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Issue: Vol. 5732 No. 1: 18 Dec. 2025
Type: Monograph
Published: 2025-12-18
DOI: 10.11646/zootaxa.5732.1.1
Page range: 1-70
Abstract views: 121
PDF downloaded: 7

Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra

Institute of Biodiversity and Environmental Conservation; Universiti Malaysia Sarawak; 94300 Kota Samarahan; Sarawak; Malaysia.; Naturalis Biodiversity Center; P.O. Box 9517; 2300 RA Leiden; The Netherlands.
Program Studi Biologi Fakultas MIPA Dan Kesehatan; Universitas Muhammadiyah Riau; Indonesia.
Restorasi Ekosistem Riau; Pangkalan Kerinci; Riau.
Research Center for Biosystematics and Evolution; National Research and Innovation Agency (BRIN); Cibinong; Kabupaten Bogor; Jawa Barat 16911; Indonesia.; Department of Biology; Brigham Young University; Provo; UT 84602; USA.
Restorasi Ekosistem Riau; Pangkalan Kerinci; Riau.
Program Studi Biologi Fakultas MIPA Dan Kesehatan; Universitas Muhammadiyah Riau; Indonesia.
Restorasi Ekosistem Riau; Pangkalan Kerinci; Riau.
Odonata Indonesia Sumatra Riau Peat Swamp Forest Gynacantha

Abstract

Sumatra, the second largest island in Sundaland, has been poorly studied for Odonata. Sumatra has ca. 72,300 km2 of peatlands which, prior to human alteration, were almost entirely covered by Peat Swamp Forest (PSF); Riau Province has the largest peatlands on the island. Less than 20% of Sumatra’s peatlands remain as PSF (pristine or degraded). PSF is known to provide habitat for specialised and/or threatened plant and animal species, including Odonata, but has received little sampling effort. No study on Odonata of any PSF area in Sumatra has been reported until now. The Kampar Peninsula in Riau Province has the largest remaining PSF (3,445.73 km2) in Sumatra, with 1,300.95 km2 specifically managed by the Restorasi Ekosistem Riau (RER) programme with the aim of protecting and restoring the degraded PSF at the core of the peninsula. In early 2020 the first author, together with Universitas Muhammadiyah Riau, commenced surveys of Odonata in the Kampar Peninsula, funded and facilitated by RER. After a long pause due to the global Covid-19 pandemic, surveys resumed in 2023. All surveys were conducted under research permits issued by the Indonesian National Research and Innovation Agency. The results of the Odonata surveys in the Kampar Peninsula are presented here. Surveys were conducted in January 2020, February 2023, May/June 2023 and September/October 2023, the spacing of and weather conditions during the surveys are discussed. Ninety-seven individual sites where specimens were collected are listed, grouped into 19 general areas. One hundred species and subspecies recorded during the surveys are listed with notes on habitat and in some cases other remarks. Taxonomic notes and remarks are given for some of the taxa recorded (Podolestes buwaldai Lieftinck, 1940, Copera vittata (Selys, 1863), Mortonagrion sp. cf. aborense (Laidlaw, 1914), M. megabinluyog Dow & Choong, 2015, M. sp. cf. forficulatum Lieftinck, 1953, Teinobasis ruficollis (Selys, 1877), T. sp. cf. suavis Lieftinck, 1953, Gynacantha dohrni Krüger, 1899 and G. maclachlani Krüger, 1899 (under the first name), Heliaeschna crassa Krüger, 1899 and H. idae (Brauer, 1865) (under the first name). In particular three forms of penile organ found in the M. aborense complex in Sumatra are discussed and illustrated with sketches; the structure of the female prothorax in T. ruficollis is discussed; the taxonomic history, status of type material, and diagnostic characters of G. dohrni and G. maclachlani are discussed in detail and illustrated with stacking images of the male anal appendages of type material of both species. Ninety-four of the taxa recorded are identified to a named species or subspecies, five are only identified with certainty to genus level, pending further taxonomic work and collection of more samples, and one taxon is identified as an unnamed but already known species. Of the distinct taxa recorded, 42 are from the Zygoptera and 58 from the Anisoptera. Most sampled habitats can be classified as either lowland swamp forest or open and disturbed; typically for both classes of habitat, the majority of species recorded are from the Coenagrionidae and Libellulidae. Eighty-nine of the 94 taxa identified to a named species/subspecies have also been recorded in Peninsular Malaysia and/or Singapore. At least 38 of the taxa recorded represent first published records for either Indonesia (five species: Amphicnemis bebar Dow, Choong & Ng, 2010, Amphicnemis hoisen Dow, Choong & Ng, 2010, Amphicnemis reri Dow, Gesriantuti & Lupiyaningdyah, 2025, M. megabinluyog and Chalybeothemis chini Dow, Choong & Orr, 2007), Sumatra including satellite islands (two taxa), mainland Sumatra (six taxa) or Riau Province (25 taxa); these first records are discussed. With these results 271 species of Odonata are known from Sumatra and 126 from Riau Province. Of the 100 distinct taxa recorded in the surveys in the Kampar Peninsula we regard 49 as forest-dependent (requiring the presence of forest for their survival) and 14 as specialists of low pH waters. A discussion of the species found in natural forest, planted forest and open habitats is included. Fifteen of the species recorded currently have IUCN Red List assessments other than Least Concern (four Data Deficient, one Endangered, three Vulnerable and seven Near Threatened. One species, A. reri, has not yet been assessed but would likely fall in one of the threatened categories with the data available. Fifteen percent is a high proportion of non-Least Concern species for a sample of Odonata in a relatively small area but fairly typical for low pH water habitats in Sundaland; all but one of the species regarded as low pH specialists have non-Least Concern Red List assessments. A comparison of the odonate fauna of the Kampar Peninsula with that of PSF areas in Borneo and Peninsular Malaysia is made and odonate diversity at low pH waters in Sundaland is discussed.

