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Issue: Vol. 5792 No. 2: 13 Apr. 2026
Type: Article
Published: 2026-04-13
DOI: 10.11646/zootaxa.5792.2.1
Page range: 201-244
Abstract views: 95
PDF downloaded: 9

Catalogue of type specimens of amphibians and reptiles in the Estación Biológica de Doñana (EBD-CSIC), Spain

Estación Biológica de Doñana (EBD-CSIC); Americo Vespucio 26; 41092 Seville; Spain
Scientific Collections Services–ICTS-RBD; Estación Biológica de Doñana (EBD-CSIC); Americo Vespucio 26; 41092 Seville; Spain
Scientific Collections Services–ICTS-RBD; Estación Biológica de Doñana (EBD-CSIC); Americo Vespucio 26; 41092 Seville; Spain
Scientific Collections Services–ICTS-RBD; Estación Biológica de Doñana (EBD-CSIC); Americo Vespucio 26; 41092 Seville; Spain; Conservation and Evolutionary Genetics Group; Estación Biológica de Doñana (EBD-CSIC); Americo Vespucio 26; 41092 Seville; Spain
Amphibia Reptilia type specimens natural history collections Doñana Biological Station Seville

Abstract

We provide a first complete list of the amphibian and reptile type specimens preserved in the scientific collections of the Estación Biológica de Doñana (EBD-CSIC) in Spain (updated to 15 September 2025). The herpetological collection houses 195 type specimens (17 holotypes, 164 paratypes, and 14 syntypes), of 33 taxa (20 species and 13 subspecies). Twenty-three of these taxa are currently considered valid. Most of the anuran type specimens are from Bolivia and Venezuela, while reptile types are mainly from Spain. Photographs of all holotypes and also for some paratypes and syntypes are provided.

 

References

  1. Araujo-Vieira, K., Lourenço, A.C.C., Lacerda, J.V.A., Lyra, M.L., Blotto, B.L., Ron, S.R., Baldo, D., Pereyra, M.O., Suárez-Mayorga, Á.M., Baêta, D., Ferreira, R.B., Barrio-Amorós, C.L., Borteiro, C., Brandão, R.A., Brasileiro, C.A., Donnelly, M.A., Dubeux, M.J.M., Köhler, J., Kolenc, F., Leite, F.S.F., Maciel, N.M., Nunes, I., Orrico, V.G.D., Peloso, P., Pezzuti, T.L., Reichle, S., Rojas-Runjaic, F.J.M., Da Silva, H.R., Sturaro, M.J., Langone, J.A., Garcia, P.C.A., Rodrigues, M.T., Frost, D.R., Wheeler, W.C., Grant, T., Pombal, J.P., Jr., Haddad, C.F.B. & Faivovich, J. (2023) Treefrog Diversity in the Neotropics: Phylogenetic Relationships of Scinaxini (Anura: Hylidae: Hylinae). South American Journal of Herpetology, 27 (sp1), 1–143. https://doi.org/10.2994/SAJH-D-22-00038.1
  2. Arribas, O.J. (1996) Taxonomic revision of the Iberian ‘Archaeolacertae” I: A new interpretation of the geographical variation of ‘Lacerta’ monticola Boulenger 1905 and ‘Lacerta’ cyreni Müller & Hellmich 1937 (Squamata: Sauria: Lacertidae). Herpetozoa, 9, 31–56.
  3. Arribas, O.J. (2017) Oteroi/oteroorum and raffonei/raffoneae: the difference between grammatical correctness and validity in Zoological Nomenclature. Butlletí de la Societat Catalana d’Herpetologia, 24, 29–32.
  4. Ayarzagüena, J. (1992) Los centrolénidos de la Guayana Venezolana. Publicaciones de la Asociación de Amigos de Doñana, 1, 1–46.
  5. Ayarzagüena, J. & Señaris J. C. (1994 “1993”) Dos nuevas especies de Hyla (Anura; Hylidae) para las cumbres tepuyanas del Estado Amazonas, Venezuela. Memoria de la Sociedad de Ciencias Naturales La Salle, 53, 127–146.
