Vol. 5589 No. 1: 19 Feb. 2025

Published: 2025-02-19

DOI: https://doi.org/10.11646/zootaxa.5589.1

Front matter

Global Diversity of Earthworms—Papers in Honor of Victor V. Pop (1936–2022), Alexander Feijoo (1963–2023), Pietro Omodeo (1919–2024) (Title page)

CSABA CSUZDI, MARIE BARTZ, DANIEL FERNÁNDEZ MARCHÁN

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.1

Page range: 1-2

Abstract views: 58

PDF downloaded: 43
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Global Diversity of Earthworms—Papers in Honor of Victor V. Pop (1936–2022), Alexander Feijoó (1963–2023), Pietro Omodeo (1919–2024) (Table of contents)

CSABA CSUZDI, MARIE BARTZ, DANIEL FERNÁNDEZ MARCHÁN

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.2

Page range: 3-5

Abstract views: 85

PDF downloaded: 50
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Introduction Special Issue on Oligochaeta

DANIEL F. MARCHÁN, CSABA CZUSDI, MARIE L. C. BARTZ

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.3

Page range: 6-7

Abstract views: 62

PDF downloaded: 43
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Editorial

Three generations in Earthworm Systematics: Homage to the work of the Pop family in the systematics of earthworms (Crassiclitellata: Lumbricidae)

CSABA CSUZDI, EMMA SHERLOCK

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.4

Page range: 8-22

Abstract views: 63

PDF downloaded: 34
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Memorial

A farewell to Alexander Feijoo Martínez (07 June 1963–05 December 2023)

GEORGE G. BROWN, CARLOS FRAGOSO, CSABA CSUZDI, PATRICK LAVELLE, ELENA VELASQUEZ, LILIANA V. CELIS

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.5

Page range: 23-32

Abstract views: 63

PDF downloaded: 39
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A tribute to a master and friend: Pietro Omodeo (1919-2024)

EMILIA ROTA

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.6

Page range: 33-67

Abstract views: 89

PDF downloaded: 66
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Article

Barcode gap delimitation and phylogenetic analyses in integrative earthworm taxonomy: Dendrobaena byblica complex

ROBABEH LATIF, ATABAK ROOHI AMINJAN, MASOUMEH MALEK, SERGEI V. SHEKHOVTSOV, TATIANA V. POLUBOYAROVA, MARIA J I BRIONES

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.7

Page range: 68-79

Abstract views: 79

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A new species of the enchytraeid genus Mesenchytraeus Eisen, 1878 (Clitellata, Enchytraeidae) from the eastern Chukot Peninsula, Russian Far East, as further evidence of former trans-Beringian faunal connections

KLÁRA DÓZSA-FARKAS, OLGA MAKAROVA

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.8

Page range: 80-90

Abstract views: 102

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Annotated Checklist of the Earthworms (Annelida: Clitellata: Megadrili) of Nepal

ANKIT KUMAR SINGH, LAXMAN KHANAL

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.9

Page range: 91-111

Abstract views: 299

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A new earthworm species of the genus Amynthas (Clitellata: Megascolecidae) from northern Taiwan, false synonymy between Amynthas corticis (Kinberg, 1867) and Amynthas sheni (Chen, 1935) and other taxonomic issues relating to A. corticis

HUEI-PING SHEN, CHIH-HAN CHANG

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.10

Page range: 112-126

Abstract views: 91

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On the biogeography of earthworms of the primitive genus Drawida Michaelsen, 1900 (Clitellata: Moniligastridae) in the Indian subcontinent

S. PRASANTH NARAYANAN, SHAILJA KUMARI, VIJO T. KURIEN, A.P. THOMAS, R. PALIWAL, J.M. JULKA

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.11

Page range: 127-141

Abstract views: 84

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Description of a new species of the earthworm genus Moniligaster Perrier, 1872 (Clitellata, Moniligastridae) from the Nilgiri Biosphere Reserve of Tamil Nadu State, India

S. PRASANTH NARAYANAN, SHAKOOR AHMED, A.P. THOMAS

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.12

Page range: 142-154

Abstract views: 63

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Discovery of a novel species of the genus Perionyx Perrier, 1872 (Clitellata: Megascolecidae) from Manipur, North-Eastern Region, India

NALINI TIWARI, SAMUEL W. JAMES, NEELIMA GUPTA, SHWETA YADAV

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.13

Page range: 155-165

Abstract views: 92

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Three Novel Species of Earthworms of Genus Metaphire Sims and Easton, 1972 from Manipur, India

NALINI TIWARI, KAUSHIK SHILPI, SAMUEL W. JAMES, NEELIMA GUPTA, SHWETA YADAV

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.14

Page range: 166-189

Abstract views: 79

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A report of the unusual presence of Haplotaxis cf. gordioides in a terrestrial subsoil and first isotopic analysis of its trophic position

