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Type: Monograph
Published: 2025-10-08
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The solitary vespid wasps of Madagascar (Hymenoptera: Vespidae: Eumeninae, Raphiglossinae and Zethinae)

Via dei Tarquini; 22—01100 Viterbo; Italy
Hymenoptera solitary wasps Africa Malagasy subregion taxonomy new synonymy

Abstract

The Malagasy fauna of Eumeninae sensu lato (= Eumeninae, Raphiglossinae and Zethinae) is revised, with identification keys, photographic reference to all species and distribution maps. Thirty-three new species are described: Afrepipona dilaticlypeus sp. nov., Afrepipona ennesima sp. nov., Afrepipona imitans sp. nov., Afrepipona misinjo sp. nov., Afrepipona resoluta sp. nov., Alastor (Alastorellus) angulatus sp. nov., Alastor (Alastorellus) curvatus sp. nov., Cyrteumenes deplanatus sp. nov., Cyrteumenes laevifrons sp. nov., Cyrteumenes propodealis sp. nov., Cyrtolabulus emirufus sp. nov., Cyrtolabulus nivosus sp. nov., Cyrtolabulus oblongus sp. nov., Cyrtolabulus obscurior sp. nov., Cyrtolabulus tegularis sp. nov., Eumenidiopsis pyriformis sp. nov., Leptomenes globosus sp. nov., Leptomenes multipictus sp. nov., Leptomenes ornatus sp. nov., Leptomenes sollicitus sp. nov., Leptomenes transuo sp. nov., Leptomenes tricolor sp. nov., Malagassodynerus concolor sp. nov., Malagassodynerus monstruosus sp. nov., Malagassodynerus pulcher sp. nov., Malgachemenes aberrans sp. nov., Malgachemenes albellus sp. nov., Malgachemenes divinus sp. nov., Malgachemenes levigatus sp. nov., Malgachemenes lobatus sp. nov., Tuleara alboviridis sp. nov., Tuleara elongata sp. nov., and Tuleara pruinosa sp. nov. The following taxonomic actions are proposed: Odynerus vatondrangyensis Giordani Soika, 1941 (= Simplepipona andreanicolor Gusenleitner, 2004 syn. nov.) is transferred to Antepipona de Saussure, 1855 (= Postepipona Giordani Soika, 1974, syn. nov., = Simplepipona Gusenleitner, 2004, syn. nov.; Antepipona vatondrangyensis comb. nov.); Odynerus pensus Giordani Soika, 1937 is transferred to Knemodynerus Blüthgen, 1940 (Knemodynerus pensus comb. nov.); Odynerus subventricosus Giordani Soika, 1941 is transferred to Malagassodynerus Gusenleitner, 1992 (= Giordania Gusenleitner, 1995, syn. nov.; Malagassodynerus subventricosus comb. nov.); Odynerus sakalavus de Saussure, 1890 is transferred to Tuleara Gusenleitner, 2000 (= Immutatus Gusenleitner, 2011, syn. nov.; Tuleara sakalava comb. nov.); Malgachemenes Gusenleitner, 1992, stat. resurr. is withdrawn from synonymy and reestablished as a distinct genus; Giordania nigra Gusenleitner, 1995, Odynerus hyalinelamellatus Giordani Soika, 1941, Odynerus melanus de Saussure, 1853 (= Odynerus malagassus de Saussure, 1900, syn. nov.) and Odynerus parvilamellatus Giordani Soika, 1941 are considered incertae sedis; Labus floricola de Saussure, 1890, Leptomenes laethificus Giordani Soika, 1941, Odynerus adulescens Giordani Soika, 1941, Odynerus antisrabensis Giordani Soika, 1941, Odynerus puer Giordani Soika, 1941 and Odynerus viridipictus Giordani Soika, 1941 are designated as nomina dubia. The females of Afrepipona segregata Selis & Carpenter, 2024 and Tuleara nigrita Gusenleitner, 2005 stat. nov., and the males of Antepipona declarata Giordani Soika, 1991, Cyrtolabulus flavorufus Selis, 2024, Cyrtolabulus rhombicus Gusenleitner, 2004 and Malgachemenes angustus Gusenleitner, 1992 are described for the first time. The genera Antodynerus de Saussure, 1855 and Eumenidiopsis Giordani Soika, 1939 are recorded from Madagascar for the first time. The present revision brings the total of Eumeninae s. l. known for Madagascar to 76 species.