  6. Ayres, C. (2015) Galápago europeo–Emys orbicularis (Linnaeus, 1758). In: Salvador A. & Marco, A. (Eds.), Enciclopedia Virtual de los Vertebrados Españoles. Museo Nacional de Ciencias Naturales, Madrid. Available from: http://www.vertebradosibericos.org/ (accessed 21 January 2026)
  7. Barrio-Amorós, C.L. (1999) Sistemática y biogeografía de los anfibios (Amphibia) de Venezuela. Acta Biológica Venezuelica, 18 (2), 1–93.
  8. Bowles, P. (2024a) Algyroides marchi. The IUCN Red List of Threatened Species, 2024, e.T840A137842545. https://doi.org/10.2305/IUCN.UK.2024-1.RLTS.T840A137842545.en
  9. Bowles, P. (2024b) Gallotia atlantica. The IUCN Red List of Threatened Species, 2024, e.T61500A137850299. https://doi.org/10.2305/IUCN.UK.2024-1.RLTS.T61500A137850299.en
  10. Bowles, P. (2024c) Iberolacerta cyreni. The IUCN Red List of Threatened Species, 2024, e.T61514A137851751. https://doi.org/10.2305/IUCN.UK.2024-1.RLTS.T61514A137851751.en
  11. Bowles, P. (2024d) Timon lepidus. The IUCN Red List of Threatened Species, 2024, e.T218293375A137858480. https://doi.org/10.2305/IUCN.UK.2024-1.RLTS.T218293375A137858480.en
  12. Bowles, P. (2024e) Podarcis lusitanicus. The IUCN Red List of Threatened Species, 2024, e.T212423579A212423660. https://doi.org/10.2305/IUCN.UK.2024-1.RLTS.T212423579A212423660.en
  13. Bowles, P. (2024f) Chalcides bedriagai. The IUCN Red List of Threatened Species, 2024, e.T61472A137847259. https://doi.org/10.2305/IUCN.UK.2024-1.RLTS.T61472A137847259.en
  14. Bowles, P. (2024g) Chalcides coeruleopunctatus. The IUCN Red List of Threatened Species, 2024, e.T200056417A201258375. https://doi.org/10.2305/IUCN.UK.2024-1.RLTS.T200056417A201258375.en
  15. Cabot, J. (1991) Prólogo. In: Cabot, J. (Coord.), Inventario de las colecciones herpetológica y ornitológica de la Estación Biológica de Doñana. Estación Biológica de Doñana-Consejo Superior de Investigaciones Científicas, Sevilla, pp. 2-3.
  16. Caeiro-Dias, G., Rocha, S., Couto, A., Pereira, C., Brelsford, A., Crochet, P.-A. & Pinho, C. (2021) Nuclear Phylogenies and Genomics of a Contact Zone Establish the Species Rank of Podarcis lusitanicus (Squamata, Lacertidae). Molecular Phylogenetics and Evolution, 164, 107270. https://doi.org/10.1016/j.ympev.2021.107270
  17. Castroviejo, J. & Mateo, J.A. (1998) Una nueva subespecie de Lacerta lepida Daudin 1802 (Sauria, Lacertidae) para la Isla de Sálvora (España). Publicaciones de la Asociación de Amigos de Doñana, 12, 1–21.
  18. Cezón, K., Castilla, F. & Pando, F. (2021). Informe de colecciones biológicas y bases de datos de biodiversidad en España. 6ª Edition. Nodo Nacional de Información en Biodiversidad GBIF-CSIC. Consejo Superior de Investigaciones Científicas (CSIC), Madrid, 59 pp. https://doi.org/10.20350/digitalCSIC/14140
  19. Crochet, P.-A. & Dubois, A. (2004) Recent changes in the taxonomy of European amphibians and reptiles. In: Gasc, J.P., Cabela, A., Crnobrnja-Isailovic, J., Dolmen, D., Grossenbacher, K., Haffner, P., Lescure, J., Martens, H., Martínez Rica, J.P., Maurin, H., Oliveira, M.E., Sofianidou, T.S., Veith, M. & Zuiderwijk, A. (Eds.), Atlas of the amphibians and reptiles in Europe (second edition). Societas Europaea Herpetologica & Muséum National d’Histoire Naturelle, Paris, pp. 495–516.