 

References

  1. Alfarisyi, A. (2018) Odonata survey at Juru Seberang, Belitung Island, Indonesia. Agrion, 22 (1), 42–47.
  2. Alfarisyi, A. (2019) Odonata survey on some of the outer islands of Belitung Regency, Belitung island, Indonesia. Faunistic Studies in Southeast Asian and Pacific Island Odonata, 29, 1–34.
  3. Asahina, S. (1982) A list of the Odonata from Thailand. Part I. Agrionidae. Kontyû, 50 (3), 454–466.
  4. Asahina, S. (1986) A list of the Odonata from Thailand. Part XV. Aeschnidae. Tombo, 29 (3/4), 71–106.
  5. Brauer, F.M. (1864) Erster Bericht über die auf der Weltfahrt der kais. Fregatte Novara gesammelten Neuropteren. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 14, 159–164.
  6. Brauer, F.M. (1865a) Dritter Bericht über die auf der weltfarht der kais. Fregatte Novara Gesammelten Libellulinen. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 15, 501–512.
  7. Brauer, F.M. (1865b) Vierter Bericht über die auf der weltfarht der kais. Fregatte Novara Gesammelten Neuropteren. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 15, 903–908.
  8. Brauer, F.M. (1865c) Herrn Baron Ransonett am rothen Meer und auf Ceylon Gesammelten Neuropteren (L.). Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 15, 1009–1018.
  9. Brauer, F.M. (1867a) Beschreibungen neuer exotisher Libellen aus den Gattungen Neurothemis, Libellula, Diplax, Celithemis und Tramea. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 17, 3–26.
  10. Brauer, F.M. (1867b) Neue exotishe Odonaten. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 17, 811–816.
  11. Brauer, F.M. (1868a) Neue und wenig Bekannte vom. Herrn Doct. Semper gesammelten Odonaten. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 18, 167–188.
  12. Brauer, F.M. (1868b) Verzeichniss der biz jetz bekannten Neuropteran im Sinne Linné’s. Zweiter Abschnitt. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 18, 359–416.
  13. Brauer, F.M. (1868c) Dritter Bericht über die von Herrn G. Semper mitgeteilten, von dessen Bruder auf den Philippinen gesammelten Neuropteren und Beschreibung einer neuen Libellen-Gattung. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 18, 541–558.
  14. Brauer, F.M. (1878) Über einige neue Gattungen und Artena us der Ordnung der Neuropteren Lin. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe Abt. 1 Mineralogie, Botanik, Zoologie, Geologie und Paläontologie, 77, 193–206.
  15. Burmeister, C.H.C. (1839) Handbuch der Entomologie. Neuroptera. 2 (2). Reimer, Berlin, xii + 294 pp. [pp. i–xii + 757–1050]
  16. de Charpentier, T. (1840) Libellulinae Europaeae descriptae ac depictae. Voss, Lipsiae, 2 + 108 pp. https://doi.org/10.5962/t.173037
  17. Choong, C.Y. (2013) Odonata of North Selangor Peat Swamp Forest, Selangor. In: Prosiding Ekspedisi Saintifik Biodiversiti Hutan Paya Gambut Selangor Utara. ASEAN Peatland Forest Programme, pp. 183–189.
  18. Choong, C.Y. & Cheah, D.S.L. (2013) Odonata of Ayer Hitam Forest Reserve, Johor, Peninsular Malaysia. Faunistic Studies in South–East Asian and Pacific Island Odonata, 2, 1–11.
  19. Choong, C.Y., Dow, R.A. & Ng, Y.F. (2017) New records of Odonata from Kelantan, Malaysia, with a checklist of species recorded from the state. Faunistic Studies in Southeast Asian and Pacific Island Odonata, 21, 1–22.
  20. Choong, Y.C., Dow, R.A. & Ng, Y.F. (2020) Additional records of Odonata from Kelantan and Terengganu, Malaysia. International Dragonfly Fund Report, 144, 1–26.
  21. Choong, C.Y., Izzat-Husna, M. & Amirrudin, B.A. (2016) Odonata (Insecta) fauna of Tasek Bera Ramsar Site, Pahang, Peninsular Malaysia. Journal of Wildlife and Parks, 31, 39–48. https://doi.org/10.64291/80e6z252
  22. Choong, C.Y., Orr, A.G. & Dow, R.A. (2008) Checklist of dragonflies of UKM Campus. including Bangi Forest Reserve, Bangi, Selangor, Malaysia. Echo, 5, 4–5. [in Agrion 12 (1)]
  23. Cleary, D.F.R., Moers, A.Ø., Eichhorn, K.A.O., van Tol, J., de Jong, R. & Menken, S.B.J. (2004) Diversity and community composition of butterflies and odonates in an ENSO–induced fire affected habitat mosaic: a case study from East Kalimantan, Indonesia. Oikos, 105, 426–446. https://doi.org/10.1111/j.0030-1299.2004.12219.x
  24. Cowley, J. (1934) Changes in the Generic Names of the Odonata. The Entomologist, 67, 200–205.
  25. Cowley, J. (1935) A New Generic Name in the Protoneuridae (Odonata). The Entomologist’s Monthly Magazine, 71, 14–15.
  26. Dosi, E.M., Grinang, J., Nyanti, L., Khoon, K.L., Harun, M.H. & Kamarudin, N. (2018) A preliminary study of the macroinvertebrate fauna of freshwater habitats in Maludam National Park, Sarawak. Mires and Peat, 22, 1–7.