 

References

  1. Ashmead, W.H. (1902) Classification of the fossorial, predacious and parasitic wasps, or the superfamily Vespoidea. (Paper no. 7). Family XXIX. Eumenidae. Canadian Entomologist, 34, 203–210. https://doi.org/10.4039/Ent34203-8
  2. Atanassov, N. (1967) Eine neue Untergattung und Art der Eumeniden (Hymenoptera: Diploptera) aus der bulgarischen Fauna. Izvestiya na Zoologicheskiya Institut Sofiya, 23, 167–174.
  3. Bequaert, J. (1918) A revision of the Vespidae of the Belgian Congo based on the Collection of the American Museum Congo Expedition, with a list of the Ethiopian Diplopterous Wasps. Bulletin of the American Museum of Natural History, 39 (1), 1–384.
  4. Bequaert, J. (1925) Eumenes dyscherus H. de Saussure, a Neotropical, not an African wasp, and other notes on synonymy. Bulletin of the Brooklyn Entomological Society, 20, 134–140.
  5. Bequaert, J. (1926) The genus Eumenes, Latreille, in South Africa, with a revision of the Ethiopian species (Hymenoptera). Annals of the South African Museum, 23, 483–577.
  6. Blüthgen, P. (1938 [1937]) Systematisches Verzeichnis der Faltenwespen Mitteleuropas, Scandinaviens und Englands. Konowia, 16, 270–295.
  7. Blüthgen, P. (1940) Über Odynerus meyeri CAM., Od. meyeri CAM. var. albolimbatus SCHULTHESS und Od. pseudolateralis M.W. (Hym. Vespidae). Entomologisk Tidskrift, 61 (1–2), 41–45.
  8. Blüthgen, P. (1958) Oplomerocephalie, eine pathologische Erscheinungsform bei gewissen Eumeniden (Hym., Dipl.). Zoologischer Anzeiger, 160, 36–38.
  9. Blüthgen, P. (1961) Die Faltenwespen Mitteleuropas (Hymenoptera, Diploptera). Abhandlungen der Deutschen Akademie der Wissenschaften zu Berlin, 1961 (2), 1–251. https://doi.org/10.1515/9783112537084
  10. Blommers, L.H.M. (2012) Taxonomy and natural history of 18 Ropalidia species (Hymenoptera, Vespidae) of Madagascar. Tijdschrift voor Entomologie, 155 (2–3), 133–192. https://doi.org/10.1163/22119434-00002010
  11. Blüthgen, P. (1956) Untersuchungen über palaearktische Eumenidae (Hym. Diploptera) II. Die Untergattung Alastor Lep. und Parastalor Blüth. der Gattung Alastor Lep. Mitteilungen aus dem Zoologische Museum zu Berlin, 32, 120–149.
  12. Bohart, R.M. & Stange, L.A. (1965) A Revision of the genus Zethus Fabricius in the Western Hemisphere (Hymenoptera: Eumenidae). University of California Publications in Entomology, 40, 1–208.
  13. Borsato, W. (1999) Eumenidi raccolti in Madagascar da A. Mochi (Hym., Eumenidae). Bollettino del Museo Regionale di Scienze Naturali di Torino, 16 (1–2), 107–112.
  14. Brancsik, K. (1891) Beiträge zur Kenntniss Nossibé’s und dessen Fauna nach Sendungen und Mittheilungen des Herrn P. Frey. Jahresheft des Naturwissenschaftlichen Vereins des Trencsiner Comitates, 14, 123–167.
  15. Cameron, P. (1910) Vespidae. In: Sjöstedt, Y. (Ed.), Wissenschaftliche Ergebnisse der Schwedischen Zoologischen Expedition nach dem Kilimandjaro, dem Meru und den umgebenden Massaisteppen Deutsch-Ostafrikas 1905-1906 unter der Leitung von Prof. Dr. Yngve Sjöstedt. Vol. 2. P. Palmquists Aktiebolag, Stockholm, pp. 169–196.