  20. Cuadrado, M., Amarillo, J.M. & Urdiales, C. (2019) La colección de vertebrados de la Estación Biológica de Doñana (CSIC, Sevilla). Especial referencia a los ejemplares depositados por el ZooBotánico de Jeréz. La singular historia de un torillo andaluz (Turnix sylvatica sylvatica). El Corzo Sociedad Gaditana de Historia Natural, 7, 19–31.
  21. De la Riva, I. & Mateo, J.A. (1991) Capítulo I. Inventario de la Colección Herpetológica de Doñana. In: Cabot, J. (Coord.), Inventario de las colecciones herpetologica y ornitológica de la Estación Biologica de Doñana (C.S.I.C). Estación Biológica de Doñana, Sevilla, pp. 9–17.
  22. De la Riva, I. (1990 “1989”) Una especie nueva de Ololygon (Anura: Hylidae) procedente de Bolivia. Revista Española de Herpetología, 4, 81–86.
  23. De la Riva, I. (1992a) A new species of Phrynopus from Bolivia (Anura: Leptodactylidae). Herpetologica, 48 (1), 11–114.
  24. De la Riva, I. (1992b) Comentarios sobre el género Gastrotheca (Anura: Hylidae) en Bolivia y descripción de una nueva especie. Revista Española de Herpetología, 6, 15–22.
  25. De la Riva, I. (1993) A new species of Scinax (Anura, Hylidae) from Argentina and Bolivia. Journal of Herpetology, 27 (1), 41–46.
  26. De la Riva, I. (1994) A new aquatic frog of the genus Telmatobius (Anura: Leptodactylidae) from Bolivian cloud forests. Herpetologica, 50 (1), 38–45.
  27. De la Riva, I., Chaparro, J.C., Castroviejo-Fisher, S. & Padial, J.M. (2017 “2018”) Underestimated anuran radiations in the high Andes: five new species and a new genus of Holoadeninae, and their phylogenetic relationships (Anura: Craugastoridae). Zoological Journal of the Linnean Society, 182 (1), 129–172. https://doi.org/10.1093/zoolinnean/zlx020
  28. Díaz-Rodríguez, J., Gehara, M., Márquez, R., Vences, M., Gonçalves, H., Sequeira, F., Martínez-Solano, I. & Tejedo, M. (2017) Integration of molecular, bioacoustical and morphological data reveals two new cryptic species of Pelodytes (Anura, Pelodytidae) from the Iberian Peninsula. Zootaxa, 4243 (1), 1–41. https://doi.org/10.11646/zootaxa.4243.1.1
  29. Dubois, A. (2007) Genitives of species and subspecies nomina derived from personal names should not be emended. Zootaxa, 1550 (1), 49–68. https://doi.org/10.11646/zootaxa.1550.1.2
  30. Duellman, W.E. & Señaris, J.C. (2003) A new species of glass frog (Anura: Centrolenidae) from the Venezuelan Guayana. Herpetologica, 59 (2), 247–252.
  31. Duellman, W.E. & Wiens, J.J. (1992) The status of the hylid frog genus Ololygon and the recognition of Scinax Wagler, 1830. Occasional Papers of the Museum of Natural History, University of Kansas, 151, 1–23.
  32. Dufresnes, C., Pribille, M., Alard, B., Gonçalves, H., Amat, F., Crochet, P.A., Dubey, S., Perrin, N., Fumagalli, L., Vences, M. & Martínez-Solano, I. (2020) Integrating hybrid zone analyses in species delimitation: lessons from two anuran radiations of the Western Mediterranean. Heredity, 124, 423–438. https://doi.org/10.1038/s41437-020-0294-z
  33. Faivovich, J., Haddad, C.F.B., Garcia, P.C.O., Frost, D.R., Campbell, J.A. & Wheeler, W.C. (2005) Systematic review of the frog family Hylidae, with special reference to Hylinae: Phylogenetic analysis and taxonomic revision. Bulletin of the American Museum of Natural History, 294, 1–240. https://doi.org/10.1206/0003-0090(2005)294[0001:SROTFF]2.0.CO;2
  34. Faivovich, J., McDiarmid, R.W. & Myers C.W. (2013) Two new species of Myersiohyla (Anura: Hylidae) from Cerro de la Neblina, Venezuela, with comments on other species of the genus. American Museum Novitates, 3792, 1–63. https://doi.org/10.1206/3792.1
  35. Fritz, U. (2002) Herpetology and herpetological type specimens at the Museum für Tierkunde Dresden with a bibliography of herpetological contributions by Fritz Jürgen Obst (Amphibia, Reptilia). Faunistiche Abhhandlungen Staatliches Museum für Tierkunde Dresden, 23 (1), 3–34.
