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Issue: Vol. 5722 No. 2: 19 Nov. 2025
Type: Article
Published: 2025-11-19
DOI: 10.11646/zootaxa.5722.2.3
Page range: 199-228
Abstract views: 55
PDF downloaded: 2

Overview of Campopleginae (Hymenoptera: Ichneumonidae) species described from 2016 to 2025 with revised taxon metrics

Universidade Federal do Tocantins (UFT); Porto Nacional; Tocantins; Brazil
Hymenoptera nomenclature parasitoid wasps new species dataset

Abstract

The subfamily Campopleginae, one of the largest groups within the family Ichneumonidae, faces persistent challenges in maintaining updated taxonomic records due to the rapid pace of new species descriptions. This study compiles and consolidates recent taxonomic information, presenting a comprehensive list of species described between 2016 and 2025, along with a dataset documenting nomenclatural changes. In total, two new genera and 242 new species of Darwin wasps were identified from 67 articles published during this period. By unifying scattered data, this work addresses a critical gap in the literature and provides a reference tool that reflects the dynamic growth of Campopleginae taxonomy.

 

References

  1. Araujo, R.O. & Giovanni, F.D. (2021) Description of the first species of Nemeritis Holmgren (Hymenoptera: Ichneumonidae: Campopleginae) from the Southern Hemisphere, with a key to the New World species. Zootaxa, 5023 (2), 263–272. https://doi.org/10.11646/zootaxa.5023.2.6
  2. Boparai, J. & Broad, G.R. (2024) The genome sequence of an ichneumonid wasp, Rhimphoctona (Xylophylax) megacephalus (Gravenhorst, 1829). Wellcome Open Research, 9, 472. https://doi.org/10.12688/wellcomeopenres.22865.1
  3. Broad, G.R. (2024) The genome sequence of an ichneumonid wasp, Hyposoter dolosus (Gravenhorst, 1829). Wellcome Open Research, 9, 499. https://doi.org/10.12688/wellcomeopenres.22895.1
  4. Broad, G.R., Shaw, M.R. & Fitton, M.G. (2018) Ichneumonid wasps: (Hymenoptera: Ichneumonidae): their classification and biology. Royal Entomological Society, London, 418 pp. https://doi.org/10.1079/9781800625471.0000
  5. Carlson, R.W. (1979) Family Ichneumonidae. In: Catalog of hymenoptera in America north of Mexico. Vol.1 Smithsonian Institution Press, Washington, pp. 315–741.
  6. Choi, J.-K. (2022) Description of a new species of the genus Venturia Schrottky (Hymenoptera: Ichneumonidae: Campopleginae) from South Korea. Journal of Species Research, 11 (2), 128–131. https://doi.org/10.12651/JSR.2022.11.2.128
  7. Choi, J.-K. (2024) New Host Plants of Charops striatus (Uchida) (Hymenoptera: Ichneumonidae: Campopleginae) from South Korea. Proceedings of the National Institute of Ecology of the Republic of Korea, 5 (2), 68–70. https://doi.org/10.22920/PNIE.2024.5.2.68
  8. Cisneros, J.C., Raja, N.B., Ghilardi, A.M., Dunne, E.M., Pinheiro, F.L., Regalado Fernández, O.R., Sales, M.A.F., Rodríguez-de La Rosa, R.A., Miranda-Martínez, A.Y., González-Mora, S., Bantim, R.A.M., De Lima, F.J. & Pardo, J.D. (2022) Digging deeper into colonial palaeontological practices in modern day Mexico and Brazil. Royal Society Open Science, 9 (3), 210898. https://doi.org/10.1098/rsos.210898
  9. Cox, M.L. & Broad, G.R. (2020) Nepiesta mandibularis (Holmgren, 1860) (Hymenoptera: Ichneumonidae, Campopleginae), reared from the larvae of Chrysolina oricalcia (Müller, 1776) (Coleoptera: Chrysomelidae, Chrysomelinae) in Essex, U.K. Entomologist’s Gazette, 71 (2), 98–102. https://doi.org/10.31184/G00138894.712.1760
  10. Dalla Torre, C.G. de (1901) Catalogus Hymenopterorum. Volumen III. Trigonalidae, Megalyridae, Stephanidae, Ichneumonidae, Agriotypidae, Evaniidae, Pelecinidae. Guilelmi Engelmann, Lipsiae, 544 pp.