  27. Dow, R.A. (2010) A review of the Teinobasis of Sundaland, with the description of Teinobasis cryptica sp. nov. from Malaysia (Odonata: Coenagrionidae). International Journal of Odonatology, 13 (2), 205–230, pl. II. https://doi.org/10.1080/13887890.2010.9748375
  28. Dow, R.A. (2011) Mortonagrion indraneil spec. nov. from Borneo, and a redescription of M. arthuri Fraser (Odonata: Zygoptera: Coenagrionidae). Zootaxa, 3093 (1), 35–46. https://doi.org/10.11646/zootaxa.3093.1.2
  29. Dow, R.A. (2016) Odonata collected at the Samunsam Wildlife Sanctuary, Kuching Division, Sarawak, Malaysia in August 2015. Faunistic Studies in South–East Asian and Pacific Island Odonata, 14, 1–12.
  30. Dow, R.A. (2021) An annotated checklist of the Odonata (Insecta) known from Sarawak with records to district level. Sarawak Museum Journal, LXXX1, No. 101 (New Series), Special Issue 10, 313–422.
  31. Dow, R.A. (2022) Previously unpublished Odonata records from Sarawak, Borneo, part VIII: New records from Kapit Division. International Dragonfly Fund Report, 169, 1–13.
  32. Dow, R.A., Advento, A.D., Turner, E.C., Caliman, J.-P., Foster, W.A., Naim, M., Snaddon, J.L. & Ps, S. (2018) Odonata from the BEFTA Project area, Riau Province, Sumatra, Indonesia. Faunistic Studies in South-East Asian and Pacific Island Odonata, 24, 1–22.
  33. Dow, R.A., Ahmad, R., Butler, S.G., Choong, C.Y., Grinang, J., Ng, Y.F., Ngiam, R.W.J., Reels, G.T., Steinhoff, P.O.M. & Unggang, J. (2021) Previously unpublished Odonata records from Sarawak, Borneo, part VI: Miri Division including checklists for Niah, Lambir Hills, Loagan Bunut and Pulong Tau National Parks. Faunistic Studies in South–East Asian and Pacific Island Odonata, 36, 1–94.
  34. Dow, R.A., Butler, S.G., Ngiam, R.W.J. & Reels, G.T. (2021) Previously unpublished Odonata records from Sarawak, Borneo, part V: Odonata from the southwest of Sarawak, including the first records from the Bungo Range National Park. International Dragonfly Fund Report, 159, 1–77.
  35. Dow, R.A. & Choong, C.Y. (2015) Mortonagrion megabinluyog spec. nov. from Brunei (Odonata: Zygoptera: Coenagrionidae). Zootaxa, 3914 (1), 89–93. https://doi.org/10.11646/zootaxa.3914.1.8
  36. Dow, R.A., Choong, C.Y., Grinang, J., Lupiyaningdyah, P., Ngiam, R.W.J. & Kalkman, V.J. (2024) Checklist of the Odonata (Insecta) of Sundaland and Wallacea (Malaysia, Singapore, Brunei, Indonesia and Timor Leste). Zootaxa, 5460 (1), 1–122. https://doi.org/10.11646/zootaxa.5460.1.1
  37. Dow, R.A., Choong, C.Y. & Ng, Y.F. (2010) A review of the genus Amphicnemis in Peninsular Malaysia and Singapore, with descriptions of two new species (Odonata: Zygoptera: Coenagrionidae). Zootaxa, 2605 (1), 45–55. https://doi.org/10.11646/zootaxa.2605.1.3
  38. Dow, R.A., Choong, C.Y. & Orr, A.G. (2007) Two new species of Chalybeothemis from Malaysia, with a redefinition of the genus (Odonata: Libellulidae). International Journal of Odonatology, 10 (2), 171–184. https://doi.org/10.1080/13887890.2007.9748297
  39. Dow, R.A., Gesriantuti, N. & Lupiyaningdyah, P. (2025) A new species of Amphicnemis Selys, 1863 from Sumatra, Indonesia and first records of two other species of Amphicnemis for the island (Odonata: Zygoptera: Coenagrionidae). Zootaxa, 5659 (2), 273–282. https://doi.org/10.11646/zootaxa.5659.2.8
  40. Dow, R.A. & Morris, J.R. (2021) The old Odonata (Insecta) Collection of the Sarawak Museum. Sarawak Museum Journal, LXXX1, No. 101 (New Series), Special Issue 10, 261–311.
  41. Dow, R.A. & Ng, Y.F. (2020) New records of Odonata from Selangor and Negeri Sembilan, Malaysia, with provisional checklists of species recorded from the states. International Dragonfly Fund Report, 146, 1–29.
  42. Dow, R.A., Ng, Y.F. & Choong, C.Y. (2012) Odonata of Sungai Bebar, Pahang, Malaysia, with four species recorded for the first time from mainland Asia. Journal of Threatened Taxa, 4 (3), 2417–2426. https://doi.org/10.11609/JoTT.o3041.2417-26
  43. Dow, R.A. & Ngiam, R.W.J. (2011) Two damselflies new to Singapore: Amphicnemis bebar Dow, Choong & Ng and Teinobasis cryptica Dow (Odonata: Zygoptera: Coenagrionidae). Nature in Singapore, 4, 393–396.
