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Issue: Vol. 5766 No. 1: 6 Mar. 2026
Type: Monograph
Published: 2026-03-06
DOI: 10.11646/zootaxa.5766.1.1
Page range: 1-87
Abstract views: 97
PDF downloaded: 11

Revision of subgeneric system and species groups of Cephennodes Reitter, with redescriptions of problematic species (Coleoptera: Staphylinidae: Scydmaeninae)

Museum of Natural History; University of Wrocław; Sienkiewicza 21; 50-335 Wrocław; Poland
Coleoptera Museum of Natural History University of Wrocław Poland

Abstract

The subgeneric division and species groups of the large genus Cephennodes Reitter, 1884 are revised. Aculeodes Jałoszyński & Nomura, 2009, syn. n. is placed as a junior synonym of Cephennodes s. str., thus reducing the number of subgenera to two. Cephennodes miharu Hoshina, 2021, syn. n. is placed as a junior synonym of C. kotori Hoshina, 2021; C. mizunoi Hoshina & Kishimoto, 2004, syn. n. as a junior synonym of C. vafer Kurbatov, 1995; C. brachyclavatides Jałoszyński, nom. n. is proposed as a replacement name for C. brachyclavatus Jałoszyński, 2025, preoccupied by C. brachyclavatus Jałoszyński, 2017. The following species are redescribed (some partly, to supplement existing data with new details) and assigned to subgenus or/and a species group: C. (s. str.) tuniseus (Pic, 1901); C. (s. str.) kotori Hoshina, 2021; C. (s. str.) moderatus (Kurbatov, 1995); C. (s. str.) akane Hoshina, 2020; C. (s. str.) maya Hoshina, 2021; C. (s. str.) yume Hoshina, 2017; C. (s. str.) yonaguniensis Hoshina, 2012; C. (s. str.) vafer Kurbatov, 1995; and C. (Fusionodes) otome Hoshina, 2016. Cephennodes tertius Jałoszyński, 2011 and C. leonensis Castellini, 2010 are placed in the nominotypical subgenus, and C. besucheti (Kurbatov, 1995) in the subgenus Fusionodes Jałoszyński, 2007. Cephennodes (s. str.) nanaka Hoshina, 2019 is discussed as a possible junior synonym of C. moderatus, and four species, C. koko Hoshina, 2021, C. nemu Hoshina, 2021, C. otome Hoshina, 2016, and C. sakura Hoshina, 2021 as probably identical with C. loebli (Kurbatov, 1995), all pending further study. Cephennodes s. str. is divided into twenty-two species groups, and the subgenus Fusionodes into six groups: Cephennodes (s. str.) araiorum group (5 spp.), bagongsilangensis group (2 spp.), bos group (8 spp.), cavifrons group (2 spp.), excavatus group (50 spp.), gokarnaensis group (3 spp.), impressifrons group (19 spp.), inflatipes group (4 spp.), latus group (2 spp.), leai group (3 spp.), lustricollis group (5 spp.), mahisapala group (2 spp.), obesus group (4 spp.), oblongopunctatus group (12 spp.), palpalis group (3 spp.), simonis group (21 spp.), simplicipes group (18 spp.), spatulipes group (14 spp.), sulcatus group (3 spp.), taurus group (14 spp.), tuberculifrons group (10 spp.), yunnanensis group (2 spp.), and Cephennodes (Fusionodes) graeseri group (9 spp.), hippopotamus group (9 spp.), loebli group (7 spp.), longilinguis group (4 spp.), microphthalmus group (6 spp.), and serratus group (2 spp.). Keys are given to identify species groups of Cephennodes (s. str.) and the subgenus Fusionodes.

 

References

  1. Besuchet, C. (1962) LXII. Coleoptera Scydmaenidae: Cephenniini. Mission zoologique de l’I.R.S.A.C. en Afrique orientale. P. Basilewsky and N. Leleup, 1957. Annales du Musée Royal du Congo Belge, Tervuren, Série in 8˚: Sciences Zoologiques, 107, 420–423.
  2. Besuchet, C. (1971) 18. Familie: Scydmaenidae. Tribus: Euthiini, Cephenniini. In: Die Kafer Mitteleuropas. Vol. 3. Adephaga 2, Palpicornia, Histeroidea, Staphylinoidea 1. Goecke & Evers, Krefeld, pp. 271–278.