  11. Dalla Torre, C.G. de (1902) Catalogus Hymenopterorum. Volumen III. Trigonalidae, Megalyridae, Stephanidae, Ichneumonidae, Agriotypidae, Evaniidae, Pelecinidae. Guilelmi Engelmann, Lipsiae, 597 pp. [pp.545–1141]
  12. Di Giovanni, F. & Riedel, M. (2017) New records of Campopleginae for Italy (Hymenoptera: Ichneumonidae). Fragmenta entomologica, 49 (1), 109–114. https://doi.org/10.4081/fe.2017.239
  13. Dunnington, D. (2025) ggspatial: Spatial Data Framework for ggplot2. Available from: https://github.com/paleolimbot/ggspatial (accessed 13 October 2025)
  14. Engel, M.S., Ceríaco, L.M., Daniel, G.M., Dellapé, P.M., Löbl, I., Marinov, M., Reis, R.E., Young, M.T., Dubois, A. & Agarwal, I. (2021) The taxonomic impediment: a shortage of taxonomists, not the lack of technical approaches. Zoological Journal of the Linnean Society, 193 (2), 381–387. https://doi.org/10.1093/zoolinnean/zlab072
  15. Feizi, A., Talebi, A.A. & Zardouei Heydari, M. (2024) Contribution to the genus Dusona Cameron, 1901 (Hymenoptera, Ichneumonidae, Campopleginae) from Iran, with three new records and an updated checklist. Journal of Crop Protection, 13 (1), 1–13.
  16. Galsworthy, A., Shaw, M.R. & Haraldseide, H. (2023) A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species. Zootaxa, 5290 (1), 1–73. https://doi.org/10.11646/zootaxa.5290.1.1
  17. García-Roselló, E., González-Dacosta, J. & Lobo, J.M. (2023) The biased distribution of existing information on biodiversity hinders its use in conservation, and we need an integrative approach to act urgently. Biological Conservation, 283, 110118. https://doi.org/10.1016/j.biocon.2023.110118
  18. Gil, O.J.A., Souza, L.A.D., Funichello, M. & Busoli, A.C. (2020) The sex ratio of the koinobiont parasitoid Microcharops anticarsiae Gupta remains female-biased on young larvae of velvetbean in the laboratory environment. Acta Scientiarum. Biological Sciences, 42, e48187. https://doi.org/10.4025/actascibiolsci.v42i1.48187
  19. Gupta, V.K. (1987) The Ichneumonidae of the Indo-Australian area (Hymenoptera). Memoirs of the American Entomological Institute, 41 (1), 1–597.