  44. Dow, R.A., Ngiam, R.W.J. & Ahmad, R. (2015) Odonata of Maludam National Park, Sarawak, Malaysia. Journal of Threatened Taxa, 7 (1), 6764–6773. https://doi.org/10.11609/JoTT.o4107.6764-73
  45. Dow, R.A. & Reels, G.T. (2011) Odonata from a remnant patch of disturbed peatswamp forest on the outskirts of Kuching, west Sarawak. Agrion, 15 (2), 50–51.
  46. Dow, R.A. & Reels, G.T. (2024) Previously unpublished Odonata records from Sarawak, Borneo, part X: Maludam and Ulu Sebuyau National Parks, with notes on Pseudagrionoptera diotima Ris, 1912. Faunistic Studies in South–East Asian and Pacific Island Odonata, 44, 1–18.
  47. Dow, R.A. & Silvius, M. (2014) Results of an Odonata survey carried out in the peatlands of Central Kalimantan, Indonesia, in 2012. Faunistic Studies in South–East Asian and Pacific Island Odonata, 7, 1–37.
  48. Dow, R.A. & Singa, H. (2022) Previously unpublished Odonata records from Sarawak, Borneo. Part IX. More Odonata from Limbang Division, including the first records from Gunung Buda National Park. International Dragonfly Fund Report, 173, 1–32.
  49. Dow, R.A. & Unggang, J. (2010) The Odonata of Binyo Penyilam, a unique tropical wetland area in Bintulu Division, Sarawak, Malaysia. Journal of Threatened Taxa, 2 (13), 1349–1358. https://doi.org/10.11609/JoTT.o2478.1349-58
  50. Drury, D. (1773) Illustrations of natural History: wherein are exhibited upwards of two hundred and forty figures of exotic insects, according to their different genera: very few of which have hitherto been figured by any author, being engraved and coloured from nature, with the greatest accuracy, and under the authors own inspection, on fifty copper-plates; with a particular description of each insect ... to which is added a translation into French. Vol. 2. White, London, vii + 90 pp.
  51. Erichson, W.F. (1848) Insecten. In: Schomburgk, R.H. (Ed.), Reisen in Britisch-Guiana in den Jahren 1840-1844: nebst einer Fauna und Flora Guiana’s nach Vorlagen von Johannes Müller, Ehrenberg, Erichson, Klotzsch, Troschel, Cabanis und Andern. Weber, Leipzig, 1260 pp.
  52. Fabricius, J.C. (1793) Entomologia Systematica Emedata et Aucta. Secundum classes, ordines, genera, species, adjectis synonymis, locis, observationibus, descriptionibus. Vol. 2. C. G. Proft, Copenhagen, viii + 519 pp. https://doi.org/10.5962/bhl.title.125869
  53. Fabricius, J.C. (1798) Entomologia Systematica Emedata et Aucta. Secundum classes, ordines, genera, species, adjectis synonymis, locis, observationibus, descriptionibus. Vol. 4. C. G. Proft, Copenhagen, 472 pp.
  54. Fraser, F.C. (1919) Descriptions of four new Indian Odonata. Records of the Indian Museum, 16, 451–455. https://doi.org/10.5962/bhl.part.25933
  55. Fraser, F.C. (1920) Some new Indian Dragonflies. Journal of the Bombay Natural History Society, 27 (1), 147–150.
  56. Fraser, F.C. (1922a) New and rare Indian Odonata in the Pusa collection. Memoirs of the Department of Agriculture in India, Entomology, 3 (7), 39–77.
  57. Fraser, F.C. (1922b) A collection of dragonflies from Bangkok. Journal of the Natural History Society of Siam, 4, 231–238.
  58. Fraser, F.C. (1926) Notes on a collection of Dragonflies (Order Odonata) from the Dutch East Indies and descriptions of four new species from the neighbouring continent. Treubia, VIII (3–4), 467–494. https://doi.org/10.14203/treubia.v8i3-4.1762
  59. Gesriantuti, N., Herlina, N. & Yunita, N. (2018) Jenis-jenis Odonata di kawasan Stadion Utama Riau, Pekan Baru. Jurnal Photon, 9 (1), 197–202. https://doi.org/10.37859/jp.v9i1.1081
  60. Hagen, H.A. (1861) Synopsis of the Neuroptera of North America, with a List of the South American Species, prepared for the Smithsonian Institution. Smithsonian Institute, Washington, D.C., 20 + 347 pp. https://doi.org/10.5962/bhl.title.22092
  61. Hagen, H.A. (1867a) Revision der von Herm Uhler beschriebenen Odonaten. Stettiner Entomologische Zeitung, 28, 87–95.
  62. Hagen, H.A. (1867b) Die Neuroptera der Insel Cuba. Stettiner Entomologische Zeitung, 28, 215–232.
  63. Hämäläinen, M. (1989) Synonymic notes on some species of the genus Mortonagrion (Zygoptera: Coenagrionidae). Indian Odonatology, 2, 1–4.
  64. Hämäläinen, M. (2000) Ten species added to the list of Peninsular Malaysian Odonata. Notulae odonatologicae, 5 (5), 53–55.
