopted as a formal criterion for ruling on Cases. The Bulletin of Zoological Nomenclature, 78 (1), 42–45. https://doi.org/10.21805/bzn.v78.a012
- Jackson, M.D., Scherz, M.D. & Zona, S. (2017) Taxonomy is not beholden to its dependencies: a rebuttal to Garnett and Christidis (2017). PeerJ Preprints, 5, e3060v1. https://doi.org/10.7287/peerj.preprints.3060v1
- Jiménez-Mejías, P., Manzano, S., Gowda, V., Krell, F.-T., Lin, M.-Y., Martín-Bravo, S., Martín-Torrijos, L., Nieto Feliner, G., Mosyakin, S.L., Naczi, R.F.C., Acedo, C., Álvarez, I., Crisci, J.V., Luceño Garcés, M., Manning, J., Moreno Saiz, J.C., Muasya, A.M., Riina, R., Meseguer, A.S., Sánchez-Mata, D. & 1543 additional coauthors (2024) Protecting stable biological nomenclatural systems enables universal communication: A collective international appeal. BioScience, 74 (7), 467–472. [an additional 1543 coauthors are listed in Aupplemntary_1_Revised_2 file: https://academic.oup.com/bioscience/article/74/7/467/7696204#supplementary-data]
- Klemmer, K. (1963) Liste der rezenten Giftschlangen. In: Die Giftschlangen der Erde. Behringwerke-Mitteilungen. Sonderband. N.G. Elwert Universitäts- und Verlagsbuchhandlung, Marburg, pp. 255–464.
- Krecsák, L., Bauer, A.M., Tomović, L., Schweiger, S. & Westerström, A. (2025a) The history of the Illyrian nose-horned Viper, Vipera ammodytes illyrica (Laurenti, 1768), with designation of a neotype. Salamandra, 61 (2), 184–194.
- Krecsák, L., Bauer, A.M., Tomović, L., Schweiger, S. & Westerström, A. (2025b) A valid neotype designation for Vipera illyrica Laurenti, 1768. Salamandra, 61 (3), 362–363.
- Krecsák, L., Bauer, A.M., Westerström, A., Wahlgren, R., Tomović, L., Stille, B. & Åhlander, E. (2024) Assessment of the Linnaean type material of the nose-horned viper, Vipera ammodytes (Linnaeus, 1758). Zootaxa, 5537 (1), 24–48. https://doi.org/10.11646/zootaxa.5537.1.2
- Križaj, I. (2011) Ammodytoxin: A window into understanding presynaptic toxicity of secreted phospholipases A2 and more. Toxicon, 58 (3), 219–229. https://doi.org/10.1016/j.toxicon.2011.06.009
- Lamb, T., De Haro, L., Lonati, D., Brvar, M. & Eddleston, M. (2017) Antivenom for European Vipera species envenoming. Clinical Toxicology, 55 (6), 557–568. https://doi.org/10.1080/15563650.2017.1300261
- Laurenti, J.N. (1768) Austriaci viennensis Specimen medicum, exhibens synopsin reptilium emendatam cum experimentis circa venena et antidota reptilium austriacorum. Johan. Thom Nob. de Trattnern, Viennae, 214 pp. https://doi.org/10.5962/bhl.title.5108
- Lien, A.M., Banki, O., Barik, S.K., Buckeridge, J.S., Christidis, L., Cigliano, M.M., Conix, S., Costello, M.J., Hobern, D., Kirk, P.M., Kroh, A., Montgomery, N., Nikolaeva, S., Orrell, T.M., Pyle, R.L., Raz, L., Thiele, K., Thomson, S.A., Van Dijk, P.P., Wambiji, N., Whalen, A., Zachos, F.E., Zhang, Z.-Q. & Garnett, S.T. (2023) Widespread support for a global species list with a formal governance system. Proceedings of the National Academy of Sciences, 120 (45), e2306899120. https://doi.org/10.1073/pnas.2306899120
- Linnaeus, C. (1758) Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Tomus I. 10th Edition. Laurentii Salvii, Stockholm, 824 pp. https://doi.org/10.5962/bhl.title.542
- Löbl, I., Klausnitzer, B., Hartmann, M. & Krell, F.-T. (2023) The silent extinction of species and taxonomists—an appeal to science policymakers and legislators. Diversity, 15 (10), 1053. https://doi.org/10.3390/d15101053
- Mayr, E., Simpson, G.G. & Eisenmann, E. (1971) Stability in zoological nomenclature. Science, 174 (4013), 1041–1042. https://doi.org/10.1126/science.174.4013.1041.b
- Mertens, R. & Müller, L. (1928) Liste der Amphibien und Reptilien Europas. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 41 (1), 3–62.