  36. Fritz, U., Keller, C. & Budde, M. (1996) Eine neue Unterart der Europäischen Sumpfschildkröte aus Südwestspanien, Emys orbicularis hispanica subsp. nov. Salamandra, 32 (3),129–152.
  37. Frost, D.R. (2024) Amphibian Species of the World: an Online Reference. Version 6.2. American Museum of Natural History, New York, New York, Electronic Database. Available from: https://amphibiansoftheworld.amnh.org/index.php (accessed 28 July 2025)
  38. García-Díez, T. & González–Fernández, J.E. (2013) The reptile type specimens preserved in the Museo Nacional de Ciencias Naturales (CSIC) of Madrid, Spain. Zootaxa, 3619 (1), 046–058. https://doi.org/10.11646/zootaxa.3619.1.2
  39. García-Díez, T., Sempere, M.R., Cabot, J., Juste, J. & Ibáñez, C. (2014) The collections of vertebrates of the Estación Biológica de Doñana (CSIC). Origin and evolution. Journal of Paleontological Techniques, 13, 111–117.
  40. Geniez, P., Mateo Miras, J.A., Joger, U., Pleguezuelos, J., Slimani, T. & El Mouden, E.H. (2010) Acanthodactylus busacki. The IUCN Red List of Threatened Species, 2010, e.T178569A7572735. https://doi.org/10.2305/IUCN.UK.2010-4.RLTS.T178569A7572735.en
  41. Geniez, P., Sá-Sousa, P., Guillaume, C.P., Cluchier, A. & Crochet, P.A. (2014) Systematics of the Podarcis hispanicus complex (Sauria, Lacertidae) III: valid nomina of the western and central Iberian forms. Zootaxa, 3794 (1), 1–51. https://doi.org/10.11646/zootaxa.3794.1.1
  42. González, P.F.P., Nogales, M., Jiménez, A.J. & Hernández, M. (1996) Phylogenetic relationships of the Canary Islands endemic lizard genus Gallotia (Sauria: Lacertidae), inferred from mitochondrial DNA sequences. Molecular Phylogenetics and Evolution, 6, 63–71. https://doi.org/10.1006/mpev.1996.0058
  43. González-Fernández J.E., Calvo M. & Rey, I. (2017) Ejemplares tipo de anfibios y reptiles conservados en las colecciones científicas del Museo Nacional de Ciencias Naturales (MNCN–CSIC) de Madrid: Revisión bibliográfica y catálogo. Boletín de la Real Sociedad Española de Historia Natural (Sección aula, museos y colecciones), 4, 107–127.