  65. Husson, S.J., Limin, S.H., Adul, Boyd, N.S., Brousseau, J.J., Collier, S., Cheyne, S.M., D’Arcy, L.J., Dow, R.A., Dowds, N.W., Dragiewicz, M.L., Ehlers Smith, D.A., Iwan, Hendri, Houlihan, P.R., Jeffers, K.A., Jarrett, B.J.M., Kulu, I.P., Morrogh-Bernard, H.C., Page, S.E., Perlett, E.D., Purwanto, A., Ripoll Capilla, B., Salahuddin, Santiano, Schreven, S.J.J., Struebig, M.J., Thornton, S.A., Tremlett, C., Yeen, Z. & Harrison, M.E. (2018) Biodiversity of the Sebangau tropical peat swamp forest, Indonesian Borneo. Mires and Peat, 22, 1–50. https://doi.org/10.19189/MaP.2018.OMB.352
  66. Kalkman, V.J. (2004) From cool hill resorts to humid rainforest: an odonatological trip to Peninsular Malaysia (July 2002). Echo, 1, in Agrion 8 (2), 26–28.
  67. Karsch, F.A.F. (1890) Beitrage zur Kenntniss der Arten und Gattungen der Libellulinen. Berliner entomologische Zeitschrift, 33, 347–392. https://doi.org/10.1002/mmnd.18900330221
  68. Karsch, F.A.F. (1891) Sumatranische Odonaten, gesammelt von Herrn Hofrath Dr. med. L. Martin in Bindjei (Deli). Entomologische Nachrichten, 17 (16), 241–247.
  69. Karsch, F.A.F. (1900) Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo. Zweiter Teil: Wissenschaftliche Reiseergebnisse. Band III. Odonaten. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 25, 211–230.
  70. Kirby, W.F. (1889a) A Revision of the Subfamily Libellulinae, with Descriptions of New Genera and Species. Transactions of the Zoological Society of London, 12 (9), 249–343. https://doi.org/10.1111/j.1096-3642.1889.tb00016.x
  71. Kirby, W.F. (1889b) Descriptions of new Genera and Species of Odonata in the Collection of the British Museum, chiefly from Africa. Proceedings of the Zoological Society of London, 1889, 297–303.
  72. Kirby, W.F. (1890) A Synonymic Catalogue of Neuroptera Odonata or Dragonflies, with an appendix of fossil species. Gurney & Jackson, London, ix + 202 pp. https://doi.org/10.5962/bhl.title.5534
  73. Kirby, W.F. (1894) On a small collection of Odonata (Dragonflies) from Queensland, with descriptions of five new species. Annals and Magazine of Natural History, Series 6, 14, 15–21. https://doi.org/10.1080/00222939408677758
  74. Kirby, W.F. (1900) On the species which have been included in Zygonyx, Hagen and De Selys. Annals and Magazine of Natural History, Series 7, 5, 539–542. https://doi.org/10.1080/00222930008678328
  75. Krüger, L. (1898) Die Odonaten von Sumatra. I. Theil. Familie Agrioniden. Stettiner Entomologische Zeitung, 59, 64–139.
  76. Krüger, L. (1899) Die Odonaten von Sumatra II. Theil. Familie Aeschniden. Stettiner Entomologische Zeitung, 59, 267–331.
  77. Krüger, L. (1902) Die Odonaten von Sumatra. III. Theil b. Familie Libelluliden. Stettiner Entomologische Zeitung, 63, 58–193.
  78. Laidlaw, F.F. (1912) List of the Odonata taken on an expedition to Mt. Batu Lawi together with descriptions of supposed new species. Journal of the Straits Branch of the Royal Asiatic Society, 63, 92–99, 1 pl.
  79. Laidlaw, F.F. (1914a) Odonata of the Abor Expedition. Records of the Indian Museum, 8, 335–349. https://doi.org/10.26515/rzsi/v8/i4/1914/163176
  80. Laidlaw, F.F. (1914b) Contributions to a study of the dragonfly fauna of Borneo. Part II. The Gomphinae and Chlorogomphinae. Proceedings of the Zoological Society of London, 1914, 51–63, pl. I. https://doi.org/10.1111/j.1096-3642.1913.tb01984.x
  81. Laidlaw, F.F. (1919) A list of the dragonflies recorded from the Indian Empire, Part II (cont.). The family Agrionidae. B. The Legions Argia and Agrion. Records of the Indian Museum, 16, 169–195. https://doi.org/10.5962/bhl.part.25920
  82. Laidlaw, F.F. & Förster, F. (1907) Report on the dragon flies. II. The Legions Platycnemis and Protoneura. Fasciculi Malayenses, Zoology, 4, 1–15.
  83. Leach, W.E. (1815) Entomology. In: Brewster, D. (Ed.), The Endinburgh Encyclopedia, 9, 57–172.
  84. Lieftinck, M.A. (1929) Contributions to the Dragonfly fauna of the Sondaic Area. Tijdschrift voor Entomologie, 72, 109–147.
  85. Lieftinck, M.A. (1930) Contributions to the dragonfly–fauna of the Dutch East Indies II. Treubia, 12 (2), 135–166.
  86. Lieftinck, M.A. (1931) A revision of the genus Epophthalmia Burm. (Odon., Corduliinae) with notes on habitats and larvae. Treubia, 13 (1), 21–80, 29 figs., pl. 1. https://doi.org/10.14203/treubia.v13i1.1930
  87. Lieftinck, M.A. (1933a) A new genus and species of Libellulinae from western Borneo (Odonata). Sarawak Museum Journal, 4 (13), 131–137.