  3. Besuchet, C. (2004) New nomenclatorial and taxonomic acts: Scydmaenidae: Cephenniini. In: Löbl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea – Histeroidea – Staphylinoidea. Apollo Books, Stenstrup, pp. 25–26.
  4. Blattný, C. (1929) Fauna Philippinica, Scydmaenidae, Coleoptera. The Philippine Journal of Science, 38 (3), 303–323, pls. 1–2.
  5. Casey, T.L. (1897) Coleopterological notices, VII. Annals of the New York Academy of Sciences, 9 (1), 285–684. https://doi.org/10.1111/j.1749-6632.1896.tb55435.x
  6. Castellini, G. (2007) Scidmenidi afrotropicali del Museo di Zoologia dell’Università di Roma “La Sapienza” (Coleoptera, Scydmaenidae). Fragmenta Entomologica, 39 (2), 225–254. https://doi.org/10.4081/fe.2007.121
  7. Castellini, G. (2010) Per la conoscenza degli Scydmaenidae afrotropicali (Coleoptera): descrizioni e note. Atti del Museo di Storia Naturale della Maremma, 22, 3–62. [2008]
  8. Cauchois, P. (1955) LV. Coleoptera Scydmaenidae. Contributions á l’étude de la faune entomologique du Ruanda-Urundi (Mission P. Basilewsky 1953). Annales du Musée Royal du Congo Belge Tervuren, Belgique, Série in 8°, 40, 82–133.
  9. Croissandeau, J. (1891) Description de six nouvelles espèces de Cephennium. Le Coléoptériste, 4, 53–55. [1890–1891]
  10. Croissandeau, J. (1894) Scydmaenidae européens et circa-méditerranéens. Annales de la Société Entomologique de France, 63, 351–400, pls. 6–14.
  11. Csiki, E. (1919) Pars 70: Scydmaenidae. In: Schenkling, S. & Junk, W. (Eds.), Coleopterorum Catalogus. Vol. 7. W. Junk, Berlin, 1–106.
  12. Feng, T. & Yin, Z.-W. (2023) A new Cephennodes Reitter (Coleoptera: Staphylinidae: Scydmaeninae) of C. lustricollis group from East China. Zootaxa, 5346 (3), 331–336. https://doi.org/10.11646/zootaxa.5346.3.7
  13. Hitchcock, E. (1858) Ichnology of New England: A report of the Sandstone of the Connecticut Valley, Especially Its Fossil Footmarks. White, Boston, Massachusetts, 220 pp.
  14. Hoshina, H. (2012) New record of the genus Cephennodes (Coleoptera, Staphylinidae, Scydmaeninae) from Yaeyama Group, the Ryukyus, Japan, with description of a new species. Japanese Journal of Systematic Entomology, 18 (2), 425–429.
  15. Hoshina, H. (2016) Discovery of a second species of the subgenus Fusionodes of the genus Cephennodes (Coleoptera: Staphylinidae: Scydmaeninae) from Honshu, Japan. Bulletin of Research on Regional Environmental Research Education Center, 23, 63–67.
  16. Hoshina, H. (2017) Description of a new second species of the genus Cephennodes (Coleoptera: Staphylinidae: Scydmaeninae) from the Ryukyus, Japan. Memoir of Faculty of Education and Regional Studies, Fukui University, Series II: Natural Science, 24, 27–31.
  17. Hoshina, H. (2019) Seven new species of the subfamily Scydmaeninae (Coleoptera: Staphylinidae) from Japan. The Memoirs of the Research and Education Center for Regional Environment, Fukui University, 25, 49–65. [2018]
  18. Hoshina, H. (2020) Taxonomic note of the genus Cephennodes (Coleoptera: Staphylinidae: Scydmaeninae) from Taiwan. The Memoirs of the Research and Education Center for Regional Environment, University of Fukui, 26 (3), 43–47.
  19. Hoshina, H. (2021) Taxonomic notes of the subfamily Scydmaeninae (Coleoptera: Staphylinidae) from Japan and Taiwan. The Memoirs of the Research and Education Center for Regional Environment, University of Fukui, 28, 81–100.
  20. Hoshina, H. & Kishimoto, T. (2004) First record of the family Scydmaenidae (Coleoptera) from Ogasawara Islands, Japan, with description of a new species. Memoir of Faculty of Education and Regional Studies, Fukui University, Series II: Natural Science, 56, 1–7.