  20. Han, Y.-Y., van Achterberg, K. & Chen, X.-X. (2021a) New species and records of Venturia Schrottky (Hymenoptera, Ichneumonidae, Campopleginae) from China and Nepal. ZooKeys, 1041, 113. https://doi.org/10.3897/zookeys.1041.64238
  21. Han, Y.-Y., van Achterberg, K.V. & Chen, X.-X. (2021b) The genus Campoplex Gravenhorst, 1829 (Hymenoptera, Ichneumonidae, Campopleginae) from China. Zootaxa, 5066 (1), 1–121. https://doi.org/10.11646/zootaxa.5066.1.1
  22. Han, Y.-Y., van Achterberg, K. & Chen, X.-X. (2021c) Five new species of the genus Sinophorus Förster (Hymenoptera, Ichneumonidae, Campopleginae) from China. Zootaxa, 5061 (1), 115–133. https://doi.org/10.11646/zootaxa.5061.1.5
  23. Han, Y.-Y., van Achterberg, K. & Chen, X.-X. (2021d) The genus Casinaria Holmgren, 1859 (Hymenoptera: Ichneumonidae, Campopleginae) from China. Zootaxa, 4974 (3), 504–536. https://doi.org/10.11646/zootaxa.4974.3.3
  24. Han, Y.-Y., van Achterberg, K. & Chen, X.-X. (2022a) Four rare genera of the subfamily Campopleginae (Hymenoptera: Ichneumonidae) from China, with description of three new species. Zootaxa, 5219 (3), 247–264. https://doi.org/10.11646/zootaxa.5219.3.3
  25. Han, Y.-Y., van Achterberg, K. & Chen, X.-X. (2022b) The rare genera Picacharops Gauld and Sliochia Gauld (Hymenoptera, Ichneumonidae, Campopleginae) found in China, with description of two new species. Zootaxa, 5133 (4), 577–584. https://doi.org/10.11646/zootaxa.5133.4.8
  26. Han, Y.-Y., van Achterberg, K. & Chen, X.-X. (2022c) Review of the genera Breviterebra Kusigemati, Charops Holmgren and Scenocharops Uchida (Hymenoptera, Ichneumonidae, Campopleginae) from China, with description of three new species. Zootaxa, 5133 (4), 527–542. https://doi.org/10.11646/zootaxa.5133.4.4
  27. Han, Y.-Y., van Achterberg, K.V. & Chen, X.-X. (2024a) DNA barcodes and morphology reveal two new species of the genus Prochas Walkley, 1959 (Ichneumonidae, Campopleginae), from China. Insects, 15 (12), 968. https://doi.org/10.3390/insects15120968
  28. Han, Y.-Y., Wei, C., Liu, C. & Dong, Y. (2024b) New Species and Records of Lemophagus Townes, 1965 (Hymenoptera, Ichneumonidae, Campopleginae), from China. Insects, 15 (12), 932. https://doi.org/10.3390/insects15120932
  29. Han, Y.-Y., Wei, C.-X., Chen, M.-M. & Zhu, L.-X. (2025a) A new species of genus Venturia Schrottky, 1902 (Hymenoptera: Ichneumonidae: Campopleginae) from China. Annales Zoologici, 75 (2), 527–533. https://doi.org/10.3161/00034541ANZ2025.75.2.008
  30. Han, Y.-Y., Wei, C.-X., Xu, R.-L. & Chen, X.-X. (2025b) First record of the genus Nemeritis Holmgren (Ichneumonidae, Campopleginae) from China, with descriptions of two new species. Zootaxa, 5679 (2), 286–292. https://doi.org/10.11646/zootaxa.5679.2.8
  31. Haraldseide, H. (2021) Two new species of Meloboris Holmgren, 1859 (Campopleginae, Ichneumonidae, Hymenoptera) from Norway. Norwegian Journal of Entomology, 68, 193–202.
  32. Haraldseide, H. (2023) Two new species of campoplegine ichneumonid wasps (Hymenoptera, Ichneumonidae, Campopleginae) from Germany. Norwegian Journal of Entomology, 70, 132–137.