  88. Lieftinck, M.A. (1934) An annotated list of the Odonata of Java, with notes on their distribution, habits and life-history. Treubia, 14, 377–462.
  89. Lieftinck, M.A. (1935a) A Synopsis of the Odonata (Dragonflies) of Sumatra. Miscellanea Zoologica Sumatrana, 92–93, 1–23.
  90. Lieftinck, M.A. (1935b) New and little known Odonata of the Oriental and Australian Regions. Treubia, 15 (2), 175–207.
  91. Lieftinck, M.A. (1937) Descriptions and records of South–east Asiatic Odonata. Treubia, 16, 55–119. https://doi.org/10.14203/treubia.v16i1.2504
  92. Lieftinck, M.A. (1940a) Revisional notes on some species of Copera Kirby. With notes on habits and larvae (Odon., Platycnemididae). Treubia, 17 (4), 281–306. https://doi.org/10.14203/treubia.v17i4.2577
  93. Lieftinck, M.A. (1940b) Descriptions and records of South-east Asiatic Odonata (II). Treubia, 17, 337–390. https://doi.org/10.14203/treubia.v17i4.2581
  94. Lieftinck, M.A. (1948a) Descriptions and records of South-east Asiatic Odonata. Treubia, 19, 221–278.
  95. Lieftinck, M.A. (1948b) The Odonata of Engano with a survey of the dragonfly—fauna of the West Sumatra chain of islands. Treubia, 19 (2), 279–304.
  96. Lieftinck, M.A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938–1939 and of the Le Roux Expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea New Series, 5, 1–271.
  97. Lieftinck, M.A. (1950) Two new species of Podolestes Selys from Borneo, with a key for the identification of the known species (Odonata, Megapodagriidae). Zoologische Mededelingen, Leiden, 31 (4), 39–47.
  98. Lieftinck, M.A. (1951) The Identity of some Malaysian Species of Ceriagrion, with Descriptions of Two New Species (Odon.). Treubia, 21 (1), 183–197. https://doi.org/10.14203/treubia.v21i1.2652
  99. Lieftinck, M.A. (1953a) Additions to the odonate fauna of the Indo-Australian Archipelago. Treubia, 22 (1), 233–269. https://doi.org/10.14203/treubia.v22i1.1567
  100. Lieftinck, M.A. (1953b) New dragonflies (Odonata) from Borneo, with notes on their habits and larvae. Treubia, 22 (2), 381–406. https://doi.org/10.14203/treubia.v22i2.2679
  101. Lieftinck, M.A. (1954) Handlist of Malaysian Odonata. A catalogue of the dragonflies of the Malay Peninsula, Sumatra, Java and Borneo, including the adjacent small islands. Treubia Supplement, 22, i–xiii + 1–202, 1 folded map excl.
  102. Lieftinck, M.A. (1965) The species–group of Vestalis amoena Selys, 1853, in Sundaland (Odonata, Calopterygidae). Tijdschrift voor Entomologie, 108 (11), 325–364.
  103. Martin, R. (1909) Aeschnines III. Collections Zoologiques du Baron Edm. De Selys Longchamps. Catalogue Systématique et Descriptif, 20, 157–223, pls. V–VI.
  104. Martin, R. (1911) Odonata Fam. Aeschnidae Subfam. Aeschninae. In: Wytsman, P. (Ed.), Genera Insectorum, 115, pp. 1–34, pls. 1–6.
  105. McLachlan, R. (1895) Some new species of Odonata of the “Légion” Lestes, with notes. Annals and Magazine of Natural History, Series 6, 16 (91), 19–28. https://doi.org/10.1080/00222939508680221
  106. Miettinen, J., Shi, C. & Liew, S.C. (2016) Land cover distribution in the peatlands of Peninsular Malaysia, Sumatra and Borneo in 2015 with changes since 1990. Global Ecology and Conservation, 6, 67–78. https://doi.org/10.1016/j.gecco.2016.02.004
  107. Müller, O.F. (1764) Fauna insectorum fridrichsdalina, sive, Methodica descriptio insectorum agri fridrichsdalensis : cum characteribus genericis et specificis, nominibus trivialibus, locis natalibus, inconibus allegatis, novisque pluribvs speciebus additis. In officina libraria I.F. Gleditschii, Hafniae et Lipsiae, xxiv + 96 pp.
  108. Newman, E. (1833) The Entomological Magazine. Vol. 1. Westley, London, 511 pp.
  109. Ngiam, R.W.J., Dow, R.A., Thio, H.B. & Chan, W.T. (2024) Biodiversity Record: New Singapore record of the dragonfly, Tetrathemis flavescens, with a note on Tetrathemis hyalina in Singapore. Nature in Singapore, 17, 1–3. https://doi.org/10.26107/NIS-2024-0082
  110. Norma-Rashid, Y., Mohd-Sofian, A. & Zakaria-Ismail, M. (2001) Diversity and distribution of Odonata (dragonflies and damselflies) in the freshwater swamp lake Tasek Bera, Malaysia. Hydrobiologica, 459, 135–146. https://doi.org/10.1023/A:1012562611307
  111. Orr, A.G. (2001) An annotated checklist of the Odonata of Brunei with ecological notes and descriptions of hitherto unknown males and larvae. International Journal of Odonatology, 4, 167–220. https://doi.org/10.1080/13887890.2001.9748168
  112. Orr, A.G. (2003) A guide to the dragonflies of Borneo: their identification and biology. Natural History Publications (Borneo), Kota Kinabalu, x + 195 pp.