  21. Jakobson, G. (1910) Zhuki Rossii i zapadnoy Evropy. Rukovodstvo k opredieleniyu zhukov. A.J. Devrien, St. Petersburg, 1024 pp., lxxxiii pls.
  22. Jałoszyński, P. (2003) Chelonoidum araiorum sp. nov., an unusual species of the Cephenniini (Coleoptera, Scydmaenidae) from the intertidal zone of the Pacific coast of Central Honshu, Japan. Bulletin of the National Science Museum, Tokyo, Series A, 29 (4), 225–228.
  23. Jałoszyński, P. (2005) Redescription of Cephennodes japonicus (Sharp) (Coleoptera: Scydmaenidae). Genus, Wrocław, 16 (3), 325–329.
  24. Jałoszyński, P. (2006) New genus of the Cephenniini from Borneo with description of Hlavaciellus vampirus n. sp. (Coleoptera: Scydmaenidae). Genus, 17 (1), 67–74.
  25. Jałoszyński, P. (2007a) The Cephenniini of China. II. Cephennodes Reitter of southern provinces, with taxonomic notes on the Cephennodes-Chelonoidum complex (Coleoptera: Scydmaenidae). Genus, Wrocław, 18 (1), 7–101.
  26. Jałoszyński, P. (2007b) The Cephenniini of China. III. Cephennodes Reitter of Sichuan and Shaanxi (Coleoptera: Scydmaenidae). Genus, Wrocław, 18 (2), 151–207.
  27. Jałoszyński, P. (2007c) Validation of Fusionodes Jałoszyński (Coleoptera: Scydmaenidae). Genus, 18 (3), 359.
  28. Jałoszyński, P. (2008a) A new bizarre species of Cephennodes from the Malay Peninsula (Coleoptera: Scydmaenidae). Genus, Wrocław, 19 (1), 27–31.
  29. Jałoszyński, P. (2008b) First record of Cephennodes Reitter from Hong Kong, with description of a new species (Coleoptera: Scydmaenidae). Genus, Wrocław, 19 (2), 171–175.
  30. Jałoszyński, P. (2008c) First record of the occurrence of Eutheiini and Cephenniini in New Guinea, with descriptions of new species of Paraneseuthia Franz and Cephennodes Reitter (Coleoptera: Scydmaenidae). Genus, Wrocław, 19 (1), 37–44.
  31. Jałoszyński, P. (2008d) Taxonomic notes on the Cephenniini (Coleoptera, Scydmaenidae): Status of Coatesia Lea, Cephennomicrus Reitter, and Neseuthia Scott. Zootaxa, 1696 (1), 25–36. https://doi.org/10.11646/zootaxa.1696.1.2
  32. Jałoszyński, P. (2009) Cephenniini of the Philippines. Part 2. Two new species of Cephennodes Reitter from Mindanao (Coleoptera: Staphylinidae). Genus, Wrocław, 20 (3), 391–397.
  33. Jałoszyński, P. (2010) Afrotropical Cephenniini: redescriptions of Cephennodes described by Schaufuss and Cauchois and three new species from Ghana, Nigeria and Uganda (Coleoptera: Staphylinidae: Scydmaeninae). Genus, Wrocław, 21 (2), 175–190.
  34. Jałoszyński, P. (2011a) Hlavaciellus Jałoszyński, 2006: eleven new species, detailed morphology and systematic position within Cephenniini (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 2763 (1), 1–33. https://doi.org/10.11646/zootaxa.2763.1.1
  35. Jałoszyński, P. (2011b) New records and new species of Cephennodes Reitter from Hong Kong (Coleoptera: Staphylinidae: Scydmaeninae). Genus, Wrocław, 22 (1), 1–8. https://doi.org/10.11646/zootaxa.5664.1.1
  36. Jałoszyński, P. (2011c) Taxonomic status of Cephenniini described by Schaufuss from Singapore (Coleoptera: Staphylinidae: Scydmaeninae). Genus, Wrocław, 22 (1), 9–16.
  37. Jałoszyński, P. (2012a) Beetles with “trochantelli”: Phylogeny of Cephenniini (Coleoptera: Staphylinidae: Scydmaeninae) focussing on Neotropical genera. Systematic Entomology, 37, 448–477. https://doi.org/10.1111/j.1365-3113.2012.00627.x
  38. Jałoszyński, P. (2012b) The Cephenniini of China. IV. The first record of Cephennodes Reitter from Hainan (Coleoptera: Staphylinidae: Scydmaeninae). Genus, Wrocław, 23 (1), 33–38.