  16. Carpenter, J.M. & Madl, M. (2009) A Catalogue of the Vespidae of the Malagasy Subregion (Insecta, Hymenoptera). Linzer biologische Beiträge, 41 (2), 1871–1935.
  17. Carpenter, J.M., Gusenleitner, J. & Madl, M. (2010) A Catalogue of the Eumeninae (Hymenoptera: Vespidae) of the Ethiopian Region excluding Malagasy Subregion. Part II: Genera Delta de Saussure 1885 to Zethus Fabricius 1804 and Species Incertae Sedis. Linzer biologische Beiträge, 42 (1), 95–315.
  18. Christ, J.L. (1791) Naturgeschichte, Klassification und Nomenclatur der Insekten, vom Bienen, Wespen und Ameisengeschlecht; als der fünften Klasse fünfte Ordnung des Linneischen Natursystems von den Insekten: Hymenoptera. Hermann, Frankfurt am Main, 535 pp. https://doi.org/10.5962/bhl.title.87724
  19. Dal Pos, D., Carpenter, J.M. & Uliana, M. (2022) The Giordani Soika collection of Eumeninae at the Natural History Museum of Venice Giancarlo Ligabue: catalogue of species and type specimens (Insecta, Hymenoptera, Vespidae). Zootaxa, 5137 (1), 1–111. https://doi.org/10.11646/zootaxa.5137.1.1
  20. Dalla Torre, K.W. (1894) Catalogus Hymenopterorum 9. Vespidae (Diploptera). G. Engelmann, Leipzig, 181 pp.
  21. Dalla Torre, K.W. (1904) Hymenoptera, Fam. Vespidae. Genera Insectorum, 19, 1–108.
  22. Day, M.C. (1979) The species of Hymenoptera described by Linnaeus in the genera Sphex, Chrysis, Vespa, Apis and Mutilla. Biological Journal of the Linnean Society, 12 (1), 45–84. https://doi.org/10.1111/j.1095-8312.1979.tb00049.x
  23. De Geer, C. (1773) Memoires pour servir a l’histoire des Insectes. Vol. 3. Pierre Hesselberg, Stockholm, viii + 696 pp., 44 pls.
  24. de Saussure, H. (1852–1853) Études sur la famille des Vespides 1. Monographie des guêpes solitaires ou de la tribu des Euméniens. Masson, V. Paris and Kessmann, J. Genève, 286 pp., 21 pls. https://doi.org/10.5962/bhl.title.39973
  25. de Saussure, H. (1854–1856) Études sur la famille des Vespides 3. La Monographie des Masariens et un supplement à la Monographie des Euméniens. Masson, V., Paris and Cherbuliez, J., Genève, 352 pp. https://doi.org/10.5962/bhl.title.39973
  26. de Saussure, H. (1863) Mélanges hyménoptérologiques II. Vespides. Mémoires de la Société de physique et d’histoire naturelle de Genève, 17 (1), 173–244, pl. 2.
  27. de Saussure, H. (1890) Histoire Naturelle des Hyménoptères. In: Grandidier, A. (Ed.), Histoire physique, naturelle et politique de Madagascar, XX, pp. I–XXI + 1–176 + pls. 1–20. https://doi.org/10.5962/bhl.title.9201
  28. de Saussure, H. (1900) Hymenoptera. Vespidae. Abhandlungen der Senckenbergischen Naturfoschenden Gesellschaft, 26 (2), 201–240.
  29. du Buysson, R. (1898) Contribution aux Chrysidides du Globe (3. Série). Revue d’Entomologie, 17 (5), 125–147.
  30. du Buysson, R. (1900) Sur quelques Hyménoptères de Madagascar. Annales de la Société Entomologique de France, 69 (1), 177–180. https://doi.org/10.1080/21686351.1900.12279867
  31. du Buysson, R. (1907) Sur quelques parasites des Euménides. Revue d’Entomologie, 25 (8), 106–109.