  113. Orr, A.G. (2006) Odonata in Bornean tropical rain forest formations: diversity, endemicity and implications for conservation management. In: Rivera, A.C. (Ed.), Forests and Dragonflies. Fourth WDA International Symposium of Odonatology, Pontevedra (Spain), July 2005. Pensoft Publishers, Sofia-Moscow, pp. 51–78.
  114. Pinhey, E. (1974) A revision of the African Agriocnemis Selys and Mortonagrion Fraser (Odonata Coenagrionidae). Occasional papers of the National Museums and Monuments of Rhodesia Series B Natural Sciences, 5 (4), 171–278.
  115. Posa, M.R.C., Wijedasa, L.S. & Corlett, R.T. (2011) Biodiversity and Conservation of tropical Peat Swamp Forests. BioScience, 61 (1), 49–57. https://doi.org/10.1525/bio.2011.61.1.10
  116. Pujiastuti, Y., Windusari, Y. & Agus, M. (2017) The distribution and composition of Odonata (Dragonfly and Damselfly) in Sriwijaya University, Inderalaya Campus South Sumatera. Journal of Biological Researches, 23 (1), 1–5. https://doi.org/10.23869/bphjbr.23.1.20171
  117. Rambur, J.P. (1842) Histoire Naturelle des Insectes. Névroptès. Roret, Paris, 17 + 534 pp.
  118. Ris, F. (1909) Libellulinen 1. Collections Zoologiques du Baron Edm. De Selys Longchamps. Catalogue Systématique et Descriptif, 9, 1–120, figs. 1–89, pl. I.
  119. Ris, F. (1911) Libellen von Sintang, Borneo. Annales de la Societe Entomologique de Belgique, 55, 231–255.
  120. Ris, F. (1912) Über Odonaten von Java und Krakatau gessammelt Edward Jacobson. Tijdschrift voor Entomologie, 55, 157–183.
  121. Ris, F. (1913) Libellulinen 7. Collections Zoologiques du Baron Edm. De Selys Longchamps. Catalogue Systématique et Descriptif, 15, 837–964, VII pls.
  122. Ris, F. (1915) Fauna simalurensis. Odonata. Tijdschrift voor Entomologie, 58, 5–21.
  123. Ris, F. (1919) Libellulinen 9. Collections Zoologiques du Baron Edm. De Selys Longchamps. Catalogue Systématique et Descriptif, 16, 1043–1278.
  124. Ris, F. (1927) Odonaten von Sumatra, gesammelt von Edward Jacobson. Zoologische Mededelingen, 10 (1), 1–49.
  125. Ris, F. (1930) Drei Notizen über ostasiatische Agrioniden (Odonata). Arkiv förZoologi, Bd. 21 A, 31, 1–32.
  126. St. Quentin, D. (1968) Odonaten vom Lingga-Archipel. Annalen des Naturhistorischen Museums in Wien, 72, 389–390.
  127. Schmidt, E. (1934) Odonata der Deutschen Limnologischen Sunda-Expedition. 1. Imagines. Mit Beschreibungen zweier neuer Ictinus aus Celebes und Neu-Guinea. Archiv für Hydrobiologie, Suppl. Bd., 13, 316–397, pls. 14–16.
  128. Seehausen, M.A. & Dow, R.A. (2016) Morphological studies and taxonomic considerations on the ‘reddish-brown-winged’ group of Neurothemis Brauer, 1867 with the description of N. taiwanensis sp. nov. (Odonata: Libellulidae). International Dragonfly Fund Report, 93, 1–101.
  129. de Selys Longchamps, E. de (1840) Monographie des Libellulidées d’Europe. Roret, Paris, 220 pp.
  130. de Selys Longchamps, E. de (1853) Synopsis des Caloptérygines. Bulletins de l’Académie royale de Belgiquè, Série 1, 20 (Annex), 73 pp.
  131. de Selys Longchamps, E. de (1854) Synopsis des Gomphines. Bulletins de l’Académie royale de Belgiquè, Série 1, 21 (2), 23–112.
  132. de Selys Longchamps, E. de (1859) Additions au synopsis des Caloptérygines. Bulletins de l’Académie royale de Belgiquè, Série 2, 7 (7), 437–451.
  133. de Selys Longchamps, E. de (1860) Synopsis des Agrionines, derniere legion: Protoneura. Bulletins de l’Académie royale de Belgiquè, Série 2, 10, 431–462. https://doi.org/10.5962/bhl.title.66021
  134. de Selys Longchamps, E. de (1862a) Synopsis des Agrionines, seconde légion: Lestes. Bulletins de l’Académie royale des sciences, des Lettres et des Beaux–Arts de Belgique, Série 2, 13 (4), 288–338.
  135. de Selys Longchamps, E. de (1862b) Synopsis des Agrionines, 3me légion: Agrion (suite et fin). Podagrion. Bulletins de l’Académie royale des sciences, des Lettres et des Beaux–Arts de Belgique, Série 2, 14 (2), 3–42.
  136. de Selys Longchamps, E. de (1863) Synopsis des Agrionines. Quatrième légion: Platycnemis. Bulletins de l’Académie royale de Belgiquè, Série 2, 16 (8), 147–176.
  137. de Selys Longchamps, E. de (1865) Synopsis des Agrionines. 5e légion: Agrion. Bulletins de l’Académie royale de Belgiquè, Série 2, 20, 375–417.