  39. Jałoszyński, P. (2012c) The Cephenniini of China. V. Cephennodes Reitter of Guizhou and Zhejiang (Coleoptera: Staphylinidae: Scydmaeninae). Genus, Wrocław, 23 (2), 229–248.
  40. Jałoszyński, P. (2013a) Phylogeny of a new supertribe Cephenniitae with generic review of Eutheiini and description of a new tribe Marcepaniini (Coleoptera: Staphylinidae: Scydmaeninae). Systematic Entomology, 39, 159–189. https://doi.org/10.1111/syen.12044
  41. Jałoszyński, P. (2013b) The first record of the tribe Cephenniini in South Africa, with new species of Cephennomicrus Reitter and Cephennodes Reitter (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 3683 (4), 411–426. https://doi.org/10.11646/zootaxa.3683.4.5
  42. Jałoszyński, P. (2015a) New nomenclatural and taxonomic acts, and comments: Staphylinidae: Scydmaeninae. In: Löbl, I. & Löbl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Revised and updated edition. Hydrophiloidea - Staphylinoidea. Vol. 2/1. Brill, Leiden and Boston, pp. 22–23. https://doi.org/10.1163/9789004296855_002
  43. Jałoszyński, P. (2015b) Staphylinidae: Scydmaeninae. In: Löbl, I. & Löbl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2/1. Hydrophiloidea - Staphylinoidea. Revised and updated edition. Brill, Leiden and Boston, p. 22.
  44. Jałoszyński, P. (2015c) The Cephenniini of China. VI. New species and new records of Cephennodes Reitter from Hainan, Guangxi and Guangdong (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 3990 (2), 221–234. https://doi.org/10.11646/zootaxa.3990.2.3
  45. Jałoszyński, P. (2015d) The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4033 (3), 393–410. https://doi.org/10.11646/zootaxa.4033.3.5
  46. Jałoszyński, P. (2016a) Four new species of Pomphopsilla Jałoszyński and a new record of Cephennodes glabella Castellini in the Democratic Republic of the Congo (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4098 (3), 593–600. https://doi.org/10.11646/zootaxa.4098.3.11
  47. Jałoszyński, P. (2016b) The Cephenniini of China. VIII. New species and new records of Cephennodes Reitter of Hunan, Jiangxi, Zhejian and Fujian (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4079 (4), 415–428. https://doi.org/10.11646/zootaxa.4079.4.2
  48. Jałoszyński, P. (2017a) Ant-like stone beetles on the roof of the world. Cephenniini of Nepal and Bhutan (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4349 (1), 1–120. https://doi.org/10.11646/zootaxa.4349.1.1
  49. Jałoszyński, P. (2017b) First record of Cephenniitae in the Bismarck Archipelago, Papua New Guinea, with description of new species of Cephennodes and Cephennomicrus of Mussau and Manus Islands (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4268 (3), 427–432. https://doi.org/10.11646/zootaxa.4268.3.7
  50. Jałoszyński, P. (2018) †Praphennium gen. nov.: extant mite killers meet their Cenomanian relatives (Coleoptera: Staphylinidae: Scydmaeninae: Cephenniini). Cretaceous Research, 90, 142–153. https://doi.org/10.1016/j.cretres.2018.04.011
  51. Jałoszyński, P. (2021) New taxonomic and nomenclatural acts in Cephenniini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4996 (1), 171–176. https://doi.org/10.11646/zootaxa.4996.1.8
  52. Jałoszyński, P. (2022) Euconnus Thomson of Japan: redescriptions of species established by Reitter, Sharp and Franz, new synonyms, and summary of current state of knowledge (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 5093 (1), 1–37. https://doi.org/10.11646/zootaxa.5093.1.1
  53. Jałoszyński, P. (2023) New species and new records of Eastern Palaearctic Stenichnus Thomson (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 5318 (3), 363–381. https://doi.org/10.11646/zootaxa.5318.3.3
  54. Jałoszyński, P. (2025a) Introduction to Scydmaeninae (Coleoptera, Staphylinidae): morphology of adults. Zootaxa, 5581 (1), 1–390. https://doi.org/10.11646/zootaxa.5581.1.1
  55. Jałoszyński, P. (2025b) John Nietner collection of Sri Lankan Scydmaeninae (Coleoptera, Staphylinidae). Zootaxa, 5633 (1), 79–109. https://doi.org/10.11646/zootaxa.5633.1.4
  56. Jałoszyński, P. (2025c) The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 5664 (1), 1–101. https://doi.org/10.11646/zootaxa.5664.1.1
  57. Jałoszyński, P. & Hoshina, H. (2003) Notes on the distribution of Cephennodes vafer Kurbatov, 1995 (Coleoptera, Scydmaenidae) in the mainland of Japan. Elytra, Tokyo, 31 (1), 195–198.