  32. Emery, C., Gribodo, G. & Kriechbaumer, J. (1894) Rassegna degl’Imenotteri raccolti nel Mozambico dal Cav. Fornasini esistenti nel Museo Zoologico della R. Università di Bologna. Memorie della R. Accademia delle Scienze dell’Istituto di Bologna, 4, 45–90.
  33. Fabricius, J.C.F. (1775) Systema Entomologiae, sistens Insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. Kortii, Flensburgi et Lipsiae [Flensburg and Leipzig], [32] + 832 pp. https://doi.org/10.5962/bhl.title.36510
  34. Fabricius, J.C.F. (1793) Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Tome 2. Christ. Gottl. Proft, Hafniae, viii + 519 pp. https://doi.org/10.5962/bhl.title.122153
  35. Fabricius, J.C.F. (1798) Supplementum entomologiae systematicae. impensis Christ. Gottl. Proft, Hafniae, [2] + 572 pp. https://doi.org/10.5962/bhl.title.65803
  36. Fabricius, J.C. (1804) Systema Piezatorum. Carolum Reichard, Brunsvigae, 470 pp. [pp. i–xiv + 15–440 + 1–30]
  37. Friederichs, K. (1919) Beobachtungen über einige solitäre Wespen in Madagaskar. Mitteilungen aus dem Zoologischen Museum in Berlin, 9 (1), 27–50.
  38. Giordani Soika, A. (1934a) Un nuovo genere egiziano di Eumenini (Hymenoptera: Vespidae). Bulletin de la Société Royale Entomologique d’Egypte, 4, 436–439.
  39. Giordani Soika, A. (1934b) Labus ed Eumenes nuovi o poco noti (Hym. Vespidae). Memorie della Società Entomologica Italiana, 12, 215–228.
  40. Giordani Soika, A. (1937a) Descrizione di un nuovo Odynerus endemico del Madagascar. Bollettino della Società Veneziana di Storia Naturale, 1 (9–10), 177–179.
  41. Giordani Soika, A. (1937b) Monografia degli Odynerus etiopici, parte seconda. Annali del Museo Civico di Storia Naturale di Genova, 59, 297–362.
  42. Giordani Soika, A. (1939) Sul genere Pachymenes e descrizione di un nuovo genere e di nuove specie (Hym. Vespidae). Memorie della Società Entomologica Italiana, 17, 85–96.
  43. Giordani Soika, A. (1940) Le specie etiopiche e malagasse del genere Zethus F. (Hym. Vespidae). Memorie della Società Entomologica Italiana, 19, 129–139.
  44. Giordani Soika, A. (1941) Studi sui Vespidi solitari. Bollettino della Società Veneziana di Storia Naturale e del Museo Civico di Storia Naturale, 2 (3), 130–279.
  45. Giordani Soika, A. (1952) Missione Biologica Sagan-Omo diretta dal Prof. Edoardo Zavattari: Hymenoptera Vespidae. Rivista di Biologia Coloniale, 11, 73–89.
  46. Giordani Soika, A. (1955) Contributions à l’étude de la faune entomologique du Ruanda-Urundi (Mission P. Basilewsky 1953) XLII. Hymenoptera Vespides solitaires. Annales du Musée Royal du Congo Belge, Zoologie, 36, 362–367.
  47. Giordani Soika, A. (1957a) Expedition to SW Arabia 1937–38. 31- Hymenoptera. British Museum (Natural History) London, 1, 471–484.
  48. Giordani Soika, A. (1957b) Biogeografia, evoluzione e sistematica dei Vespidi solitari della Polinesia Meridionale. Bollettino del Museo Civico di Storia Naturale di Venezia, 10, 183–221.
  49. Giordani Soika, A. (1965) Notulae vespidologicae XXIII. Descrizione di tre nuovi Eumenidi dell’Africa occidentale. Bollettino della Società Entomologica Italiana, 95 (3–4), 46–51.
  50. Giordani Soika, A. (1973) Designazione di lectotipi ed elenco dei tipi di Eumenidi, Vespidi e Masaridi da me descritti negli anni 1934–1960. Bollettino del Museo Civico di Storia Naturale di Venezia, 24, 7–53.