  33. Heberling, J.M., Miller, J.T., Noesgaard, D., Weingart, S.B. & Schigel, D. (2021) Data integration enables global biodiversity synthesis. Proceedings of the National Academy of Sciences, 118 (6), e2018093118. https://doi.org/10.1073/pnas.2018093118
  34. Herrera-Florez, A.F., Geraldo, M., Filho, M.M.D. & Penteado-Dias, A. (2017) A new species of Jomine Graf-Kumagai (Hymenoptera: Ichneumonidae: Campopleginae) from Brazil. Zootaxa, 4226 (2), 292–300. https://doi.org/10.11646/zootaxa.4226.2.9
  35. INCT (2025) Instituto Nacional de Ciência e Tecnologia dos Hymenoptera Parasitóides (INCT Hympar). Available from: https://hympar.com.br/ (accessed 18 September 2025)
  36. Karimi, S., Ghassemi-Kahrizeh, A., Hosseinzadeh, A., Lotfalizadeh, H. & Riedel, M. (2023) A new genus and two new species with some new records of the subfamily Campopleginae (Hymenoptera: Ichneumonidae) from Iran. Egyptian Journal of Biological Pest Control, 33 (1), 117. https://doi.org/10.1186/s41938-023-00761-5
  37. Klopfstein, S., Broad, G., Urfer, K., Vårdal, H. & Haraldseide, H. (2022) An interactive key to the European genera of Campopleginae (Hymenoptera, Ichneumonidae) and 20 new species for Sweden. Entomologisk tidskrift, 143, 121–156.
  38. Klopfstein, S., Riedel, M. & Schwarz, M. (2019a) Checklist of ichneumonid parasitoid wasps in Switzerland (Hymenoptera, Ichneumonidae): 470 species new for the country and an appraisal of the alpine diversity. Alpine Entomology, 3, 51–81. https://doi.org/10.3897/alpento.3.31613
  39. Klopfstein, S., Santos, B.F., Shaw, M.R., Alvarado, M., Bennett, A.M.R., Dal Pos, D., Giannotta, M., Herrera Florez, A.F., Karlsson, D., Khalaim, A.I., Lima, A.R., Mikó, I., Sääksjärvi, I.E., Shimizu, S., Spasojevic, T., Van Noort, S., Vilhelmsen, L. & Broad, G.R. (2019b) Darwin wasps: a new name heralds renewed efforts to unravel the evolutionary history of Ichneumonidae. Entomological Communications, 1, ec01006. https://doi.org/10.37486/2675-1305.ec01006
  40. Li, T., Chang, G.-B., Yang, Z.-H., Sun, S.-P., Tian, Y. & Sheng, M.-L. (2021) Two new species of Cymodusa Holmgren (Hymenoptera, Ichneumonidae) with a key to species known from China and Oriental region. Journal of Hymenoptera Research, 88, 103–114. https://doi.org/10.3897/jhr.88.75304
  41. Meier, N., Urfer, K., Haraldseide, H., Vårdal, H. & Klopfstein, S. (2022) Open access in a taxonomic sense: a morphological and molecular guide to Western Palaearctic Dusona (Hymenoptera, Ichneumonidae). Journal of Hymenoptera Research, 91, 83–183. https://doi.org/10.3897/jhr.91.83318
  42. Miralles, A., Bruy, T., Wolcott, K., Scherz, M.D., Begerow, D., Beszteri, B., Bonkowski, M., Felden, J., Gemeinholzer, B., Glaw, F., Glöckner, F.O., Hawlitschek, O., Kostadinov, I., Nattkemper, T.W., Printzen, C., Renz, J., Rybalka, N., Stadler, M., Weibulat, T., Wilke, T., Renner, S.S. & Vences, M. (2020) Repositories for Taxonomic Data: Where We Are and What is Missing Friedman, M. (Ed.), Systematic Biology, 69 (6), 1231–1253. https://doi.org/10.1093/sysbio/syaa026
  43. Onody, H.C. & Penteado-Dias, A.M. (2016) A new species of Cryptophion (Hymenoptera: Ichneumonidae) from southeastern Brazil. Zoologia (Curitiba), 33 (1), e20150101. https://doi.org/10.1590/S1984-4689zool-20150101
  44. Onody, H.C. & Penteado-Dias, A.M. (2019) Description of the first species of Cymodusa Holmgren from Brazil (Hymenoptera, Ichneumonidae, Campopleginae). Zootaxa, 4545 (3), 434–440. https://doi.org/10.11646/zootaxa.4545.3.7
  45. Ranjith, A.P. & Kedar, S.C. (2025) New species of Lemophagus Townes, 1965 (Hymenoptera, Ichneumonidae, Campopleginae) from India reared as a larval parasitoid of Crioceris nigroornata Clarke, 1866 (Coleoptera, Chrysomelidae). Journal of Hymenoptera Research, 98, 743–755. https://doi.org/10.3897/jhr.98.154760
  46. Riedel, M. (2017) Die westpaläarktischen Arten der Gattung Campoletis Förster (Hymenoptera, Ichneumonidae, Campopleginae). Spixiana, 40 (1), 95–137.