AGNIESZKA JÓZEFOWSKA, PATRICK MARTIN, OLAF SCHMIDT

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.15

Page range: 190-200

Abstract views: 70

PDF downloaded: 27
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Remarks on the Dendrobaena alpina (Rosa, 1884) species complex in the Carpathian Basin (Oligochaeta, Lumbricidae)

TÍMEA SZEDERJESI, ANTONIA ADRIANA POP, EDIT HORVÁTH, VIRÁG KRÍZSIK, CSABA CSUZDI

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.16

Page range: 201-209

Abstract views: 92

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Checklist of earthworms of North Macedonia (Clitellata: Megadrili)

FILIP J. POPOVIĆ, TANJA B. TRAKIĆ, TÍMEA SZEDERJESI, DAVORKA K. HACKENBERGER, JOVANA M. SEKULIĆ

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.17

Page range: 210-234

Abstract views: 86

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Clarification of the controversy around the problematic species “Eophila gestroi” (Cognetti, 1905) in the Balkan

FILIP J. POPOVIĆ, TANJA B. TRAKIĆ, JOVANA M. SEKULIĆ, JORGE DOMÍNGUEZ, DANIEL F. MARCHÁN

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.18

Page range: 235-241

Abstract views: 86

PDF downloaded: 4
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Morphological variability, distribution and taxonomic relationships of Dendrobaena illyrica (Cognetti, 1906) (Lumbricidae)

FILIP J. POPOVIĆ, JOVANA M. SEKULIĆ, TANJA B. TRAKIĆ

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.19

Page range: 242-247

Abstract views: 69

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Earthworm fauna of Cape Kamenjak (Istria, Croatia) with a focus on the endemic species Octodrilus istrianus Mršić, 1991

DAVORKA HACKENBERGER KUTUZOVIĆ, DANIEL F. MARCHÁN, BRANIMIR HACKENBERGER KUTUZOVIĆ

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.20

Page range: 248-256

Abstract views: 84

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Earthworm fauna (Oligochaeta: Lumbricidae) of Croatian Pannonian mountains—Papuk and Požeška gora

DAVORKA HACKENBERGER KUTUZOVIĆ, TANJA TRAKIĆ, OLGA JOVANOVIĆ GLAVAŠ, TENA PLAVAC, BRANIMIR HACKENBERGER KUTUZOVIĆ

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.21

Page range: 257-264

Abstract views: 84

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On the valid name of Savigny’s „earthworm” Enterion terrestre (Crassiclitellata, Lumbricidae)

CSABA CSUZDI, EMMA SHERLOCK, TÍMEA SZEDERJESI

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.22

Page range: 265-269

Abstract views: 64

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Not the brightest earthworm in the genus: two non-bioluminescent Avelona (Lumbricidae, Crassiclitellata) with description of a new species

SYLVAIN GÉRARD, ALEJANDRO MARTÍNEZ NAVARRO, THIBAUD DECAËNS, MICKAËL HEDDE, DANIEL FERNÁNDEZ MARCHÁN

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.23

Page range: 270-281

Abstract views: 102

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Earthworms in grass lawns and native Atlantic Forest fragments at Embrapa Forestry, Colombo, Paraná state, Brazil

RAFAELA TAVARES DUDAS, WILIAN C. DEMETRIO, RODRIGO ROANI, TALITA FERREIRA, BEATRIZ VANOLLI, LUIZ P.P.M. STIEBLER, MYLLA T.M. SOUZA, SIMONE FONTOURA, ORLANDO ASSIS, SILVIO USHIWATA, LAIARA MOREIRA, MARIA E. OSSOWSKI, EMMELINE FRANÇA, EDUARDO ZORZI, RICARDO M. L. FERNANDES, JULIA NIEMEYER, CINTIA NIVA, MARIE L.C. BARTZ, SAMUEL JAMES, GEORGE G. BROWN

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.24

Page range: 282-300

Abstract views: 115

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Four new large Glossoscolex (Clitellata: Glossoscolecidae) earthworms from Brazil

RAFAELA TAVARES DUDAS, ALEXANDER FEIJOO M., GEORGE G. BROWN

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.25

Page range: 301-320

Abstract views: 115

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New earthworm species of Glossoscolex Leuckart, 1835 and Fimoscolex Michaelsen, 1900 (Clitellata: Glossoscolecidae) from southeastern Brazil

RAFAELA TAVARES DUDAS, SAMUEL WOOSTER JAMES, MARIE L.C. BARTZ, GEORGE GARDNER BROWN

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.26

Page range: 321-345

Abstract views: 98

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Annotated checklist of the earthworms (Annelida: Clitellata: Oligochaeta) from Hispaniola: a brief review and remarks

ONIEL ÁLVAREZ-ABREU, ROBERTO CARRERA-MARTÍNEZ

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.27

Page range: 346-362

Abstract views: 83

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A review of literature and a reevaluation of Diplocardia verrucosa Ude 1895 resulted in the discovery of a new species in southern New Mexico and west Texas, USA

GEORGE A. DAMOFF, ROBERTO CARRERA-MARTÍNEZ

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.28

Page range: 363-381

Abstract views: 77

PDF downloaded: 33
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The unseen diversity of the semi-aquatic earthworms of the genus Sparganophilus (Oligochaeta: Sparganophilidae) from the Southeastern Appalachian Piedmont

ROBERTO CARRERA-MARTÍNEZ, MELANIE K. TAYLOR, DANIEL JONES, SEAN D. SCHOVILLE, BRUCE A. SNYDER, MAC A. CALLAHAM

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.29

Page range: 382-409

Abstract views: 126

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Shooting the phantom: what is Octolasion tyrtaeum?