  44. Guayasamin, J.M., Castroviejo-Fisher, S., Trueb, L., Ayarzagüena, J., Rada, M. & Vilà, C. (2009) Phylogenetic systematics of Glassfrogs (Amphibia: Centrolenidae) and their sister taxon Allophryne ruthveni. Zootaxa, 2100 (1), 1–97. https://doi.org/10.11646/zootaxa.2100.1.1
  45. Hedges, S.B., Duellman, W.E. & Heinicke, M.P. (2008) New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa, 1737 (1), 1–182. https://doi.org/10.11646/zootaxa.1737.1.1
  46. ICZN/International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. Available from: https://www.iczn.org/the-code/the-code-online/ (accessed 28 July 2025)
  47. IUCN/The International Union for Conservation of Nature] (2025) The IUCN Red List of Threatened Species. Version 2025-1. Available from: https://www.iucnredlist.org (accessed 20 July 2025)
  48. IUCN SSC Amphibian Specialist Group (2020a) Vitreorana gorzulae. The IUCN Red List of Threatened Species, 2020, e.T89098052A109546415. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T89098052A109546415.en
  49. IUCN SSC Amphibian Specialist Group (2020b) Vitreorana helenae. The IUCN Red List of Threatened Species, 2020, e.T55017A109531839. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T55017A109531839.en
  50. IUCN SSC Amphibian Specialist Group (2020c) Microkayla kempffi. The IUCN Red List of Threatened Species, 2020, e.T57215A154333734. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T57215A154333734.en
  51. IUCN SSC Amphibian Specialist Group (2020d) Scinax castroviejoi. The IUCN Red List of Threatened Species, 2020, e.T55943A98407339. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T55943A98407339.en
  52. IUCN SSC Amphibian Specialist Group (2020e) Telmatobius yuracare. The IUCN Red List of Threatened Species, 2020, e.T57369A154335458. https://doi.org/10.2305/IUCN.UK.2020-3.RLTS.T57369A154335458.en
  53. IUCN SSC Amphibian Specialist Group (2022a) Hyalinobatrachium iaspidiense. The IUCN Red List of Threatened Species, 2022, e.T201029146A89198728. https://doi.org/10.2305/IUCN.UK.2022-2.RLTS.T201029146A89198728.en
  54. IUCN SSC Amphibian Specialist Group (2022b) Pelodytes punctatus (amended version of 2020 assessment). The IUCN Red List of Threatened Species, 2022, e.T173111497A218083218. https://doi.org/10.2305/IUCN.UK.2022-2.RLTS.T173111497A218083218.en
  55. IUCN SSC Amphibian Specialist Group (2023a) Gastrotheca lauzuricae. The IUCN Red List of Threatened Species, 2023, e.T55341A193846651. https://doi.org/10.2305/IUCN.UK.2023-1.RLTS.T55341A193846651.en
  56. IUCN SSC Amphibian Specialist Group (2023b) Scinax chiquitanus. The IUCN Red List of Threatened Species, 2023, e.T55946A61407114. https://doi.org/10.2305/IUCN.UK.2023-1.RLTS.T55946A61407114.en
  57. Köhler, J., Koch, C., Garlacz, R., Preick, M., De la Riva, I. & Vences, M. (2024) Getting back to name-bearing types: archival DNA and morphology clarify the identity of Hyla splendens Schmidt, 1857 and challenge the taxonomy of Bolivian populations of Gastrotheca (Anura: Hemiphractidae). Salamandra, 60 (4), 263–282.
  58. Kok, P. & Rivas Fuenmayor, G. (2008) Typhlophis ayarzaguenai Señaris, 1998 is a junior synonym of Typhlophis squamosus (Schlegel, 1839). Amphibia-Reptilia, 29 (4), 555–558. https://doi.org/10.1163/156853808786230406
  59. Lansac, C., Aguayo, R. & De la Riva, I. (2021) An assessment of the taxonomic validity of three species of marsupial frogs (Anura: Hemiphractidae: Gastrotheca) from the Yungas of Bolivia based on external morphology and cranial osteology. Zootaxa, 5068 (2), 211–239. https://doi.org/10.11646/zootaxa.5068.2.3
  60. Luiselli, L. & Vamberger, M. (2024) Emys orbicularis (Europe assessment). The IUCN Red List of Threatened Species, 2024, e.T7717A207667247. https://doi.org/10.2305/IUCN.UK.2024-1.RLTS.T7717A207667247.en
  61. Martínez, G., León, R., Jiménez-Robles, O., González De la Vega, J.P., Gabari, V., Rebollo, B., Sánchez-Tójar, A., Fernández-Cardenete, J.R. & Gállego, J. (Eds.) (2025) Moroccoherps.com 2.0. Amphibians and Reptiles of Morocco and Western Sahara. Available from: https://www.moroccoherps.com/ (accessed 20 July 2025)
  62. Martínez-Freiría, F., Freitas, I., Velo-Antón, G., Lucchini, N., Fahd, S., Larbes, S., Pleguezuelos, J.M., Santos, X. & Brito, J.C. (2021). Integrative taxonomy reveals two species and intraspecific differentiation in the Vipera latastei–monticola complex. Journal of Zoological Systematics and Evolutionary Research, 59, 2278–2306. https://doi.org/10.1111/jzs.12534
  63. Martínez-Freiría, F., Santos, X. & Pleguezuelos, J.M. (2024) Vipera latastei. The IUCN Red List of Threatened Species, 2024, e.T221202039A137859691. https://doi.org/10.2305/IUCN.UK.2024-1.RLTS.T221202039A137859691.en
  64. Martínez-Solano, I. (2024) Pelodytes atlanticus. The IUCN Red List of Threatened Species, 2024, e.T114139495A227223836. https://doi.org/10.2305/IUCN.UK.2024-2.RLTS.T114139495A227223836.en
  65. Mateo, J.A. (2015) El lagarto atlántico (Gallotia atlantica) en Gran Canaria. Boletín de la Asociación Herpetológica Española, 26 (2), 61–63.