  47. Riedel, M. (2018) Revision of the Western Palaearctic species of the genus Casinaria Holmgren (Hymenoptera, Ichneumonidae, Campopleginae). Linzer Biologische Beiträge, 50 (1), 687–716.
  48. Riedel, M. (2022) New records of Darwin wasps (Hymenoptera, Ichneumonidae) from Norway, with descriptions of three new species. Norwegian Journal of Entomology.
  49. Riedel, M., Ameri, A., Talebi, A.A. & Ebrahimi, E. (2019a) Contribution to the Ichneumonidae (Hymenoptera) of Iran, with descriptions of seven new species. Linzer Biologische Beiträge, 51 (1), 361–390.
  50. Riedel, M., Diller, E. & Japoshvili, G. (2018) The Ichneumonid fauna (Hymenoptera: Ichneumonidae) of Lagodekhi Reserve, Sakartvelo (Georgia), with descriptions of four new species. Linzer Biologische Beiträge, 50 (2), 1447–1507.
  51. Riedel, M., Mohammadi-Khoramabadi, A. & Khayrandish, M. (2019b) Two new species of Campopleginae (Hymenoptera: Ichneumonidae) from Iran. Zoology in the Middle East, 65 (3), 256–260. https://doi.org/10.1080/09397140.2019.1615751
  52. Riedel, M., Pénigot, W., Schwarz, M., Diller, E., Johannsson, N. & Japoshvili, G. (2023) Darwin wasps (Hymenoptera, Ichneumonidae) of the Kintrishi National Park, Sakartvelo (Georgia), with descriptions of six new species. Linzer biologische Beiträge, 55 (1), 61–153.
  53. Santos, A.D., Brandão, C.R.F. & Onody, H.C. (2025) Host associations of Brazilian Darwin wasps: current knowledge and the IchHostBR dataset. The Science of Nature, 112 (2), 24. https://doi.org/10.1007/s00114-025-01974-4
  54. Santos, A.D., Onody, H.C. & Brandão, C.R.F. (2019) Taxonomic contributions to the genus Charops Holmgren, 1859 (Hymenoptera: Ichneumonidae), with description of seven new species from Brazil. Zootaxa, 4619 (1), 45–76. https://doi.org/10.11646/zootaxa.4619.1.2
  55. Santos, A.D., Onody, H.C. & Palacio, E. (2022) First record of Cryptophion Viereck, 1913 (Ichneumonidae: Campopleginae) from Colombia, with description of a new species. Zootaxa, 5209 (4), 483–489. https://doi.org/10.11646/zootaxa.5209.4.7
  56. Shaw, M.R., Horstmann, K. & Whiffin, A.L. (2016) Two hundred and twenty-five species of reared western Palaearctic Campopleginae (Hymenoptera: Ichneumonidae) in the National Museums of Scotland, with descriptions of new species of Campoplex and Diadegma, and records of fifty-five species new to Britain. Entomologist’s Gazette, 67 (3), 177–222.
  57. Townes, H. (1944) A catalogue and reclassification of the Nearctic Ichneumonidae (Hymenoptera). Memoirs of the American Entomological Society, 11, 1–477.
  58. Townes, H.K. (1945) A Catalogue and Reclassification of the Nearctic Ichneumonidae (Hymenoptera). Part II. The Subfamilies Mesoleiinae, Plectiscinae, Orthocentrinae, Diplazontinae, Metopiinae, Ophioninae, Mesochorinae. Memoirs of the American Entomological Society, 11, 478–925.