TÍMEA SZEDERJESI, CSABA CSUZDI

Published: 2025-02-19

DOI: 10.11646/zootaxa.5589.1.30

Page range: 410-415

Abstract views: 107

PDF downloaded: 5
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Vol. 5589 No. 1: 19 Feb. 2025

Front matter

Global Diversity of Earthworms—Papers in Honor of Victor V. Pop (1936–2022), Alexander Feijoo (1963–2023), Pietro Omodeo (1919–2024) (Title page)

Global Diversity of Earthworms—Papers in Honor of Victor V. Pop (1936–2022), Alexander Feijoó (1963–2023), Pietro Omodeo (1919–2024) (Table of contents)

Introduction Special Issue on Oligochaeta

Editorial

Three generations in Earthworm Systematics: Homage to the work of the Pop family in the systematics of earthworms (Crassiclitellata: Lumbricidae)

Memorial

A farewell to Alexander Feijoo Martínez (07 June 1963–05 December 2023)

A tribute to a master and friend: Pietro Omodeo (1919-2024)

Article

**Barcode gap delimitation and phylogenetic analyses in integrative earthworm taxonomy: Dendrobaena byblica complex**

A new species of the enchytraeid genus Mesenchytraeus Eisen, 1878 (Clitellata, Enchytraeidae) from the eastern Chukot Peninsula, Russian Far East, as further evidence of former trans-Beringian faunal connections

Annotated Checklist of the Earthworms (Annelida: Clitellata: Megadrili) of Nepal

A new earthworm species of the genus Amynthas (Clitellata: Megascolecidae) from northern Taiwan, false synonymy between Amynthas corticis (Kinberg, 1867) and Amynthas sheni (Chen, 1935) and other taxonomic issues relating to A. corticis

**On the biogeography of earthworms of the primitive genus Drawida Michaelsen, 1900 (Clitellata: Moniligastridae) in the Indian subcontinent**

**Description of a new species of the earthworm genus Moniligaster Perrier, 1872 (Clitellata, Moniligastridae) from the Nilgiri Biosphere Reserve of Tamil Nadu State, India**

**Discovery of a novel species of the genus Perionyx Perrier, 1872 (Clitellata: Megascolecidae) from Manipur, North-Eastern Region, India**

**Three Novel Species of Earthworms of Genus Metaphire Sims and Easton, 1972 from Manipur, India**

**A report of the unusual presence of Haplotaxis cf. gordioides in a terrestrial subsoil and first isotopic analysis of its trophic position**

**Remarks on the Dendrobaena alpina (Rosa, 1884) species complex in the Carpathian Basin (Oligochaeta, Lumbricidae)**

Checklist of earthworms of North Macedonia (Clitellata: Megadrili)

**Clarification of the controversy around the problematic species “Eophila gestroi” (Cognetti, 1905) in the Balkan**

**Morphological variability, distribution and taxonomic relationships of Dendrobaena illyrica (Cognetti, 1906) (Lumbricidae)**

**Earthworm fauna of Cape Kamenjak (Istria, Croatia) with a focus on the endemic species Octodrilus istrianus Mršić, 1991**

Earthworm fauna (Oligochaeta: Lumbricidae) of Croatian Pannonian mountains—Papuk and Požeška gora

**On the valid name of Savigny’s „earthworm” Enterion terrestre (Crassiclitellata, Lumbricidae)**

**Not the brightest earthworm in the genus: two non-bioluminescent Avelona (Lumbricidae, Crassiclitellata) with description of a new species**

Earthworms in grass lawns and native Atlantic Forest fragments at Embrapa Forestry, Colombo, Paraná state, Brazil

**Four new large Glossoscolex (Clitellata: Glossoscolecidae) earthworms from Brazil**

**New earthworm species of Glossoscolex Leuckart, 1835 and Fimoscolex Michaelsen, 1900 (Clitellata: Glossoscolecidae) from southeastern Brazil**

Annotated checklist of the earthworms (Annelida: Clitellata: Oligochaeta) from Hispaniola: a brief review and remarks

**A review of literature and a reevaluation of Diplocardia verrucosa Ude 1895 resulted in the discovery of a new species in southern New Mexico and west Texas, USA**

**The unseen diversity of the semi-aquatic earthworms of the genus Sparganophilus (Oligochaeta: Sparganophilidae) from the Southeastern Appalachian Piedmont**

**Shooting the phantom: what is Octolasion tyrtaeum?**