  66. Mateo, J.A., Geniez, P., López-Jurado, L.F. & Bons, J. (1999) Chorological analysis and morphological variations of Saurians of the genus Uromastyx (Reptilia: Agamidae) in western Sahara. Description of two new taxa. Revista Española de Herpetologia, 12, 97–109.
  67. Mateo J.A., Geniez, P., Veiret, P. & López-Jurado, L.F. (2023) Reptiles de Macaronesia, Azores, Madeira, Salvajes, Canarias, Cabo Verde y Litoral Sáhara Atlantico. Asociación Herpetológica Española, Madrid, 473 pp.
  68. Michels, J.P. & Bauer, A.M. (2004) Some corrections to the scientific names of amphibians and reptiles. Bonner Zoologische Beiträge, 52, 83–94.
  69. Myers, C.W. & Donnelly, M.A. (1997) A tepui herpetofauna on a granitic mountain (Tamacuari) in the borderland between Venezuela and Brazil: report from the Phipps Tapirapecó Expedition. American Museum Novitates, 3213, 1–71.
  70. Oubotar, P., Schargel, W. & Rivas, G. (2019) Typhlophis squamosus. The IUCN Red List of Threatened Species, 2019, e.T203205A2762090. https://doi.org/10.2305/IUCN.UK.2019-3.RLTS.T203205A2762090.en
  71. Padial, J.M., Castroviejo-Fisher, S., Merchán, M., Cabot, J. & Castroviejo, J. (2003) The herpetological collection from Bolivia in the Estación Biológica de Doñana (Spain). Graellsia, 59 (1), 5–14.
  72. Pleguezuelos, J.M., Marques, R. & Lizana, M. (Eds.) (2002) Atlas y Libro Rojo de los anfibios y reptiles de España. Dirección General de Conservación de la Naturaleza-Asociación Herpetológica Española (2ª impresión), Madrid, 587 pp.
  73. Pollo, C. (2009). Eslizón ibérico–Chalcides bedriagai. In: Salvador, A. & Marco, A. (Eds.), Enciclopedia Virtual de los Vertebrados Españoles. Museo Nacional de Ciencias Naturales, Madrid. Available from: http://www.vertebradosibericos.org/ (accesed 21 January 2026)
  74. Ruiz-Carranza, P.M. & Lynch, J.D. (1991) Ranas Centrolenidae de Colombia I. Propuesta de una nueva clasificación genérica. Lozania, 57, 1–30.
  75. Sabaj, M.H. (2020) Codes for Natural History Collections in Ichthyology and Herpetology. Copeia, 108 (3), 593–669. https://doi.org/10.1643/ASIHCODONS2020
  76. Sabaj, M.H. (2025) Codes for Natural History Collections in Ichthyology and Herpetology (online supplement). Version 9.8 American Society of Ichthyologists and Herpetologists, Washington, DC. Available from: https://www.asih.org/resources/standard-symbolic-codes (accessed 29 July 2025)
  77. Salvador, A. (1982) A revision of the lizards of the genus Acanthodactylus (Sauria: Lacertidae). Bonner Zoologische Monographien, 16, 1–167.
  78. Salvador, A. (1998). Chalcides bedriagai (Boscá, 1880). In: Salvador, A. (Coord.), Fauna Ibérica. Vol. 10. Museo Nacional de Ciencias Naturales-CSIC, Madrid, pp. 149–156.