  51. Giordani Soika, A. (1974a) Eumenidi di Socotra ed Abd-el-Kuri. Bollettino del Museo Civico di Storia Naturale di Venezia, 25, 69–85.
  52. Giordani Soika, A. (1974b) Revisione della sottofamiglia Raphiglossinae (Hym. Vesp.). Bollettino del Museo Civico di Storia Naturale di Venezia, 25, 107–146.
  53. Giordani Soika, A. (1977) Sui generi Leptomenes G.S., Stroudia Grib. ed Eumenidiopsis (G.S.) (Hym. Eumenidae). Bollettino del Museo Civico di Storia Naturale di Venezia, 28, 105–151.
  54. Giordani Soika, A. (1979) Revisione delle specie etiopiche e malgasce della sottofamiglia Discoeliinae (Hym.). Bollettino del Museo Civico di Storia Naturale di Venezia, 30, 19–65.
  55. Giordani Soika, A. (1989a) Terzo contributo alla conoscenza degli Eumenidi afrotropicali (Hymenoptera). Lavori della Società Veneziana di Scienze Naturali, 14 (1), 19–68.
  56. Giordani Soika, A. (1989b) Eumenidi raccolti nell’Africa occidentale, Camerun e Gabon da A. Pauly (Hymenoptera: Vespoidea, Eumenidae). Notes Fauniques de Gembloux, 20, 23–67.
  57. Giordani Soika, A. (1991a) Vespoidei raccolti da L.A. Nilsson nel Madagascar con una tabella per la determinazione delle Ropalidia del Madagascar (Hymenoptera Vespoidea). Lavori della Società Veneziana di Scienze Naturali, 16, 79–90.
  58. Giordani Soika, A. (1991b) Notulae Vespidologicae 47. Nuovi Alastor afrotropicali con descrizione di un nuovo sottogenere (Hymenoptera Vespoidea). Bollettino della Società Entomologica Italiana, 123 (1), 51–54.
  59. Gribodo, G. (1895) Seconda contribuzione alla conoscenza della fauna Imenotterologica del Mozambico. Memorie della Reale Accademia delle Scienze dell’Istituto di Bologna, Serie 5, 5, 323–366.
  60. Guérin-Méneville, F.E. (1844) Iconographie du règne animal de G. Cuvier, ou représentation d’après nature de l’une des es- pèces les plus remarquables et souvent non encore figurées, de chaque genre d’animaux. Avec un texte descriptif mis au courant de la science. Ouvrage pouvant servir d’atlas à tous les traités de zoologie. Vol. 7. Baillière, Paris, 576 pp.
  61. Gusenleitner, F. & Ockermüller, E. (2024) Dr. Josef Gusenleitner, ein bedeutender oberösterreichischer Entomologe, hat uns verlassen (*17.9.1929 †6.7.2023). Linzer biologische Beiträge, 56 (1), 379–482.
  62. Gusenleitner, J. (1970) Bemerkenswertes über Faltenwespen II (Diploptera, Hymenopt.). Nachrichtenblatt der Bayerischen Entomologen, 18 (4–6), 112–116.
  63. Gusenleitner, J. (1992) Zwei neue Eumeniden-Gattungen und -Arten aus Madagaskar (Vespoidea, Hymenoptera). Linzer biologische Beiträge, 24 (1 & 2), 91–96 & 931.
  64. Gusenleitner, J. (1995) Über Eumenidae aus Kontinentalafrika und Madagaskar (Hymenoptera, Vespoidea, Eumenidae). Linzer biologische Beiträge, 27 (1), 203–215.
  65. Gusenleitner, J. (1998) Weitere neue Arten der Familie Eumenidae aus Kontinentalafrika und Madagaskar (Hymenoptera, Eumenidae). Linzer biologische Beiträge, 30 (1), 183–194.
  66. Gusenleitner, J. (2000) Eine Aufsammlung von Eumeniden auf Madagaskar (Hymenoptera, Vespoidea, Eumenidae). Linzer biologische Beiträge, 32 (2), 923–926.