- Molina, M. (2001) More notes on the Verreaux brothers. Pula: Botswana Journal of African Studies, 16 (1), 30–36.
- Nikolaeva, S.V., Saunders, W.B., Mapes, R. & Allcock, A.L. (2015) Case 3703 Nautilus pompilius Linnaeus, 1758 (Mollusca, Cephalopoda, Nautilida): proposed designation of a neotype. The Bulletin of Zoological Nomenclature, 72 (4), 274–285. https://doi.org/10.21805/bzn.v72i4.a21
- Nilson, G., Tuniyev, B., Andrén, C. & Orlov, N. (1999) Vipers of the Caucasus: taxonomic considerations. Kaupia Darmstädter Beiträge zur Naturgeschichte, 8, 103–106.
- O’Shea, M. & Kaiser, H. (2018) Erroneous environs or aberrant activities? Reconciling unexpected collection localities for three New Guinea Worm-eating Snakes (Toxicocalamus, Serpentes, Elapidae) using historical accounts. Herpetological Review, 49, 189–207.
- Osipov, A. & Utkin, Y. (2023) What are the neurotoxins in hemotoxic snake venoms? International Journal of Molecular Sciences, 24 (3), 2919. https://doi.org/10.3390/ijms24032919
- Petrova, S.D., Atanasov, V.N. & Balashev, K. (2012) Vipoxin and Its components: structure-function relationship. In: Christov, C. & Karabencheva-Christova, T. (Eds.), Advances in Protein Chemistry and Structural Biology. Academic Press, San Diego, pp. 117–153. https://doi.org/10.1016/B978-0-12-398312-1.00005-6
- Plasinger, I., Righetti, D. & Di Cerbo, A.R. (2016) La conservazione della vipera dal corno, Vipera ammodytes, in Alto Adige: analisi dei fattori di minaccia con indicazioni di gestione dell’habitat (Reptilia: Viperidae) In: Bonato, L., Trabucco, R. & Bon, M. (Eds.), Atti 7° Convegno Faunisti Veneti, Verona, 15–16 Novembre 2014. Bolletino del Museo di Storia Naturale di Venezia, 66 (Supplement), pp. 116–122.
- Radonić, V., Budimir, D., Bradarić, N., Lukšić, B., Sapunar, D. & Vilović, K. (1997) Envenomation by the horned viper (Vipera ammodytes). Military Medicine, 162 (3), 179–182. https://doi.org/10.1093/milmed/162.3.179
- Rösler, H., Grismer, L.L., Ineich, I. & Kaiser, H. (2019) The purportedly Indo-Australian gecko species Cnemaspis timoriensis (Duméril & Bibron, 1836) is actually the first named species of the neotropical genus Gonatodes Fitzinger, 1843 (Squamata: Gekkonidae). Zootaxa, 4576 (3), 483–509. https://doi.org/10.11646/zootaxa.4576.3.4
- Rösler, H., Ineich, I., Ávila-Pires, T.C.S., Hoogmoed, M.S. & Kaiser, H. (2025) Case 3798. Gymnodactylus humeralis Guichenot, 1855 (currently Gonatodes humeralis; Reptilia, Sphaerodactylidae): proposed conservation of the specific name by suppression of Gymnodactylus timoriensis Duméril & Bibron, 1836. Bulletin of Zoological Nomenclature. [in press]
- Roussos, S.A. (2015) Integrative Evolutionary Biogeography of the Nose-horned Viper (Viperidae: Squamata) in the Cycladic Archipelago and Continental Greece. Unpublished PhD Thesis, Texas Tech University, Lubbock, Texas, 172 pp.
- Schwarz, E. (1936) Untersuchungen über die Systematik und Verbreitung der europäischen und mediterranen Ottern. In: Bieling, R., Demnitz, A., Schaumann, O., Schloßberger, H., von Schuckmann, W. & Schwarz, E. (Eds.), Die Europäischen und Mediterranen Ottern und Ihre Gifte. Vol. 7. Behringwerk-Mitteilungen. Selbstverlag der Behringwerke, Marburg-Lahn, pp. 159–355, 1 map.