  79. Salvador, A., Díaz-Paniagua, C., Recuero Gil, E., Martinez Solano, I., Bosch, J., García París, M., Lizana, M., Tejedo, M., Beja, P., Hernández Sastre, P., Márquez, R., Reques, R. & Pérez, V. (2024) Pelodytes ibericus. The IUCN Red List of Threatened Species, 2024, e.T58055A227224480. https://doi.org/10.2305/IUCN.UK.2024-2.RLTS.T58055A227224480.en
  80. Sánchez-Herráiz, M.J., Barbadillo-Escrivá, L. J., Machordom, A. & Sanchíz, B. (2000) A new species of pelodytid frog from the Iberian Peninsula. Herpetologica 56 (1), 105–118.
  81. Sánchez-Vialas, A., Calvo-Revuelta, M. & De la Riva, I. (2022) Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa, 5202 (1), 1–197. https://doi.org/10.11646/zootaxa.5202.1.1
  82. Sánchez-Vialas, A., Calvo-Revuelta, M., Castroviejo-Fisher, S. & De la Riva, I. (2020) Synopsis of the Amphibians of Equatorial Guinea based upon the Authors’ Field Work and Spanish Natural History Collections. Proceedings of the California Academy of Sciences, 66 (8), 137–230.
  83. Sánchez-Vialas, A., Calvo-Revuelta, M., Rubio, J. L., Palacios, F. & García-París, M. (2019). Taxonomic and nomenclatural status of Iberian Algyroides (Lacertidae). Amphibia-Reptilia, 40 (1), 91–102. https://doi.org/10.1163/15685381-20181016
  84. Señaris, C., Urdiales, C., Sempere M. & Leonard, J.A. (2023) Colección herpetológica de Latinoamérica en la Estación Biológica de Doñana. Memorias del XII Congreso Latinoamericano de Herpetología, 530–531.
  85. Señaris, J.C. & Ayarzagüena, J. (1994 “1993”) Una nueva especie de Centrolenella (Anura: Centrolenidae) del Auyán-tepui, Edo. Bolívar, Venezuela. Memoria de la Sociedad de Ciencias Naturales La Salle, 53, 121–126.
  86. Señaris, J.C. (1998) A new species of Typhlophis (Serpentes: Anomalepididae) from Bolívar State, Venezuela. Amphibia-Reptilia, 19 (3), 303-310. https://doi.org/10.1163/156853898X00205
  87. Señaris, J.C. (2022a) Metaphryniscus sosai (amended version of 2020 assessment). The IUCN Red List of Threatened Species, 2022, e.T54834A198628341. https://doi.org/10.2305/IUCN.UK.2022-1.RLTS.T54834A198628341.en
  88. Señaris, J.C. (2022b) Cochranella duidaeana (amended version of 2020 assessment). The IUCN Red List of Threatened Species, 2022, e.T54958A198630291. https://doi.org/10.2305/IUCN.UK.2022-1.RLTS.T54958A198630291.en
  89. Señaris, J.C. (2022c) Stefania percristata (amended version of 2020 assessment). The IUCN Red List of Threatened Species, 2022, e.T56031A198645648. https://doi.org/10.2305/IUCN.UK.2022-1.RLTS.T56031A198645648.en
  90. Señaris, J.C. (2022d) Myersiohyla aromatica (amended version of 2020 assessment). The IUCN Red List of Threatened Species, 2022, e.T55399A198641882. https://doi.org/10.2305/IUCN.UK.2022-1.RLTS.T55399A198641882.en
  91. Señaris, J.C. (2022e) Myersiohyla inparquesi (amended version of 2020 assessment). The IUCN Red List of Threatened Species, 2022, e.T87735771A198667165. https://doi.org/10.2305/IUCN.UK.2022-1.RLTS.T87735771A198667165.en
  92. Señaris, J.C., Ayarzagüena, J. & Gorzula, S.J. (1994) Los sapos de la familia Bufonidae (Amphibia: Anura) de las tierras altas de la Guayana Venezolana: descripción de un nuevo género y tres especies. Publicaciones de la Asociación de Amigos de Doñana, 3, 1–37.
  93. Señaris, J.C., Ayarzagüena, J. & Gorzula, S.J. (1997 “1996”) Revision taxonómica del genero Stefania (Anura: Hylidae) en Venezuela con la descripción de cinco nuevas especies. Publicaciones de la Asociación de Amigos de Doñana, 7, 1–57.