  58. Jałoszyński, P. & Nomura, S. (2008) Cephenniini (Coleoptera: Scydmaenidae) of Doi Inthanon and Khao Yai National Parks, Thailand. Entomological Reviews of Japan, 63 (2), 71–88.
  59. Jałoszyński, P. & Nomura, S. (2009) The Cephenniini (Coleoptera, Scydmaenidae) of Vietnam. Bulletin of the National Museum of Nature and Science, Series A, 35 (3), 167–225.
  60. Jałoszyński, P., Nomura, S. & Kurbatov, S. (2005) New records of Chelonoidum graeseri (Reitter) in Russia and South Korea (Coleoptera, Scydmaenidae). Elytra, Tokyo, 33 (2), 543–546.
  61. Jeon, G.N. & Ahn, K.-J. (2023) Two intertidal Staphylinidae species (Coleoptera) new to Korea. Journal of Asia-Pacific Biodiversity, 16 (4), 675–677. https://doi.org/10.1016/j.japb.2023.01.003
  62. Kurbatov, S. (1995) Sur les Euthiini et Cephenniini (Coleoptera, Scydmaenidae) de l’extrême-est de la Russie et du Japon. Revue suisse de Zoologie, 102 (4), 943–959. https://doi.org/10.5962/bhl.part.80485
  63. Lazorko, W. (1962) Zwei neue Cephennium-Arten (Col. Scydmaenidae) mit einer Übersicht der ukrainischen Arten der Tribus Cephenniini. Entomologische Arbeiten aus dem Museum Georg Frey, 13, 273–320.
  64. Lea, A.M. (1915) Notes on Australian and Tasmanian Scydmaenidae, with descriptions of new species. Proceedings of the Royal Society of Victoria, Melbourne, New Series, 27 (2), 198–231. [1914]
  65. LeConte, J.L. (1852) Synopsis of the Scydmaenidae of the United States. Proceedings of the Academy of Natural Scences, Philiadelphia, 6 (5), 149–157.
  66. Makhan, D. (2005) Four new Scydmaenidae species from Mount Jinyun, China (Coleoptera). Calodema, 3, 6–13.
  67. Makhan, D. (2009) Hawkeswoodcephennodes gen. nov. from China (Coleoptera: Scydmaenidae) and a response to the criticisms of Pawel Jaloszynski (2007). Calodema, 108, 1–7.
  68. Marsh, G.A. (1957) Family Scydmaenidae. In: Hatch, M.H. (Ed.), The Beetles of the Pacific Northwest. Part II. Staphyliniformia. University of Washington Publications in Biology. University of Washington Press, Seattle, Washington, pp. 271–280.
  69. Motschulsky, V. (1851) Lettres adressées à Monsieur le premier secrétaire de la Société Impériale des Naturalistes de Moscou. Bulletin de la Société Impériale des Naturalistes de Moscou, 24 (4), 592–616.
  70. Motschulsky, V. (1858) Entomologie spéciale. Insectes des Indes orientales. Études Entomologiques, 7, 20–122, 1 pl.
  71. Müller, P.W.I. & Kunze, G. (1822) Monographie der Ameisenkäfer (Scydmaenus Latreille). Schriften der Naturforschenden Gesellschaft zu Leipzig, 1, 175–204, pl. 5.
  72. Na, D., Byeon, U.-J. & Park, J.-S. (2024) Review of the tribe Cephenniini Reitter (Coleoptera: Staphylinidae: Scydmaeninae) in Korea. Journal of insect biodiversity, 58 (1), 15–27. https://doi.org/10.12976/jib/2024.58.1.3
  73. Newton, A.F. (2017) Nomenclatural and taxonomic changes in Staphyliniformia (Coleoptera). Insecta Mundi, 0595, 1–52.
  74. Nietner, J. (1856) Entomological papers - being descriptions of new Ceylon Coleoptera with such observations on their habits as appear any way interesting. Journal of the Asiatic Society of Bengal, 25 (5–6), 381–394, 523–554.