  59. Townes, H.K. & Townes, M. (1966) A catalogue and reclassification of the Neotropic Ichneumonidae. Memoirs of the American Entomological Institute, 8, 1–367.
  60. Vas, Z. (2019a) Contributions to the taxonomy, identification, and biogeography of Casinaria Holmgren and Venturia Schrottky (Hymenoptera: Ichneumonidae: Campopleginae). Zootaxa, 4664 (3), 351–364. https://doi.org/10.11646/zootaxa.4664.3.3
  61. Vas, Z. (2019b) Contributions to the taxonomy, identification, and biogeography of Eriborus Förster, 1869 and Nepiesta Förster, 1869 (Hymenoptera: Ichneumonidae: Campopleginae). Opuscula Zoologica, Budapest, 50 (1), 87–98. https://doi.org/ 10.18348/opzool.2019.1.87
  62. Vas, Z. (2019c) Contributions to the taxonomy, identification, and biogeography of the Western Palaearctic species of Campoletis Förster (Ichneumonidae: Campopleginae). Zootaxa, 4565 (3), 373–382. https://doi.org/10.11646/zootaxa.4565.3.4
  63. Vas, Z. (2019d) New species and new records of Campopleginae from the Palaearctic region (Hymenoptera: Ichneumonidae). Folia entomologica hungarica, 80, 247–271. https://doi.org/10.17112/FoliaEntHung.2019.80.247
  64. Vas, Z. (2019e) New species and new records of ichneumon wasps from the Eastern Mediterranean and the Black Sea regions (Hymenoptera: Ichneumonidae). Acta Zoologica Academiae Scientiarum Hungaricae, 65 (1), 19–30. https://doi.org/10.17109/AZH.65.1.19.2019
  65. Vas, Z. (2020a) Contributions to the taxonomy and biogeography of Nemeritis Holmgren (Hymenoptera: Ichneumonidae: Campopleginae). Zootaxa, 4758 (3), 486–500. https://doi.org/10.11646/zootaxa.4758.3.4
  66. Vas, Z. (2020b) New species and records of Afrotropical, Australasian, Oriental and Palaearctic Casinaria Holmgren, 1859 (Hymenoptera: Ichneumonidae: Campopleginae). Opuscula Zoologica, 51 (2), 133–152. https://doi.org/10.18348/opzool.2020.2.133
  67. Vas, Z. (2020c) New species and records of Afrotropical, Oriental and Palaearctic Venturia Schrottky, 1902 (Hymenoptera: Ichneumonidae: Campopleginae). Opuscula Zoologica, 51 (2), 97–114. https://doi.org/ 10.18348/opzool.2020.2.97
  68. Vas, Z. (2020d) New species and records of Charops Holmgren, 1859 (Hymenoptera: Ichneumonidae: Campopleginae). Acta Zoologica Academiae Scientiarum Hungaricae, 66 (3), 247–267. https://doi.org/10.17109/AZH.66.3.247.2020
  69. Vas, Z. (2020e) New species and records of Palaearctic Campopleginae, Cryptinae, Hybrizontinae, and Tersilochinae (Hymenoptera: Ichneumonidae). Opuscula Zoologica, 51 (1), 3–13. https://doi.org/10.18348/opzool.2020.1.3
  70. Vas, Z. (2021a) First record of Cymodusa Holmgren from the Afrotropical region with the description of a new species (Hymenoptera: Ichneumonidae: Campopleginae). Zootaxa, 5067 (3), 447–450. https://doi.org/10.11646/zootaxa.5067.3.8
  71. Vas, Z. (2021b) New Species and Records of Afrotropical Campoletis Förster, 1869 (Hymenoptera: Ichneumonidae: Campopleginae). Annales Musei historico-naturalis hungarici, 113, 39–49. https://doi.org/10.53019/AnnlsMusHistNatHung.2021.113.39