- Singh, N. (2025) Taxonomy in trouble—an impediment to life on Earth. Zootaxa, 5642 (3), 395–400. https://doi.org/10.11646/zootaxa.5642.3.8
- Sochurek, E. (1974) Vipera ammodytes gregorwallneri n. ssp. Herpetologische Blätter, 1, 1–4.
- Thanou, E., Jablonski, D. & Kornilios, P. (2023) Genome‐wide single nucleotide polymorphisms reveal recurrent waves of speciation in niche‐pockets, in Europe’s most venomous snake. Molecular Ecology, 32 (13), 3624–3640. https://doi.org/10.1111/mec.16944
- Thiele, K.R., Conix, S., Pyle, R.L., Barik, S.K., Christidis, L., Costello, M.J., van Dijk, P.P., Kirk, P., Lien, A. & Thomson, S.A., Zachos, F.E. & Zhang, Z.-Q. & Garnett, S.T. (2021) Towards a global list of accepted species. I. Why taxonomists sometimes disagree, and why this matters. Organisms Diversity & Evolution, 21 (4), 615–622. https://doi.org/10.1007/s13127-021-00495-y
- Thomson, S.A., Pyle, R.L., Ahyong, S.T., Alonso-Zarazaga, M., Ammirati, J., Araya, J.F., Ascher, J.S., Audisio, T.L., Azevedo-Santos, V.M., Bailly, N., Baker, W.J., Balke, M., Barclay, M.V.L., Barrett, R.L., Benine, R.C., Bickerstaff, J.R.M., Bouchard, P., Bour, R., Bourgoin, T., Boyko, C.B., Breure, A.S.H., Brothers, D.J., Byng, J.W., Campbell, D., Ceríaco, L.M.P., Cernák, I., Cerretti, P., Chang, C.-H., Cho, S., Copus, J.M., Costello, M.J., Cseh, A., Csuzdi, C., Culham, A., D’Elía, G., d’Udekem d’Acoz, C., Daneliya, M.E., Dekker, R., Dickinson, E.C., Dickinson, T.A., van Dijk, P.P., Dijkstra, K.-D.B., Dima, B., Dmitriev, D.A., Duistermaat, L., Dumbacher, J.P., Eiserhardt, W.L., Ekrem, T., Evenhuis, N.L., Faille, A., Fernández-Triana, J.L., Fiesler, E., Fishbein, M., Fordham, B.G., Freitas, A.V.L., Friol, N.R., Fritz, U., Frøslev, T., Funk, V.A., Gaimari, S.D., Garbino, G.S.T., Garraffoni, A.R.S., Geml, J., Gill, A.C., Gray, A., Grazziotin, F.G., Greenslade, P., Gutiérrez, E.E., Harvey, M.S., Hazevoet, C.J., He, K., He, X., Helfer, S., Helgen, K.M., van Heteren, A.H., Hita Garcia, F., Holstein, N., Horváth, M.K., Hovenkamp, P.H., Hwang, W.S., Hyvönen, J., Islam, M.B., Iverson, J.B., Ivie, M.A., Jaafar, Z., Jackson, M.D., Jayat, J.P., Johnson, N.F., Kaiser, H., Klitgård, B.B., Knapp, D.G., Kojima, J., Kõljalg, U., Kontschán, J., Krell, F.-T., Krisai-Greilhuber, I., Kullander, S., Latella, L., Lattke, J.E., Lencioni, V., Lewis, G.P., Lhano, M.G., Lujan, N.K., Luksenburg, J.A., Mariaux, J., Marinho-Filho, J., Marshall, C.J., Mate, J.F., McDonough, M.M., Michel, E., Miranda, V.F.O., Mitroiu, M.-D., Molinari, J., Monks, S., Moore, A.J., Moratelli, R., Murányi, D., Nakano, T., Nikolaeva, S., Noyes, J., Ohl, M., Oleas, N.H., Orrell, T., Páll-Gergely, B., Pape, T., Papp, V., Parenti, L.R., Patterson, D., Pavlinov, I.Ya., Pine, R.H., Poczai, P., Prado, J., Prathapan, D., Rabeler, R.K., Randall, J.E., Rheindt, F.E., Rhodin, A.G.J., Rodríguez, S.M., Rogers, D.C., Roque, F. de O., Rowe, K.C., Ruedas, L.A., Salazar-Bravo, J., Salvador, R.B., Sangster, G., Sarmiento, C.E., Schigel, D.S., Schmidt, S., Schueler, F.W., Segers, H., Snow, N., Souza-Dias, P.G.B., Stals, R., Stenroos, S., Stone, R.D., Sturm, C.F., Štys, P., Teta, P., Thomas, D.C., Timm, R.M., Tindall, B.J., Todd, J.A., Triebel, D., Valdecasas, A.G., Vizzini, A., Vorontsova, M.S., de Vos, J.M., Wagner, P., Watling, L., Weakley, A., Welter-Schultes, F., Whitmore, D., Wilding, N., Will, K., Williams, J., Wilson, K., Winston, J.E., Wüster, W., Yanega, D., Yeates, D.K., Zaher, H., Zhang, G., Zhang, Z.-Q. & Zhou, H.-Z. (2018) Taxonomy based on science is necessary for global conservation. PLoS Biology, 16 (3), e2005075. https://doi.org/10.1371/journal.pbio.2005075
- Tomović, L. (2006) Systematics of the nose-horned viper (Vipera ammodytes, Linnaeus, 1758). Herpetological Journal, 16, 191–201.