  94. Speybroeck, J., Beukema, W., Dufresnes, C., Fritz, U., Jablonski, D., Lymberakis, P., Martínez-Solano, I., Razzetti, E., Vamberger, M., Vences, M., Vörös, J. & Crochet, P.-A. (2020) Species list of the European herpetofauna–update by the Taxonomic Committee of the Societas Europaea Herpetologica. Amphibia-Reptilia, 41 (2), 139–189. https://doi.org/10.1163/15685381-bja10010
  95. Tamar, K., Geniez, P., Brito, J.C. & Crochet, P.-A. (2017) Systematic revision of Acanthodactylus busacki (Squamata: Lacertidae) with a description of a new species from Morocco. Zootaxa, 4276 (3), 357–386. https://doi.org/10.11646/zootaxa.4276.3.3
  96. Trape, J.-F., Chirio, L., G. Broadley, D.G. & Wüster, W. (2009) Phylogeography and systematic revision of the Egyptian cobra (Serpentes: Elapidae: Naja haje) species complex, with the description of a new species from West Africa. Zootaxa, 2236 (1), 1–25. https://doi.org/10.11646/zootaxa.2236.1.1
  97. TTWG [Rhodin, A.G.J., Iverson, J.B., Bour, R. Fritz, U., Georges, A., Shaffer, H.B. & van Dijk, P.P.] (2017) Turtles of the World: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. 8th Edition. In: Rhodin, A.G.J., Iverson, J.B., van Dijk, P.P., Saumure, R.A., Buhlmann, K.A., Pritchard, P.C.H. & Mittermeier, R.A. (Eds.), Conservation Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN/SSC Tortoise and FreshwaterTurtle Specialist Group. Chelonian Research Monographs, 7, pp. 1–292. https://doi.org/10.3854/crm.7.checklist.atlas.v8.2017
  98. Uetz, P., Freed, P., Aguilar, R., Reyes, F., Kudera, J. & Hošek, J. (Eds.) (2025) The Reptile Database. Available from: http://www.reptile-database.org (accessed 27 July 2025)
  99. Valverde, J.A. (1958) Una nueva lagartija del género Algiroides Bribron procedente de la Sierra de Cazorla (Sur de España). Archivos del Instituto de Aclimatación, Almería, 7, 127–135.
  100. Valverde, J.A. (1966) Notas sobre vertebrados II. Sobre las subespecies de Chalcides bedriagai (Boscá, 1880). Boletín de la Real Sociedad Española de Historia Natural, Sección Biologica, 64, 169–170.
  101. Valverde, J. A. (1968) Nuevo nombre para un Chalcides ibérico (Reptilia, Scincidae). Boletín de la Real Sociedad Española de Historia Natural, Sección Biologica, 66, 135.
  102. Valverde, J.A. (1989) Notas sobre vertebrados. VII. Una nueva cobra del NW de África, Naja haje legionis, ssp. nov. (Elapidae, Serpentes). Actas Jornadas IX Real Sociedad Española de Historia Natural, 2, 214–230.
  103. Velo-Antón, G., García-París, M. & Cordero Rivera, A. (2008) Patterns of nuclear and mitochondrial DNA variation in Iberian populations of Emys orbicularis (Emydidae): conservation implications. Conservation Genetics, 9, 1263–1274. https://doi.org/10.1007/s10592-007-9457-1
  104. Wagner, P., Crochet, P.-A., Martínez-Freiría, F. & Geniez, P. (2021) Uromastyx occidentalis. The IUCN Red List of Threatened Species, 2021, e.T199813A166739154. https://doi.org/10.2305/IUCN.UK.2021-3.RLTS.T199813A166739154.en
  105. Wilms, T., Wagner, P., Spawls, S., Beraduccii, J. & Malonza, P.K. (2021) Naja haje. The IUCN Red List of Threatened Species, 2021, e.T184071A1748178. https://doi.org/10.2305/IUCN.UK.2021-2.RLTS.T184071A1748178.en

How to Cite

Señaris, C., Urdiales, C., Sempere, M. & Leonard, J.A. (2026) Catalogue of type specimens of amphibians and reptiles in the Estación Biológica de Doñana (EBD-CSIC), Spain. Zootaxa, 5792 (2), 201–244. https://doi.org/10.11646/zootaxa.5792.2.1