  75. O’Keefe, S.T. (1998) Notes on the classification of North American ant-like stone beetles (Coleoptera: Scydmaenidae). The Coleopterists Bulletin, 52, 259–269.
  76. Orousset, J. (2024) Les types de Scydmaeninae décrits par Maurice Pic (Coleoptera, Staphylinidae). Bulletin de la Société entomologique de France, 129 (3), 331–348. https://doi.org/10.32475/bsef_2353
  77. Park, S.-J., Hoshina, H. & Ahn, K.-J. (2006) Taxonomy of the tribe Cephenniini Reitter (Coleoptera: Scydmaenidae) in Korea. Entomological Research, 36 (2), 107–111. https://doi.org/10.1111/j.1748-5967.2006.00017.x
  78. Pic, M. (1901) Notes diverses et diagnoses. Échange, Revue Linnéenne, 17 (193), 2–4.
  79. Reitter, E. (1877) Neue caucasische Coleopteren, gesammelt von Hans Leder (Zweites Stück). Deutsche Entomologische Zeitschrift, 21 (2), 289–296. https://doi.org/10.1002/mmnd.4800210213
  80. Reitter, E. (1884) Beitrag zur Pselaphiden- und Scydmaeniden-Fauna von Java und Borneo. II. Stück. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 33, 387–428, pl. 20. [1883] https://doi.org/10.1002/mmnd.48018830104
  81. Reitter, E. (1887) Neue Coleopteren aus Europa, den angrenzenden Ländern und Sibirien, mit Bemerkungen über bekannte Arten. Dritter Theil. Deutsche Entomologische Zeitschrift, 31 (1), 241–288. https://doi.org/10.1002/mmnd.48018870141
  82. Reitter, E. (1908) Siebzehn Coleopteren-Neuheiten aus der palaearktischen Fauna. Wiener Entomologische Zeitung, 27 (1), 17–24. https://doi.org/10.5962/bhl.part.11639
  83. Roubal, J. (1913) Verschiedene koleopterologiscbe Notizen: 17-21. Coleopterologische Rundschau, 2 (11), 185.
  84. Schaufuss, L.W. (1884) Die Scydmaeniden nord-ost-Afrika’s, der Sunda-Inseln und Neu-Guinea’s im Museo Civico di Storia Naturale zu Genua. Annali del Museo Civico di Storia Naturale di Genova, 2a (1), 387–424.
  85. Schaufuss, L.W. (1888) Über Pselaphiden und Scydmaeniden des Königl. zoologischen Museums zu Berlin und verwandte Arten. Berliner Entomologische Zeitschrift, 31 (2), 287–320. [1887] https://doi.org/10.1002/mmnd.18880310216
  86. Schaufuss, L.W. (1889) Neue Scydmaeniden im Museum Ludwig Salvator. Berliner entomologische Zeitschrift, 33 (1), 1–42. https://doi.org/10.1002/mmnd.18890330103
  87. Sharp, D. (1886) The Scydmaenidae of Japan. Entomologist’s Monthly Magazine, 23, 46–51.
  88. Stevanović, M. (2009) A new species of the genus Cephennodes Reitter, 1883 from Thailand (Coleoptera, Staphylinidae, Scydmaeninae). Natura Croatica, 18 (2), 263–266.
  89. Strand, E. (1935) Revision von Gattungsnamen paläarktischer Coleoptera. Folia Zoologica et Hydrobiologica, 7, 282–299.
  90. Thomson, C.G. (1859) Skandinaviens Coleoptera, synoptiskt bearbetade. Vol. 1. Berlingska Boktryckeriet, Lund, 290 pp. https://doi.org/10.5962/bhl.title.138677
  91. Yin, Z.-W. & Zhou, D.-Y. (2024) Improved diagnosis for Cephennodes pseudobos Jałoszyński (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 5528 (1), 811–813. https://doi.org/10.11646/zootaxa.5528.1.54
  92. Zimmerman, E.C. (1942) A new scydmaenid from Hawaii (Coleoptera). Proceedings of the Hawaiian Entomological Society, 11, 238–241.

How to Cite

Jałoszyński, P. (2026) Revision of subgeneric system and species groups of Cephennodes Reitter, with redescriptions of problematic species (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 5766 (1), 1–87. https://doi.org/10.11646/zootaxa.5766.1.1