  67. Gusenleitner, J. (2004) Über Eumenidae aus Madagaskar (Hymenoptera: Eumenidae). Linzer biologische Beiträge, 36 (2), 1075–1082.
  68. Gusenleitner, J. (2005) Über bemerkenswerte Faltenwespen aus der äthiopischen Region Teil 3 (Hymenoptera, Vespoidea, Eumenidae). Linzer biologische Beiträge, 37 (2), 1177–1198.
  69. Gusenleitner, J. (2011) Über bemerkenswerte Faltenwespen aus der äthiopischen Region Teil 7 (Hymenoptera: Vespidae: Eumeninae). Linzer biologische Beiträge, 43 (1), 423–441.
  70. Gusenleitner, J. (2012) Über bemerkenswerte Faltenwespen aus der äthiopischen Region (Hymenoptera: Vespidae: Eumeninae). Teil 8. Linzer biologische Beiträge, 44 (1), 523–540.
  71. Hermes, M.G., Melo, G.A.R. & Carpenter, J.M. (2014) The higher-level phylogenetic relationships of the Eumeninae (Insecta, Hymenoptera, Vespidae), with emphasis on Eumenes sensu lato. Cladistics, 30 (5), 453–484. https://doi.org/10.1111/cla.12059
  72. Hensen, R.V. & Blommers, L.H.M. (1987) Review of the Malagasy species of Belonogaster Saussure (Hymenoptera, Vespidae). Tijdschrift voor Entomologie, 130, 11–31.
  73. Herrich-Scäffer, G.A.W. (1839) s.n. In: Fauna insectorum Germanicae initia oder Deutschlands Insecten. Heft 173. Felseckerschen Buchhandlung, Nürnberg, pls. 1–15, unnumbered text pages, pls. 16–24, pp. 1–32. https://doi.org/10.5962/bhl.title.15007
  74. ICZN (1970) Opinion 893. Eumenidae names of Saussure (Hymenoptera): Grant of availability to certain names proposed for secondary divisions of genera. Bulletin of Zoological Nomenclature, 26, 187–191.
  75. ICZN (1999) International code of zoological nomenclature. 4th Edition. International Trust for Zoological Nomenclature, London, xxix + 306 pp. https://doi.org/10.5962/bhl.title.50608
  76. Kohl, F.F. (1906) Zoologische Ergebnisse der Expedition der Kaiserlichen Akademie der Wissenschaften nach Südarabien und Sokótra im Jahre 1898-1899. Hymenopteren. Denkschriften der Kaiserlichen Akademie der Wissenschaften Wien, Mathematisch- naturwissenschaftlichen Klasse, 71, 169–301.
  77. Latreille, P.A. (1810) Considérations génèrale sur l’order naturel des animaux composant les classes des Crustacés, des Arachnides, et des Insectes: avec un tableau méthodique de leurs genres, disposes en familles. Schoell, Paris, [2] + 444 pp. https://doi.org/10.5962/bhl.title.39620
  78. Lepeletier de Saint-Fargeau, A.L.M. (1841) Historie Naturelle des Insectes. Hyménoptéres. 2. Librairie Encyclopédique de Roret, Paris, 704 pp.
  79. Linnaeus, C. (1758) Systema naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Salvius, L., Holmiae [Stockholm], iv + 823 pp. https://doi.org/10.5962/bhl.title.542
  80. Linnaeus, C. (1767) Systema naturae per regna tria naturae secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Pars II. Editio duodecima reformata. Salvius, L., Holmiae [Stockholm], [2] + 795 + [37] pp. [pp. (2) + 533–1327 + (37)] https://doi.org/10.5962/bhl.title.157601
  81. Madl, M. (1995) Vespidae von der Insel Sainte Marie (Madagaskar) (Hymenoptera). Linzer biologische Beiträge, 27 (2), 1099–1102.
  82. Magretti, P. (1884a) Raccolte imenotterologiche nell’Africa orientale. Relazione preventive. Bollettino della Società Entomologica Italiana, 50 (5), 241–253.