- Tudor, M. (2010) The present situation of the nose-horned viper populations (Vipera ammodytes montandoni) Boulenger 1904 from Dobrudja (Romania and Bulgaria). Ovidius University Annals of Natural Sciences, Biology, Ecology Series, 14, 113–118.
- Ursenbacher, S., Schweiger, S., Tomović, L., Crnobrnja-Isailović, J., Fumagalli, L. & Mayer, W. (2008) Molecular phylogeography of the nose-horned viper (Vipera ammodytes, Linnaeus (1758)): Evidence for high genetic diversity and multiple refugia in the Balkan Peninsula. Molecular Phylogenetics and Evolution, 46 (3), 1116–1128. https://doi.org/10.1016/j.ympev.2007.11.002
- Vences, M., Guayasamin, J.M., Miralles, A. & Riva, I.D.L. (2013) To name or not to name: criteria to promote economy of change in Linnaean classification schemes. Zootaxa, 3636 (2), 201–244. https://doi.org/10.11646/zootaxa.3636.2.1
- World Health Organization (2013) Guidelines for the production, control and regulation of snake antivenom immunoglobulins, Annex 5. Replacement of Annex 2 of WHO Technical Report Series, No. 964. WHO Technical Report Series, 1003, 197–388.
- Wüster, W., Kaiser, H., Hoogmoed, M.S., Ceríaco, L.M.P., Dirksen, L., Dufresnes, C., Glaw, F., Hille, A., Köhler, J., Koppetsch, T., Milto, K.D., Shea, G.M., Tarkhnishvili, D., Thomson, S.A., Vences, M. & Böhme, W. (2024) How not to describe a species: lessons from a tangle of anacondas (Boidae: Eunectes Wagler, 1830). Zoological Journal of the Linnean Society, 201 (4), zlae099. https://doi.org/10.1093/zoolinnean/zlae099
- Wüster, W. & McCarthy, C.J. (1996) Venomous snake systematics: implications for snake bite treatment and toxinology. In: Bon, C. & Goyffon, M. (Eds.), Envenomings and the Treatments. Fondation Marcel Mérieux, Lyon, pp. 13–23. https://doi.org/10.1016/0041-0101(96)83649-X
- Wüster, W., Thomson, S.A., O’Shea, M. & Kaiser, H. (2021) Confronting taxonomic vandalism in biology: conscientious community self-organization can preserve nomenclatural stability. Biological Journal of the Linnean Society, 133 (3), 645–670. https://doi.org/10.1093/biolinnean/blab009
- Wüster, W. & Tillack, F. (2023) On the importance of types and the perils of ‘en passant’ taxonomy: a brief history of the typification of Coluber naja Linnaeus, 1758 (Serpentes: Elapidae) and its implications, with the designation of a lectotype. Zootaxa, 5346 (4), 403–419. https://doi.org/10.11646/zootaxa.5346.4.3
- Zhou, Z., Newman, C., Buesching, C.D., Meng, X., Macdonald, D.W. & Zhou, Y. (2016) Revised taxonomic binomials jeopardize protective wildlife legislation. Conservation Letters, 9 (5), 313–315. https://doi.org/10.1111/conl.12289