  72. Vas, Z. (2021c) New species and records of Afrotropical Campopleginae III. (Hymenoptera: Ichneumonidae). Folia entomologica hungarica, 82, 23–42. https://doi.org/10.17112/FoliaEntHung.2021.82.23
  73. Vas, Z. (2022a) Contributions to the taxonomy and biogeography of Afrotropical Eriborus Förster, 1869 (Hymenoptera: Ichneumonidae: Campopleginae). Acta Zoologica Academiae Scientiarum Hungaricae, 68 (2), 169–178. https://doi.org/10.17109/AZH.68.2.169.2022
  74. Vas, Z. (2022b) Contributions to the taxonomy, identification, and biogeography of the Palaearctic species of Cymodusa Holmgren (Hymenoptera: Ichneumonidae: Campopleginae). Zootaxa, 5162 (3), 268–276. https://doi.org/10.11646/zootaxa.5162.3.4
  75. Vas, Z. (2022c) New species and new records of ichneumon wasps from Africa (Hymenoptera: Ichneumonidae: Campopleginae, Cryptinae, Ophioninae). Folia entomologica hungarica, 83, 25–39. https://doi.org/10.17112/FoliaEntHung.2022.83.25
  76. Vas, Z. (2022d) New species and records of Afrotropical Campopleginae IV. (Hymenoptera: Ichneumonidae). Folia Entomologica Hungarica, 83, 63–90. https://doi.org/10.17112/FoliaEntHung.2022.83.63
  77. Vas, Z. (2023a) ‘Revisiting’ North Korea: new species and new records of Campopleginae (Hymenoptera: Ichneumonidae). Annales Musei Historico-Naturalis Hungarici, 115, 215–235. https://doi.org/10.53019/AnnlsMusHistNatHung.2023.115.215
  78. Vas, Z. (2023b) Contribution to the taxonomy, identification, and biogeography of the Palaearctic species of Bathyplectes Förster and Leptoperilissus Schmiedeknecht (Hymenoptera: Ichneumonidae: Campopleginae). Zootaxa, 5270 (1), 105–114. https://doi.org/10.11646/zootaxa.5270.1.5
  79. Vas, Z. (2023c) New species and new records of ichneumon wasps, with descriptions of two new species of Melalophacharops Uchida, 1928 (Hymenoptera: Ichneumonidae). Folia Entomologica Hungarica, 84, 47–61. https://doi.org/10.17112/FoliaEntHung.2023.84.47
  80. Vas, Z. (2023d) Still from Bíró’s cornucopia: new species and new records of Campopleginae from Australia (Hymenoptera: Ichneumonidae). Folia Entomologica Hungarica, 84, 63–97. https://doi.org/10.17112/FoliaEntHung.2023.84.63
  81. Vas, Z. (2024a) New species and new records of Oriental Campopleginae and Nesomesochorinae (Hymenoptera: Ichneumonidae). Folia entomologica hungarica, 85, 61–79. https://doi.org/10.17112/FoliaEntHung.2024.85.61
  82. Vas, Z. (2024b) New species and records of Afrotropical Campopleginae V. (Hymenoptera: Ichneumonidae). Folia Entomologica Hungarica, 85, 15–25. https://doi.org/10.17112/FoliaEntHung.2024.85.15
  83. Vas, Z. (2024c) New species and records of Palaearctic, Afrotropical and Neotropical ichneumon wasps (Hymenoptera: Ichneumonidae). Annales Musei Historico-Naturalis Hungarici, 116, 239–256. https://doi.org/10.53019/AnnlsMusHistNatHung.2024.116.239
  84. Vas, Z. (2024d) New species of Campoletis Förster, 1869 from the Neotropical region (Hymenoptera: Ichneumonidae: Campopleginae). Animal Taxonomy and Ecology, 70 (1), 22–29. https://doi.org/10.1556/1777.2024.12681