  83. Magretti, P. (1884b) Risultati di raccolte imenotterologiche nell’Africa orientale. Annali del Museo Civico di Storia Naturale di Genova, 21, 523–636.
  84. Meade-Waldo, G. (1913) New Species of Diploptera in the Collection of the British Museum. Part IV. Annals and Magazine of Natural History, Series 8, 11 (61), 44–54. https://doi.org/10.1080/00222931308693291
  85. Nilsson, L.A. & Rabakonandrianina, E. (1988) Chemical signalling and monopolization of nectar resources by territorial Pachymelus limbatus (Hymenoptera, Anthophoridae) male bees in Madagascar. Journal of Zoology, London, 215, 475–489. https://doi.org/10.1111/j.1469-7998.1988.tb02854.x
  86. Pérez, J. (1907) Hyménoptéres [mission] J. Bonnier et Ch. Perez, golfe Persique, 1901. Bulletin Scientifique de la France et de la Belgique, 41, 485–505. https://doi.org/10.5962/bhl.part.21339
  87. Polašek, O. & de Beer, L. (2023) A new species of social wasp from Madagascar with an inverted nest architecture (Hymenoptera, Vespidae). Journal of Hymenoptera Research, 96, 1031–1044.
  88. https://doi.org/10.3897/jhr.96.103379
  89. Retzius, S.I. (1783) Genera et Species Insectorum Generosissimis Auctoris Scriptis. Extraxit, digessit, latine quoad partem reddidit, et terminologiam insectorum Linneam addidit. Apud Sigfried Lebrecht Crusium, Lipsiae, XXXII + 220 pp.
  90. Salt, G. (1927) Notes on the Strepsiptera and their hymenopterous hosts. Psyche, 34 (5), 182–192.
  91. https://doi.org/10.1155/1927/25470
  92. Salt, G. & Bequaert, J. (1929) Stylopized Vespidae. Psyche, 36 (3), 249–282.
  93. https://doi.org/10.1155/1929/78563
  94. Saunders, S.S. (1850) Descriptions of some new aculeate Hymenoptera from Epirus. Transactions of the Entomological Society of London, New Series, 1, 69–75, pls. 5–6.
  95. https://doi.org/10.1111/j.1365-2311.1850.tb02485.x
  96. Schulz, W.A. (1912) Aelteste und alte Hymenopteren skandinavischer Autoren. Berliner Entomologische Zeitschrift, 57 (1–2), 52–102.
  97. https://doi.org/10.1002/mmnd.19120570109
  98. Selis, M. (2023) Taxonomy of the Afrotropical genus Leptomenes Giordani Soika, with description of a new species from South Africa (Hymenoptera: Vespidae: Eumeninae). Zootaxa, 5228 (5), 584–594.
  99. https://doi.org/10.11646/zootaxa.5228.5.3
  100. Selis, M. (2024) Revision of the genus Cyrtolabulus van der Vecht in Madagascar (Hymenoptera: Vespidae: Eumeninae). Zootaxa, 5406 (1), 66–86.
  101. https://doi.org/10.11646/zootaxa.5406.1.3
  102. Selis, M. & Carpenter, J.M. (2023) Revision of the Afrotropical species of the genus Anterhynchium de Saussure (Hymenoptera: Vespidae: Eumeninae). Zootaxa, 5233 (1), 1–102.
  103. https://doi.org/10.11646/zootaxa.5233.1.1
  104. Selis, M. & Carpenter, J.M. (2024) Revision of the Afrotropical genus Afrepipona Giordani Soika and description of Afrepsilon gen. nov. (Hymenoptera: Vespidae: Eumeninae). European Journal of Taxonomy, 944, 1–80.
  105. https://doi.org/10.5852/ejt.2024.944.2607
  106. Shorthouse, D.P. (2010) SimpleMappr, an online tool to produce publication-quality point maps. Available from: https://simplemappr.net (accessed 6 June 2025)
  107. Smith, F. (1857) Catalogue of the Hymenopterous Insects in the Collection of the British Museum. Part 5. Vespidae. Printed by order of the Trustees, London, 147 pp.