  138. Selys-Longchamps, E. de (1870) Sous-famille des Cordulines, Sélys (1). In: Comptes-rendus des séances de la société entomologique de Belgique: Assemblée mensuelle du 5 novembre 1870. Annales de la Société entomologique de Belgique, 14, iv–vii. https://doi.org/10.5962/bhl.part.13349
  139. de Selys Longchamps, E. de (1872) Note sur plusieurs Odonates de Madagascar et des Iles Mascareignes. Revue zoologique, Series 2, 23, 175–183.
  140. de Selys Longchamps, E. de (1873) Troisiemes additions au synopsis des Gomphines. Bulletins de l’Académie royale de Belgiquè, Série 2, 35 (6), 732–774.
  141. de Selys Longchamps, E. de (1876) Synopsis des Agrionines. 5e légion: Agrion. Le Genre Agrion. Bulletins de l’Académie royale de Belgiquè, Série 2, 41, 247–322 + 496–539 + 1233–1309.
  142. de Selys Longchamps, E. de (1877) Synopsis des Agrionines (suite et fin). (5me légion: Agrion). Les trois grands genres Telebasis, Argiocnemis et Hemiphlebia. Bulletin de l’Académie royale de Belgique, Série 2, 43 (2), 97–159.
  143. de Selys Longchamps, E. de (1878a) Odonates de la région de la Nouvelle Guinée. Mittheilungen aus dem Königlichen zoologischen Museum zu Dresden, 3, 289–323.
  144. de Selys Longchamps, E. de (1878b) Quatrièmes additions au synopsis des gomphines. (I). Bulletins de l’Académie royale Belgique, Série 2, 46, 408–471.
  145. de Selys Longchamps, E. de (1879a) Nouvelles observations sur les Odonates de la region de la Nouvelle-Guinée. Annali del Museo civico di storia naturale di Genova, 14, 287–324.
  146. de Selys Longchamps, E. de (1879b) Quatrièmes additions au synopsis des Caloptérygines. Bulletins de l’Academie royale Belgique, Série 2, 47, 349–409.
  147. de Selys Longchamps, E. de (1882) Sur la distribution des insectes Odonates en Afrique. Compte Rendu de la Session Association française pour l’avancement des sciences (Congrés d’Alger 1881), 10, 663–668.
  148. de Selys Longchamps, E. de (1886) Revision du synopsis des Agrionines. Première partie comprenant les lègions Pseudostigma – Podagrion – Platycnemis et Protonevra. Mémoires couronnés et autres mémoires publiés par l’Académie royale des sciences, des lettres et des beaux-arts de Belgique, 38 (4), 1–233. https://doi.org/10.3406/marb.1886.2330
  149. de Selys Longchamps, E. de (1889) Odonates de Sumatra comprenant les espèces recueillies à Pulo Nias par M. le Dr. E. Modigliani. Annali del Museo Civico di Storia Naturale di Genova, Serie 2, 7, 444–484.
  150. de Selys Longchamps, E. de (1891) Viaggio di Leonardo Fea in Birmania e région vicine. XXXII. Odonates. Annali del Museo civico di storia naturale di Genova, 30, 433–518.
  151. de Selys Longchamps, E. de & Hagen, H.A. (1858) Monographie des Gomphines. Muquardt, Bruxelles & Leipzig, 460 pp., 23 pls. https://doi.org/10.5962/bhl.title.60465
  152. Steinhoff, P.O.M., Butler, S.G. & Dow, R.A. (2016) Description of the final instar larva of Orthetrum borneense Kimmins, 1936 (Odonata, Libellulidae), using rearing and molecular methods. Zootaxa, 4083 (1), 99–108. https://doi.org/10.11646/zootaxa.4083.1.5
  153. Sulzer, J.H. (1776) Abgekürtze Geschichte der Insekten nach dem Linnaeischen System. Erster Theil & Zweeter Theil. H. Steiner & Co., Winterthur, xxviii + 274 pp. & 71 pp., pls. I–XXXII excl. https://doi.org/10.5962/bhl.title.11198
  154. Thompson, D.J. & van Tol, J. (1993) Damselflies and dragonflies from four forest types in Brunei. Brunei Museum Journal, 8, 57–72.
  155. Tsuda, S. & Kitagawa, K. (1988) Odonata of southeast Asia collected by the late Mr. M. Iwasaki. Part 2. Odonata of North Sumatra, Indonesia. Gracile, 39, 1–5. [in Japanese with English title]
  156. Weekers, P.H.H. & Dumont, H.J. (2004) A molecular study of the relationship between the coenagrionoid genera Erythromma and Cercion, with the creation of Paracercion Gen. Nov. for the east Asiatic “Cercion” (Zygoptera: Coenagrionidae). Odonatologica, 33 (2), 181–188.
  157. Williamson, E.B. (1907) The dragonflies (Odonata) of Burma and Lower Siam.–II. Subfamilies Cordulegasterinae, Chlorogomphinae, and Gomphinae. Proceedings U.S. National Museum, 33, 267–317. https://doi.org/10.5479/si.00963801.33-1571.267

How to Cite

Dow, R.A., Gesriantuti, N., Iqbal, M., Ingdyah, P.L., Periera, J.F., Zulya, S.O. & Sanders, B.M. (2025) Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa, 5732 (1), 1–70. https://doi.org/10.11646/zootaxa.5732.1.1