  85. Vas, Z. (2024e) The zoological results of Gy. Topál’s collectings in South Argentina. 27. Ichneumonidae: Campopleginae: Diadegma Förster, 1869 (Hymenoptera). Annales Musei historico-naturalis hungarici, 116, 257–274. https://doi.org/10.53019/AnnlsMusHistNatHung.2024.116.257
  86. Vas, Z. (2025) The zoological results of Gy. Topál’s collectings in South Argentina. 28. Ichneumonidae: Campopleginae: Hyposoter Förster, 1869 and Campoletis Förster, 1869 (Hymenoptera). Annales Musei historico-naturalis hungarici, 117, 47–64. https://doi.org/10.53019/AnnlsMusHistNatHung.2025.117.47
  87. Vas, Z. & Di Giovanni, F. (2020) New species and records of Afrotropical Campopleginae (Hymenoptera: Ichneumonidae). Folia Entomologica Hungarica, 81, 105–114. https://doi.org/10.17112/FoliaEntHung.2020.81.105
  88. Vas, Z. & Di Giovanni, F. (2021) New species and records of Afrotropical Campopleginae II.(Hymenoptera: Ichneumonidae). Folia Entomologica Hungarica, 82, 7–22. https://doi.org/10.17112/FoliaEntHung.2021.82.7
  89. Vas, Z., Rezaei, S., Fallahzadeh, M., Mohammadi-Khoramabadi, A., Saghaei, N. & Ljubomirov, T. (2022) Contributions to the taxonomy, identification, and biogeography of Palaearctic Campopleginae (Hymenoptera: Ichneumonidae), with the descriptions of four new species from Iran. Zootaxa, 5134 (2), 261–274. https://doi.org/10.11646/zootaxa.5134.2.5
  90. Vas, Z. & Toshko, L. (2022) New species and new records of Campopleginae from Bulgaria (Hymenoptera: Ichneumonidae). Annales Musei Historico Naturalis Hungarici, 114, 1–8. https://doi.org/10.53019/AnnlsMusHistNatHung.2022.114.1
  91. Watanabe, K. (2019) Revision of the Genus Pyracmon (Insecta: Hymenoptera: Ichneumonidae) from Japan, with Description of a New Species. Species Diversity, 24 (2), 281–285. https://doi.org/10.12782/specdiv.24.281
  92. Watanabe, K. (2020) Revision of the genus Cymodusa Holmgren, 1859 (Hymenoptera: Ichneumonidae: Campopleginae) from Japan. Japanese Journal of Systematic Entomology, 26 (2), 201–205.
  93. Wei, Y.-W., Zhou, Y.-B., Zou, Q.-C. & Sheng, M.-L. (2020) A new species of Campoletis Förster (Hymenoptera, Ichneumonidae) with a key to species known from China, Japan and South Korea. ZooKeys, 1004, 99–108. https://doi.org/10.3897/zookeys.1004.57913
  94. Wickham, H. (2016) Data Analysis. In: Wickham, H. (Ed.), ggplot2: Elegant Graphics for Data Analysis. Springer International Publishing, Cham, pp. 189–201. https://doi.org/10.1007/978-3-319-24277-4_9
  95. Winston, J.E. (1999) Describing species: practical taxonomic procedure for biologists. Columbia University Press.
  96. Yu, D.S.K. & Horstmann, K. (1997) A catalogue of world Ichneumonidae (Hymenoptera). Memoirs of the American Entomological Institute, 58 (1), 1–1558.
  97. Yu, D.S.K., van Achterberg, C. & Horstmann, K. (2016) Taxapad 2016, Ichneumonoidea 2015. www.taxapad.com, Nepean, Ontario. [database on flash-drive]

How to Cite

Santos, A.D.D. (2025) Overview of Campopleginae (Hymenoptera: Ichneumonidae) species described from 2016 to 2025 with revised taxon metrics. Zootaxa, 5722 (2), 199–228. https://doi.org/10.11646/zootaxa.5722.2.3