  108. Spinola, M. (1841) Hyménoptères recuellis a Cayenne en 1839 par M. Leprieur, pharmacien de la Marine Royale. Seconde Partie. Porte-Aiguillons. Annales de la Societe Entomologique de France, 10, 85–149.
  109. van der Vecht, J. (1959) On some Fabrician types of Indo-Australian Vespidae (Hymenoptera). Archives Neerlandaises Zoologi, 13 (Supplement 1), 234–247, 4 figs.
  110. van der Vecht, J. (1963) Studies on Indo-Australian and East Asiatic Eumenidae (Hymenoptera: Vespoidea). Zoologische Verhandelingen Leiden, 60, 1–116.
  111. van der Vecht, J. (1967) The status of certain genus-group names in the Eumenidae (Hymenoptera; Vespoidea). Bulletin of Zoological Nomenclature, 24, 27–33.
  112. https://doi.org/10.5962/bhl.part.15358
  113. van der Vecht, J. (1969) A new name for the genus Cyrtolabus van der Vecht (Hymenoptera, Eumenidae). Entomologische Berichten, Amsterdam, 29, 1–2.
  114. van der Vecht, J. (1970) Further notes on Fabrician types of Vespoidea (Hymenoptera). Proceedings of the Koninklijke nederlandse Akademie Wetenschappen Amsterdam, Series C, 73 (1), 15–24.
  115. van der Vecht, J. & Carpenter, J.M. (1990) A catalogue of the genera of the Vespidae (Hymenoptera). Zoologische Verhandelingen, 260, 1–62.
  116. Villers, C. de (1789) Caroli Linnaei Entomologia, faunae Suecicae descriptionibus aucta: DD. Scopoli, Geoffroy, de Geer, Fabricii, Schrank, &c., Speciebus vel in Systemate non Enumeratis, vel Nuperrime detectis, vel Speciebus Galliae Australis locupletata, Generum Specierumque rariorum iconibus ornata. Vol. 3. Sumptibus Piestre et Delamolliere, Lugduni, 657 pp., pls. VII–X.
  117. von Schulthess, A. (1907) Vespiden von Madagaskar, den Comoren und Ostafrika. In: Voeltzkow, A. (Ed.), Reise in Ostafrika in den Jahren 1903–1905 mit Mitteln der Hermann und Elise geb. Heckmann Wentzel-Stiftung ausgeführt von Alfred Voeltzkow. Wissenschaftliche Ergebnisse. 2. Band. E. Scweizerbartsche Verlagsbuchhandlung, Stuttgart, pp. 63–73.
  118. von Schulthess, A. (1913) Vespiden aus dem Stockholmer Museum II. Kritik der Cameron’schen Vespidenbestimmungen von Professor Sjöstedt’s Kilimandjaro-Expedition. Arkiv för Zoologi, 8 (2–3: 17), 1–24.
  119. von Schulthess, A. (1919) Verzeichnis von Dr. K. Friederichs in Madagaskar erbeuteter Hymenopteren. In: Friederichs, K., Beobachtungen über einige solitäre Wespen in Madagaskar. Mitteilungen aus dem Zoologischen Museum in Berlin, 9 (1), 46–50.
  120. Voss, E. (1925) Die Unterfamilien Attelabinae und Apoderinae. (Col. Curc.) (18. Beitrag zur Kenntnis der Curculioniden). Stettiner Entomologische Zeitung, 85 (1–2), 1–78 + 191–304, pls. 1–3.
  121. Zavattari, E. (1909) Imenotteri del Kashmir. Bollettino dei Musei di Zoologia ed Anatomia comparata della Regia Università di Torino, 24 (605), 1–6.
  122. Zirngiebl, L. (1953) Zur Wespen-Fauna der Pfalz. Mitteilungen Pollichia, Series 3, 1, 160–179.

How to Cite

Selis, M. (2025) The solitary vespid wasps of Madagascar (Hymenoptera: Vespidae: Eumeninae, Raphiglossinae and Zethinae). Zootaxa, 5705 (1), 1–171. https://doi.org/10.11646/zootaxa.5705.1.1